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<document id="7D559E363CDFF0168B4BD0216B529F66" ID-DOI="10.5281/zenodo.5276613" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276613" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630016730227" checkinUser="carolina" docAuthor="Assing, V." docDate="2009" docId="EA1EF20AFFE5FFB3CEABFAE1FDF1FEDA" docLanguage="en" docName="LinzerbiolBeitr.2009.41.2.1285-1305.pdf" docOrigin="Linzer biologische Beiträge 41 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Leptusa (Stictopisalia) amisensis PACE 1982" docType="treatment" docVersion="3" lastPageNumber="1289" masterDocId="16278A72FFE1FFB6CE19FF9DFFE9FF92" masterDocTitle="New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1305" masterPageNumber="1285" pageNumber="1289" updateTime="1699137003817" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0379B7A6F69840EABA87F07B3CEA5708">New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:date id="556C723F57FD4EF092E72E33C237FEF6">2009</mods:date>
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<mods:number id="C4F7D3F18200F847B6FAEA6902073BC2">41</mods:number>
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<treatment id="EA1EF20AFFE5FFB3CEABFAE1FDF1FEDA" LSID="urn:lsid:plazi:treatment:EA1EF20AFFE5FFB3CEABFAE1FDF1FEDA" httpUri="http://treatment.plazi.org/id/EA1EF20AFFE5FFB3CEABFAE1FDF1FEDA" lastPageId="5" lastPageNumber="1289" pageId="4" pageNumber="1289">
<subSubSection id="2AAD1097FFE5FFB2CEABFAE1FCC5FA00" box="[178,812,1404,1426]" pageId="4" pageNumber="1289" type="nomenclature">
<paragraph id="6208431CFFE5FFB2CEABFAE1FCC5FA00" blockId="4.[178,812,1404,1426]" box="[178,812,1404,1426]" pageId="4" pageNumber="1289">
<heading id="3940F470FFE5FFB2CEABFAE1FCC5FA00" box="[178,812,1404,1426]" fontSize="8" level="2" pageId="4" pageNumber="1289" reason="2">
<taxonomicName id="A5B7389FFFE5FFB2CEABFAE1FD96FA00" ID-CoL="6Q2GJ" authority="PACE 1982" authorityName="PACE" authorityYear="1982" box="[178,639,1404,1426]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1289" phylum="Arthropoda" rank="species" species="amisensis" subGenus="Stictopisalia">
<emphasis id="50C39F0EFFE5FFB2CEABFAE1FD96FA00" bold="true" box="[178,639,1404,1426]" pageId="4" pageNumber="1289">
<emphasis id="50C39F0EFFE5FFB2CEABFAE1FEECFA03" bold="true" box="[178,261,1404,1425]" italics="true" pageId="4" pageNumber="1289">Leptusa</emphasis>
(
<emphasis id="50C39F0EFFE5FFB2CF0AFAE1FE7AFA00" bold="true" box="[275,403,1404,1426]" italics="true" pageId="4" pageNumber="1289">Stictopisalia</emphasis>
)
<emphasis id="50C39F0EFFE5FFB2CFBBFAE1FDEEFA03" bold="true" box="[418,519,1404,1425]" italics="true" pageId="4" pageNumber="1289">amisensis</emphasis>
<bibRefCitation id="06263EEDFFE5FFB2CC14FAE1FDAEFA02" author="PACE R" box="[525,583,1404,1426]" pageId="4" pageNumber="1289" pagination="579 - 594" refId="ref11005" refString="PACE R. (1982): Leptusa raccolte dal Dr Claude Besuchet nella Penisola Iberica, nella regione balcanica, in Turchia e in estremo oriente (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 89: 579 - 594." type="journal article" year="1982">PACE</bibRefCitation>
1982
</emphasis>
</taxonomicName>
(
<figureCitation id="FA8C5F99FFE5FFB2CC95FAE1FD31FA00" box="[652,728,1404,1426]" captionStart="Figs 49-54" captionStartId="16.[178,218,1491,1511]" captionTargetBox="[186,1079,879,1436]" captionTargetId="figure-319@16.[178,1084,873,1476]" captionTargetPageId="16" captionText="Figs 49-54: Leptusa ionopolitana PACE (49-51), L. amisensis PACE (52), and L. sengleti PACE (53- 54): (49, 53) median lobe of aedeagus in lateral view; (50-52) spermatheca; (54) male sternite VII. Scale bars: 54: 0.2 mm; 49-53: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276633" httpUri="https://zenodo.org/record/5276633/files/figure.png" pageId="4" pageNumber="1289">Fig. 52</figureCitation>
,
<figureCitation id="FA8C5F99FFE5FFB2CCFBFAE1FCCDFA00" box="[738,804,1404,1426]" captionStart="Map 2" captionStartId="4.[178,223,652,672]" captionTargetBox="[178,1085,212,628]" captionTargetId="figure-1@4.[178,1085,212,628]" captionTargetPageId="4" captionText="Map 2: Distributions of four species of the subgenus Stictopisalia in northern Turkey, based on revised records: Leptusa ionopolitana PACE (open circles), L. fibula ASSING (filled circles), L. amisensis PACE (open square), and L. flagellulifera nov.sp. (filled squares)." figureDoi="http://doi.org/10.5281/zenodo.5276617" httpUri="https://zenodo.org/record/5276617/files/figure.png" pageId="4" pageNumber="1289">Map 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AAD1097FFE5FFB2CEABFA3FFD3FFA6C" pageId="4" pageNumber="1289" type="reference_group">
<paragraph id="6208431CFFE5FFB2CEABFA3FFD1CFA25" blockId="4.[178,1085,1442,1534]" box="[178,757,1442,1463]" pageId="4" pageNumber="1289">
<treatmentCitationGroup id="42A76432FFE5FFB2CEABFA3FFD1CFA25" box="[178,757,1442,1463]" pageId="4" pageNumber="1289">
<taxonomicName id="A5B7389FFFE5FFB2CEABFA3FFE10FA25" ID-CoL="722GD" authorityName="Assing" authorityYear="2009" box="[178,505,1442,1463]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1289" phylum="Arthropoda" rank="species" species="ionopolitana" subGenus="Stictopisalia">
<emphasis id="50C39F0EFFE5FFB2CEABFA3FFF13FA25" box="[178,250,1442,1463]" italics="true" pageId="4" pageNumber="1289">Leptusa</emphasis>
(
<emphasis id="50C39F0EFFE5FFB2CF1EFA3FFE91FA25" box="[263,376,1442,1463]" italics="true" pageId="4" pageNumber="1289">Stictopisalia</emphasis>
)
<emphasis id="50C39F0EFFE5FFB2CF9CFA3FFE10FA25" box="[389,505,1442,1463]" italics="true" pageId="4" pageNumber="1289">ionopolitana</emphasis>
</taxonomicName>
<treatmentCitation id="E316650DFFE5FFB2CFE6FA3FFD1BFA25" author="PACE R" box="[511,754,1442,1463]" page="588" pageId="4" pageNumber="1289" year="1982">
<taxonomicName id="A5B7389FFFE5FFB2CFE6FA3FFD1BFA25" ID-CoL="6Q2GJ" authority="PACE 1982: 588" authorityName="PACE" authorityPageNumber="588" authorityYear="1982" box="[511,754,1442,1463]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1289" phylum="Arthropoda" rank="species" species="amisensis">
<emphasis id="50C39F0EFFE5FFB2CFE6FA3FFDBEFA25" box="[511,599,1442,1463]" italics="true" pageId="4" pageNumber="1289">amisensis</emphasis>
<bibRefCitation id="06263EEDFFE5FFB2CC44FA3EFD1BFA25" author="PACE R" box="[605,754,1443,1463]" pageId="4" pageNumber="1289" pagination="579 - 594" refId="ref11005" refString="PACE R. (1982): Leptusa raccolte dal Dr Claude Besuchet nella Penisola Iberica, nella regione balcanica, in Turchia e in estremo oriente (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 89: 579 - 594." type="journal article" year="1982">PACE 1982: 588</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6208431CFFE5FFB2CEABFA21FD3FFA6C" blockId="4.[178,1085,1442,1534]" pageId="4" pageNumber="1289">
<materialsCitation id="D2DF4941FFE5FFB2CEABFA21FD3FFA6C" collectionCode="T, V, R, MHNG" collectorName="Cl. Besuchet" country="Turkey" location="Samsun" pageId="4" pageNumber="1289" specimenCount="1" specimenCount-female="1" stateProvince="Samsun" typeStatus="holotype">
<collectionCode id="04A6DBD9FFE5FFB2CEABFA21FF29FA42" box="[178,192,1468,1488]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="1289">T</collectionCode>
y p e m a t e r i a l e x a m i n e d
<typeStatus id="BD0CFDBEFFE5FFB2CC7BFA21FD46FA42" box="[610,687,1468,1488]" pageId="4" pageNumber="1289" type="holotype">Holotype</typeStatus>
<specimenCount id="74B18895FFE5FFB2CCDCFA5DFD38FA47" box="[709,721,1472,1493]" pageId="4" pageNumber="1289" type="female">♀</specimenCount>
: &quot;
<collectingCountry id="1AA0038CFFE5FFB2CCEAFA21FCD6FA42" box="[755,831,1468,1488]" name="Turkey" pageId="4" pageNumber="1289">Turquie</collectingCountry>
,
<collectingRegion id="A0738DFEFFE5FFB2CD49FA21FC73FA42" box="[848,922,1468,1488]" country="Turkey" name="Samsun" pageId="4" pageNumber="1289">Samsun</collectingRegion>
, Samsun-Kavak, 20.
<collectionCode id="04A6DBD9FFE5FFB2CED7FA4EFF08FA75" box="[206,225,1491,1511]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="1289" type="Museum">V</collectionCode>
.67,
<collectorName id="CF4226CAFFE5FFB2CF11FA4EFE94FA75" box="[264,381,1491,1511]" pageId="4" pageNumber="1289">Cl. Besuchet</collectorName>
/
<typeStatus id="BD0CFDBEFFE5FFB2CF88FA4EFE19FA75" box="[401,496,1491,1511]" pageId="4" pageNumber="1289" type="holotype">Holotypus</typeStatus>
<taxonomicName id="A5B7389FFFE5FFB2CFEEFA4EFCFFFA75" ID-CoL="6Q2GJ" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[503,790,1491,1511]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1289" phylum="Arthropoda" rank="subSpecies" species="ionopolitana" status="subsp. nov." subSpecies="amisensis">Leptusa ionopolitana amisensis</taxonomicName>
m, det.
<collectionCode id="04A6DBD9FFE5FFB2CD7CFA4EFC9EFA75" box="[869,887,1491,1511]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="1289">R</collectionCode>
. Pace 1981 /
<taxonomicName id="A5B7389FFFE5FFB2CDECFA4EFE68FA6C" ID-CoL="6Q2GJ" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1289" phylum="Arthropoda" rank="subSpecies" species="ionopolitana" status="subsp. nov." subSpecies="amisensis">Leptusa ionopolitana amisensis</taxonomicName>
<taxonomicNameLabel id="4BF02275FFE5FFB2CF9EFA77FE2AFA6C" box="[391,451,1514,1534]" pageId="4" pageNumber="1289" rank="subSpecies">n. ssp.</taxonomicNameLabel>
, det.
<collectionCode id="04A6DBD9FFE5FFB2CFEAFA77FDECFA6C" box="[499,517,1514,1534]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="1289">R</collectionCode>
. Pace 1981&quot; (
<collectionCode id="04A6DBD9FFE5FFB2CC98FA77FD25FA6C" box="[641,716,1514,1534]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="1289" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2AAD1097FFE5FFB3CEABF991FDF1FEDA" lastPageId="5" lastPageNumber="1290" pageId="4" pageNumber="1289" type="description">
<paragraph id="6208431CFFE5FFB3CEABF991FDF1FEDA" blockId="4.[178,1085,1548,1599]" lastBlockId="5.[178,1085,219,328]" lastPageId="5" lastPageNumber="1290" pageId="4" pageNumber="1289">
C o m m e n t: The original description of
<taxonomicName id="A5B7389FFFE5FFB2CC95F990FCE4F9B0" authorityName="PACE" authorityYear="1982" box="[652,781,1549,1570]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1289" phylum="Arthropoda" rank="species" species="amisensis">
<emphasis id="50C39F0EFFE5FFB2CC95F990FCE4F9B0" box="[652,781,1549,1570]" italics="true" pageId="4" pageNumber="1289">L. amisensis</emphasis>
</taxonomicName>
is based on a single female [sic] from &quot;
<collectingCountry id="1AA0038CFFE5FFB2CF31F9B4FE94F9AD" box="[296,381,1577,1599]" name="Turkey" pageId="4" pageNumber="1289">Turquie</collectingCountry>
,
<collectingRegion id="A0738DFEFFE5FFB2CF91F9B4FE33F9AD" box="[392,474,1577,1599]" country="Turkey" name="Samsun" pageId="4" pageNumber="1289">Samsun</collectingRegion>
, Samsun-Kavak&quot; (
<bibRefCitation id="06263EEDFFE5FFB2CC83F9B4FCF8F9AD" author="PACE R" box="[666,785,1577,1599]" pageId="4" pageNumber="1289" pagination="579 - 594" refId="ref11005" refString="PACE R. (1982): Leptusa raccolte dal Dr Claude Besuchet nella Penisola Iberica, nella regione balcanica, in Turchia e in estremo oriente (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 89: 579 - 594." type="journal article" year="1982">PACE 1982</bibRefCitation>
). Although similar in colora- tion and other external characters to
<taxonomicName id="A5B7389FFFE4FFB3CC22FF41FD32FF62" authorityName="Assing" authorityYear="2009" box="[571,731,219,241]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1290" phylum="Arthropoda" rank="species" species="ionopolitana">
<emphasis id="50C39F0EFFE4FFB3CC22FF41FD32FF62" box="[571,731,219,241]" italics="true" pageId="5" pageNumber="1290">L. ionopolitana</emphasis>
</taxonomicName>
and also geographically close, it seems to represent a distinct species, as is suggested by the different shape of the spermatheca (
<figureCitation id="FA8C5F99FFE4FFB3CF0DFE88FE88FEB9" box="[276,353,277,299]" captionStart="Figs 49-54" captionStartId="16.[178,218,1491,1511]" captionTargetBox="[186,1079,879,1436]" captionTargetId="figure-319@16.[178,1084,873,1476]" captionTargetPageId="16" captionText="Figs 49-54: Leptusa ionopolitana PACE (49-51), L. amisensis PACE (52), and L. sengleti PACE (53- 54): (49, 53) median lobe of aedeagus in lateral view; (50-52) spermatheca; (54) male sternite VII. Scale bars: 54: 0.2 mm; 49-53: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276633" httpUri="https://zenodo.org/record/5276633/files/figure.png" pageId="5" pageNumber="1290">Fig. 52</figureCitation>
). Males from the
<typeStatus id="BD0CFDBEFFE4FFB3CC0CFE8BFDA8FEB9" box="[533,577,278,299]" pageId="5" pageNumber="1290">type</typeStatus>
locality (
<figureCitation id="FA8C5F99FFE4FFB3CCB8FE88FD0CFEB9" box="[673,741,277,299]" captionStart="Map 2" captionStartId="4.[178,223,652,672]" captionTargetBox="[178,1085,212,628]" captionTargetId="figure-1@4.[178,1085,212,628]" captionTargetPageId="4" captionText="Map 2: Distributions of four species of the subgenus Stictopisalia in northern Turkey, based on revised records: Leptusa ionopolitana PACE (open circles), L. fibula ASSING (filled circles), L. amisensis PACE (open square), and L. flagellulifera nov.sp. (filled squares)." figureDoi="http://doi.org/10.5281/zenodo.5276617" httpUri="https://zenodo.org/record/5276617/files/figure.png" pageId="5" pageNumber="1290">Map 2</figureCitation>
) or its vicinity would be required to clarify the status of
<taxonomicName id="A5B7389FFFE4FFB3CF8CFEAEFDFBFEDA" authorityName="PACE" authorityYear="1982" box="[405,530,307,328]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1290" phylum="Arthropoda" rank="species" species="amisensis">
<emphasis id="50C39F0EFFE4FFB3CF8CFEAEFDFBFEDA" box="[405,530,307,328]" italics="true" pageId="5" pageNumber="1290">L. amisensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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