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<document ID-CLB-Dataset="3028" ID-DOI="http://dx.doi.org/10.3897/zookeys.1076.73103" ID-GBIF-Dataset="23f2f4fc-b426-488e-9495-009ef213e87b" ID-PMC="PMC8677706" ID-Pensoft-Pub="1313-2970-1076-109" ID-Pensoft-UUID="AB29B5B44C51580697DD02915D60AC34" ID-PubMed="34992492" ID-ZooBank="7A0C416920654FD9AC8D4470A0468B63" ModsDocID="1313-2970-1076-109" checkinTime="1639110008831" checkinUser="pensoft" docAuthor="Brunke, Adam J." docDate="2021" docId="EB0F25A806945B9B8CD30A0AB2C937C6" docLanguage="en" docName="ZooKeys 1076: 109-124" docOrigin="ZooKeys 1076" docPubDate="2021-12-09" docSource="http://dx.doi.org/10.3897/zookeys.1076.73103" docTitle="Fluviphirus elevatus Brunke 2021, comb. nov." docType="treatment" docVersion="6" id="AB29B5B44C51580697DD02915D60AC34" lastPageNumber="109" masterDocId="AB29B5B44C51580697DD02915D60AC34" masterDocTitle="New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="124" masterPageNumber="109" pageNumber="109" updateTime="1732750053587" updateUser="ExternalLinkService">
<mods:mods id="CB0DA6C2330405C455A33D1A211BBF5C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="125A518EB6FF56AA85110DA969EAAEBB">
<mods:title id="1791A08992B24997CF8EBE4645039F0C">New relictual genera in Cyrtoquediini and Indoquediini (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="0AB91D942EF0E65D6A045C9A542115B1" type="personal">
<mods:role id="FAD0DBDF5FED0F07C2AC4E300B4B29D5">
<mods:roleTerm id="944C5CCE25065C4782E66C68C6536404">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="287D578421EF35F20C0C9A46CAA943CF">Brunke, Adam J.</mods:namePart>
<mods:nameIdentifier id="D1FCA854CDB776FA148AE40B38211691" type="ORCID">https://orcid.org/0000-0003-1158-936X</mods:nameIdentifier>
<mods:affiliation id="1EAA4CABA266BD63A2C38BBFDD8D6F79">Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, 960 Carling Avenue, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier id="83E297326E3A402603ADE118E0519E60" type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="05D2B74445CDA609831F6660284FF824">text</mods:typeOfResource>
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<mods:title id="ACD10C2E23C176E4D2F91DDE77121246">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="DC71A2F75A2FC31A04402D0933B2A7E7">
<mods:date id="AEFD55D37D2953FF3BE356858AE27F96">2021</mods:date>
<mods:detail id="B38CD62F2E8348674A5BE17236AEA289" type="pubDate">
<mods:number id="5014D3F00FFE85B92C0F5CE89D8B0728">2021-12-09</mods:number>
</mods:detail>
<mods:detail id="921FA2961FB18C51580B7EF0B0D70DA2" type="volume">
<mods:number id="AA6C7B3467EB8120C24659D09968DACF">1076</mods:number>
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<mods:start id="FADB046FD3D54397CDFB1812A9DD502D">109</mods:start>
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<mods:location id="EEBEE4579DC55F00CE1BD7E1CFADCEFB">
<mods:url id="998253DC3EBFF738363B0466919026DB">http://dx.doi.org/10.3897/zookeys.1076.73103</mods:url>
</mods:location>
<mods:classification id="BE7EEF704CEFAF44809FDD967DB829EE">journal article</mods:classification>
<mods:identifier id="627CD4053626C09BA29AF4D18ABBF78E" type="DOI">http://dx.doi.org/10.3897/zookeys.1076.73103</mods:identifier>
<mods:identifier id="3DA61F3DC2E9B627BBE580EC1F1B2F23" type="Pensoft-Pub">1313-2970-1076-109</mods:identifier>
<mods:identifier id="CA7B72AB96738851C11BAB0D94EF3ED6" type="ZooBank">7A0C416920654FD9AC8D4470A0468B63</mods:identifier>
<mods:identifier id="496C2D9244F8DA738AC7C1B1A7B77D2E" type="Pensoft-UUID">AB29B5B44C51580697DD02915D60AC34</mods:identifier>
</mods:mods>
<subSection id="B6AD772F3063C84FEA00D4E968A01B46" pageId="0" pageNumber="109" type="taxonomy">
<treatment id="EB0F25A806945B9B8CD30A0AB2C937C6" ID-GBIF-Taxon="191261348" LSID="urn:lsid:plazi:treatment:EB0F25A806945B9B8CD30A0AB2C937C6" httpUri="http://treatment.plazi.org/id/EB0F25A806945B9B8CD30A0AB2C937C6" lastPageNumber="109" pageId="0" pageNumber="109" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae" scope_tribe="Indoquediini">
<subSubSection id="4E31922CA82D08A296742E8F22407D0B" pageId="0" pageNumber="109" type="nomenclature">
<paragraph id="9AAF7E667C1D2B09EB4CB8CC90C34E65" pageId="0" pageNumber="109">
<taxonomicName id="91C84BB33368D276263DE57DF9922FEC" ID-CoL="9GCPN" LSID="EB0F25A8-0694-5B9B-8CD3-0A0AB2C937C6" authority="(Hatch)" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" genus="Fluviphirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluviphirus elevatus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="elevatus" status="comb. nov.">Fluviphirus elevatus (Hatch)</taxonomicName>
<taxonomicNameLabel id="740AC14B06BE78EC88D59D988415BEB8" pageId="0" pageNumber="109">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="288D76DE75372B13502C5ADCB0EE3A19" pageId="0" pageNumber="109" type="description">
<paragraph id="4F8F43AF0D8FDA6FC656E7B3D2D22E06" pageId="0" pageNumber="109">
<figureCitation id="365DCC2868381B503760F19973C4777E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">Fig. 3A-G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2A02AAD7A732D38377DFD488F680A952" pageId="0" pageNumber="109" type="reference_group">
<paragraph id="E9ABBAE535AE05ECE8F787D876F0DCA8" pageId="0" pageNumber="109">
<taxonomicName id="01DA220D5127104352C20AEDB2E42809" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Sauridus) elevatus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="elevatus" status="comb. nov." subGenus="Sauridus">Quedius (Sauridus) elevatus</taxonomicName>
Hatch, 1957
</paragraph>
<paragraph id="AA5B91B52CF669FCB745D8FD493C602F" pageId="0" pageNumber="109">
<taxonomicName id="B1A15D7AB1D8FC60EAE942DBEB60C82D" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) elevatus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="elevatus" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) elevatus</taxonomicName>
Hatch:
<bibRefCitation id="FC6D7752C33AF081CBE9B2966C198F27" DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Universidad de Guadalajara y Conabio, Mexico" pageId="0" pageNumber="109" refId="B17" refString="Smetana, A, 1971a. https://doi.org/10.4039/entm10379fv" url="https://doi.org/10.4039/entm10379fv" year="1971 a">Smetana 1971a</bibRefCitation>
,
<bibRefCitation id="D166A6CD7B01B635C68093C5881E5359" DOI="https://doi.org/10.4039/Ent1031833-12" author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="109" pagination="1833 - 1848" refId="B18" refString="Smetana, A, 1971b. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 1. The Canadian Entomologist 103: 1833 - 1848, DOI: https://doi.org/10.4039/Ent1031833-12" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 1." url="https://doi.org/10.4039/Ent1031833-12" volume="103" year="1971 b">b</bibRefCitation>
(subgenus
<taxonomicName id="4380AF00E411E229F8CC782B6B30397A" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus)" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="subGenus" status="comb. nov." subGenus="Raphirus">Quedius Raphirus</taxonomicName>
,
<taxonomicName id="63415BDC799A11EC5B524009A519C944" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) elevatus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="elevatus" status="comb. nov." subGenus="Raphirus">Quedius Raphirus elevatus</taxonomicName>
species group)
</paragraph>
<paragraph id="64F546398D6F4E4E80292DAD99513424" pageId="0" pageNumber="109">
<taxonomicName id="BF079786827B4EBBD066CB77718D6F67" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius elevatus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="elevatus" status="comb. nov.">' Quedius ' elevatus</taxonomicName>
Hatch:
<bibRefCitation id="24816002192925D68382F54C11499649" DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="109" pagination="396 - 421" refId="B7" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46: 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. 2021</bibRefCitation>
(in undescribed genus of
<taxonomicName id="FBB0562EC8132CD4DC265907250A31EF" authorityName="Brunke" authorityYear="2021" baseAuthorityName="Brunke" baseAuthorityYear="2021" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="tribe" status="comb. nov." tribe="Indoquediini">Indoquediini</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="FCB8DC6E6F022EA5CA5B15AC60DAEAEB" pageId="0" pageNumber="109" type="type locality">
<paragraph id="D490DF6E9AE36EDCCA88BC0F6E5FA273" pageId="0" pageNumber="109">Type locality.</paragraph>
<paragraph id="C2F2F7582C9972F5EFF9E4D218D21821" pageId="0" pageNumber="109">Snoqualmie, Washington, United States.</paragraph>
</subSubSection>
<subSubSection id="11351865232CA41281332D0DD10394F7" pageId="0" pageNumber="109" type="type material">
<paragraph id="C98FA63EDB20AD7C6F5D9DC585FA5CAB" pageId="0" pageNumber="109">Type material.</paragraph>
<paragraph id="178132A6C3F8EBFECA3CD32349BFE8E1" pageId="0" pageNumber="109">The type material of this distinctive species was not examined.</paragraph>
</subSubSection>
<subSubSection id="74CF5DAAE11F69F3699CDF7B3C87F69B" pageId="0" pageNumber="109" type="type material">
<paragraph id="895A2D571B46BCB3949547F5C2751AFB" pageId="0" pageNumber="109">Non-type material.</paragraph>
<paragraph id="6A4C1C71378643465936B5C69C613412" pageId="0" pageNumber="109">
<emphasis id="3363F0A0E1D4998B39919C3B5853515C" bold="true" pageId="0" pageNumber="109">Canada: British Columbia</emphasis>
: 8 mi W Creston, ex. river debris, 10.VI.1968, J.M. Campbell &amp; A. Smetana (8, CNC); 20 mi E Hope, ex. river debris, 3.VI.1968, J.M. Campbell &amp; A. Smetana (1, CNC); 4 mi W Midway, ex. river debris, 6.VI.1968, J.M. Campbell &amp; A. Smetana (6, CNC); 16 mi W Osoyoos, ex. river debris, 5.VI.1968, J.M. Campbell &amp; A. Smetana (1, CNC); Paulson, beaver house, 7.VI.1968, J.M. Campbell &amp; A. Smetana (1, CNC); 4 mi W Rossland, 9.VI.1968, J.M. Campbell &amp; A. Smetana (2, CNC); 2 mi S Salmo, ex. river debris, 9.VI.1968, J.M. Campbell &amp; A. Smetana (3, CNC); 2 mi E Burnt Flats [Burnt Flat Junction], 9.VI.1968 (2, CNC).
<emphasis id="CB61221498A68E39CD36210CB2B09F0D" bold="true" pageId="0" pageNumber="109">United States: California</emphasis>
:
<emphasis id="02D23F966FCE8D68F88F09AEED3B7672" italics="true" pageId="0" pageNumber="109">Marin Co.</emphasis>
, Lagunitas Creek at Tocaloma, 18.III.1983, A. Smetana (17, CNC); same except 19.III.1983 (6, CNC).
<emphasis id="22BF437890E43C8765B24DA4D56AF4E7" bold="true" pageId="0" pageNumber="109">Oregon</emphasis>
: Union Co., Blue Mts., Phillips Creek Road, 9 km NW Elgin (2, CNC).
</paragraph>
</subSubSection>
<subSubSection id="6EFB3EF917CCF7EDFC9233042FB7623B" pageId="0" pageNumber="109" type="diagnosis">
<paragraph id="161F1AEE6A32FD6E32B187B481E83E87" pageId="0" pageNumber="109">Diagnosis.</paragraph>
<paragraph id="876C6DD1710852A02CB39F02C27206B3" pageId="0" pageNumber="109">As given above for the genus.</paragraph>
</subSubSection>
<subSubSection id="2D1890952B53E0B4D67438B4477E9E50" pageId="0" pageNumber="109" type="redescription">
<paragraph id="17C22CCEE9E8D31248815BA26C2BD475" pageId="0" pageNumber="109">Redescription.</paragraph>
<paragraph id="3E1F780C23D901200C227B7BC714A8BA" pageId="0" pageNumber="109">
The species was redescribed by
<bibRefCitation id="920F31893BB80EE4113EF64D7938CC94" DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Universidad de Guadalajara y Conabio, Mexico" pageId="0" pageNumber="109" refId="B17" refString="Smetana, A, 1971a. https://doi.org/10.4039/entm10379fv" url="https://doi.org/10.4039/entm10379fv" year="1971 a">Smetana (1971a)</bibRefCitation>
but this is here supplemented with additional characters specific to the male and female: male with sternite VII unmodified; sternite VIII with broad shallow emargination; tergite X constricted in apical half, with weakly emarginate apex, apical half with short fine setae on lateral parts of disc and strong, coarse setae along apical margin (Fig.
<figureCitation id="52A54077BDECFB6073DA13E310C85CA1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3E</figureCitation>
); sternite IX with moderately slender, asymmetrical base, elongate with deep and narrow emargination (Fig.
<figureCitation id="B0ED829598BB06D780FE3627B783F4A5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3F</figureCitation>
); median lobe of aedeagus in ventral view narrowed to sharp apex, apical portion with longitudinal median excavation (Fig.
<figureCitation id="135AF3DC8ED6366EC529D701E891104C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3B</figureCitation>
), paramere with broad base, with elongate triangular apical part and narrow apex (Fig.
<figureCitation id="0239D963AD743DBF50B74576B61521F8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3B</figureCitation>
); aedeagus in lateral view with paramere swollen, slightly deflexed dorsad, median lobe sinuate, narrow, with fin-like apex (Fig.
<figureCitation id="0D9802D2DF534260C8DFD0F2BAE6E822" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3C</figureCitation>
); apex of paramere with longitudinal, median cluster of peg setae, extended basad on slight ridge (Fig.
<figureCitation id="CB6986FBCE35C0734E233710311B5115" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3D</figureCitation>
). Female with tergite X narrowly elongate, with two-pronged apex, prongs separated by U-shaped emargination (Fig.
<figureCitation id="62EA9980249ED2B2BB53B5384A69AC1B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Fluviphirus elevatus (Hatch) A habitus B aedeagus, ventral view C aedeagus, lateral view D apex of paramere, underside E male tergite X F male sternite IX G female tergite X. Scale bars: 1 mm (A); 0.5 mm (B-C, E-G); 0.25 mm (D)." figureDoi="10.3897/zookeys.1076.73103.figure3" httpUri="https://binary.pensoft.net/fig/621082" pageId="0" pageNumber="109">3G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8883889E9F643AC429414EDD375FE56" pageId="0" pageNumber="109" type="distribution">
<paragraph id="B3C37AFB234E18FEF7FC66B05F9910DA" pageId="0" pageNumber="109">Distribution.</paragraph>
<paragraph id="06DE38BC1B6EBCDD6BA8E4BEB6C734CB" pageId="0" pageNumber="109">
<emphasis id="564D9FE7B4ED852E0F7CA41A0382C989" bold="true" pageId="0" pageNumber="109">Canada</emphasis>
: BC.
<emphasis id="1C320AFEF16E1AFBC6DB4FA34B134842" bold="true" pageId="0" pageNumber="109">United States</emphasis>
: CA, ID, NV, OR, WA
</paragraph>
</subSubSection>
<subSubSection id="2A6215806956301CA90826320E7AF75B" pageId="0" pageNumber="109" type="bionomics">
<paragraph id="BBB67D7AC782043BB108D9EF8EC96F8F" pageId="0" pageNumber="109">Bionomics.</paragraph>
<paragraph id="9BAAD9E67CDB75AFD032BC6513D4CBCA" pageId="0" pageNumber="109">
<bibRefCitation id="C9918EFBC7D4BDC29AC3D803C06FB3FD" DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Universidad de Guadalajara y Conabio, Mexico" pageId="0" pageNumber="109" refId="B17" refString="Smetana, A, 1971a. https://doi.org/10.4039/entm10379fv" url="https://doi.org/10.4039/entm10379fv" year="1971 a">Smetana (1971a</bibRefCitation>
,
<bibRefCitation id="A2D7B2C5DC278360901E350FF3AA7289" DOI="https://doi.org/10.4039/Ent1031833-12" author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="109" pagination="1833 - 1848" refId="B18" refString="Smetana, A, 1971b. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 1. The Canadian Entomologist 103: 1833 - 1848, DOI: https://doi.org/10.4039/Ent1031833-12" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 1." url="https://doi.org/10.4039/Ent1031833-12" volume="103" year="1971 b">b</bibRefCitation>
) reported this species from debris near water, especially along larger creeks and rivers. Longer series were found in river drift left behind after periods of high water levels. Several specimens have been found in beaver houses, but it is not known whether they regularly occur there.
</paragraph>
</subSubSection>
<subSubSection id="CE18C59B6F5AA24AD29F8D3AF2BDEAFA" pageId="0" pageNumber="109" type="comments">
<paragraph id="D70F3B532EEEE03DFD151E579B989826" pageId="0" pageNumber="109">Comments.</paragraph>
<paragraph id="A99ED349AD3CCCBEA65521EC2C42AEF7" pageId="0" pageNumber="109">
The paratype specimens mentioned by
<bibRefCitation id="E06FA69A2256335E0D7DAC99BF862266" author="Hatch, MH" journalOrPublisher="Novitates Zoologicae" pageId="0" pageNumber="109" refId="B12" refString="Hatch, MH, 1957. The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Press, Seattle, ix + 1-384 pp." title="The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Press, Seattle, ix + 1 - 384 pp." year="1957">Hatch (1957)</bibRefCitation>
from Lenore, Idaho and Pullman, Washington were not examined but indicate this
<normalizedToken id="D91405D06445F5C0585260679E5F0B98" originalValue="species’">species'</normalizedToken>
distribution is rather broad across the entire Western Cordillera. The occurrence of
<taxonomicName id="DCB7F3206C2DCDC5E33A53659379EF4B" lsidName="F. elevatus" pageId="0" pageNumber="109" rank="species" species="elevatus">
<emphasis id="29F1A522F686DBC45BCE5D86FB918C79" italics="true" pageId="0" pageNumber="109">F. elevatus</emphasis>
</taxonomicName>
along large river banks at a wide range of elevations suggests a single broadly distributed species. Specimens from California were paler than most of those from British Columbia and Oregon but no consistent differences were observed in the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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