treatments-xml/data/B0/18/43/B01843688BD95701858B3AA8CADA8447.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak &amp; Sutcharit, Chirasak" docDate="2023" docId="B01843688BD95701858B3AA8CADA8447" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma morleti Dautzenberg &amp; Fischer 1906" docType="treatment" docVersion="1" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1674655519517" updateUser="pensoft">
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<mods:title>Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Prasankok, Pongpun</mods:namePart>
<mods:affiliation>School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-24</mods:number>
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<mods:number>1142</mods:number>
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<mods:end>144</mods:end>
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<treatment LSID="urn:lsid:plazi:treatment:B01843688BD95701858B3AA8CADA8447" httpUri="http://treatment.plazi.org/id/B01843688BD95701858B3AA8CADA8447" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
23. 
<taxonomicName ID-CoL="7SZXB" LSID="B0184368-8BD9-5701-858B-3AA8CADA8447" authority="Dautzenberg &amp; Fischer, 1906" authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma morleti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="morleti">Rhiostoma morleti Dautzenberg &amp; Fischer, 1906</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Live snails of the species group IV A Rhiostoma morleti, from Vieng Xai, Houaphanh, Laos B Rhiostoma laosense sp. nov. from Hineboun, Khammouane, Laos. Photographs: K. Inkhavilay." figureDoi="10.3897/zookeys.1142.90097.figure43" httpUri="https://binary.pensoft.net/fig/799901" pageId="0" pageNumber="1">Figs 43A</figureCitation>
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Distribution map of Rhiostoma morleti - species group." figureDoi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1">, 44</figureCitation>
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">, 46</figureCitation>
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Shell of Rhiostoma morleti A specimen CUMZ 10002 / 2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos B, C specimens CUMZ 10002 / 1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos D specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos E, F specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos." figureDoi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1">, 47</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos planorbulus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="planorbulus">Pterocyclos planorbulus</taxonomicName>
Morlet, 1891: 247 (not Lamarck). Locality: Long-son [Lang Son Province, Vietnam].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma morleti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="morleti">Rhiostoma morleti</taxonomicName>
Dautzenberg &amp; Fischer, 1906: 429-431, pl. 10, figs 1-4. Type locality: Luang-Prabang, Laos; Ha Giang, Tonkin [Ha Giang Province, Vietnam]. Kobelt 1911: 755, 756, pl. 110, figs 1-4. 
<bibRefCitation author="Thach, NN" journalOrPublisher="48 HrBooks Company, Ohio, USA" pageId="0" pageNumber="1" refId="B165" refString="Thach, NN, 2016. Vietnamese New Mollusks with 59 New Species. 48HrBooks Company, Ohio, USA" title="Vietnamese New Mollusks with 59 New Species." year="2016">Thach 2016</bibRefCitation>
: 38, fig. 125. 
<bibRefCitation DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
: 168, 169, fig. 1g-k.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
MNHN-IM-2000-20961 (1 shell; Fig. 
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">46A</figureCitation>
) from Laos, chaine de partage entre le 
<normalizedToken originalValue="Mékong">Mekong</normalizedToken>
et le 
<normalizedToken originalValue="Ménam">Menam</normalizedToken>
(figured in 
<bibRefCitation author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="343 - 471" refId="B32" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M.H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles. Journal de Conchyliologie 53 (1905): 343 - 471" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles." volume="53" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
: pl. 10, figs 1, 2). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
MNHN-IM-2000-33837 (2 shells; Fig. 
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">46B</figureCitation>
) from Ha-Giang. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
MNHN-IM-2000-33838 (1 shell; Fig. 
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">46C</figureCitation>
) from Ha-Giang. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
RBINS-MT-658845 (2 shells; Fig. 
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">46D</figureCitation>
) from Not-Son, Tonkin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Laos" county="Ban Na Wid" location="Kraisorn Cave" municipality="Vieng Xai District" specimenCount="1" stateProvince="Houaphanh Province">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Laos">Laos</collectingCountry>
</emphasis>
: 
<collectingCounty>Ban Na Wid</collectingCounty>
, 
<collectingMunicipality>Vieng Xai District</collectingMunicipality>
, 
<collectingRegion country="Laos" name="Houaphan">Houaphanh Province</collectingRegion>
: CUMZ 10004/1 (20 shells), 10004/2 (3 shells; Fig. 
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">46E, F</figureCitation>
). 
<location LSID="urn:lsid:plazi:treatment:B01843688BD95701858B3AA8CADA8447:43999192518055CB401B7A6336045F40" country="Laos" county="Ban Na Wid" municipality="Vieng Xai District" name="Kraisorn Cave" stateProvince="Houaphanh Province">Kraisorn Cave</location>
, 
<location LSID="urn:lsid:plazi:treatment:B01843688BD95701858B3AA8CADA8447:43DD2786387CF5B18769A2E9A06D400D" country="Laos" county="Ban Na Wid" municipality="Vieng Xai District" name="Vieng Xai District" stateProvince="Houaphanh Province">Vieng Xai District</location>
, 
<collectingRegion country="Laos" name="Houaphan">Houaphanh Province</collectingRegion>
, Laos: CUMZ 10002/1 (15 shells; Fig. 
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Shell of Rhiostoma morleti A specimen CUMZ 10002 / 2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos B, C specimens CUMZ 10002 / 1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos D specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos E, F specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos." figureDoi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1">47B, C</figureCitation>
), 10002/2 (2 shells; Fig. 
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Shell of Rhiostoma morleti A specimen CUMZ 10002 / 2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos B, C specimens CUMZ 10002 / 1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos D specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos E, F specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos." figureDoi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1">47A</figureCitation>
). Km 31, 
<location LSID="urn:lsid:plazi:treatment:B01843688BD95701858B3AA8CADA8447:AC20CD3B539C2C18D95C3F9BF6AC0B5E" country="Laos" county="Ban Na Wid" municipality="Vieng Xai District" name="Vieng Xai District" stateProvince="Houaphanh Province">Vieng Xai District</location>
, 
<collectingRegion country="Laos" name="Houaphan">Houaphanh Province</collectingRegion>
: CUMZ 10003 (7 shells; Fig. 
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Shell of Rhiostoma morleti A specimen CUMZ 10002 / 2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos B, C specimens CUMZ 10002 / 1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos D specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos E, F specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos." figureDoi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1">47D</figureCitation>
). Tam Xang (cave), 
<location LSID="urn:lsid:plazi:treatment:B01843688BD95701858B3AA8CADA8447:CF628953035E1554F6022C0B6724D414" country="Laos" county="Ban Na Wid" municipality="Vieng Xai District" name="Mueng Kham District" stateProvince="Houaphanh Province">Mueng Kham District</location>
, 
<collectingRegion country="Laos" name="Xiangkhouang">Xiangkhouang Province</collectingRegion>
, Laos: CUMZ 10209 (4 shells; Fig. 
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Shell of Rhiostoma morleti A specimen CUMZ 10002 / 2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos B, C specimens CUMZ 10002 / 1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos D specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos E, F specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos." figureDoi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1">47E, F</figureCitation>
)
</materialsCitation>
. 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
</emphasis>
: Phong-Tho, Tonkin: SMF 130576 (4 shells), RBINS 659868 (2 shells), 659871 (2 shells).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell large, flattened to depressed shell and without detached whorl. Breathing device incomplete or canal-shaped and pointed apically. Peristome double; outer lip with narrow canal; inner lip with shallow incision or small hole.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is similar to 
<taxonomicName lsidName="R. cochinchinense" pageId="0" pageNumber="1" rank="species" species="cochinchinense">
<emphasis italics="true" pageId="0" pageNumber="1">R. cochinchinense</emphasis>
</taxonomicName>
in general shell form, but with an incomplete tube or canal-shaped breathing device, generally brownish to whitish pattern and with a peripheral band, which are the distinguishing characters. In comparison, 
<taxonomicName lsidName="R. cochinchinense" pageId="0" pageNumber="1" rank="species" species="cochinchinense">
<emphasis italics="true" pageId="0" pageNumber="1">R. cochinchinense</emphasis>
</taxonomicName>
has an incomplete tube-shaped breathing device, a uniformly whitish shell, and lacks a peripheral band.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
.
</emphasis>
Shell usually large, width 18.9-28.1 mm, height 8.7-13.0 mm, thickened, and flattened shape. Apex acute; spire flat to slightly elevated. Whorls 4 to 5, convex, increasing regularly; suture narrow and deep; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thick corneous and brown. Shell colour varies from uniformly whitish or brownish pattern, with narrow to wide dark-brown spiral band on periphery. Detached whorl absent (rarely very short). Peristome circular and double; lip thickened and slightly expanded. Aperture opened sub-laterally. Breathing device incomplete or canal-shaped, protruding apically, and usually not attached to preceding whorl; outer lip protruded and with narrow canal or forming nearly closed tube; inner lip with shallow incision or small hole inside shell. Umbilicus widely opened. Operculum calcareous, cup-shaped, and multispiral (Figs 
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Shell of Rhiostoma morleti A syntype MNHN-IM- 2000 - 20961 from chaine de partage entre le Mekong et le Menam B syntype MNHN-IM- 2000 - 33837 from Ha-Giang C syntype MNHN-IM- 2000 - 33838 from Ha-Giang D syntype RBINS-MT- 658845 from Not-Son, Tonkin E, F specimens CUMZ 10004 / 2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (A-C)." figureDoi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1">46</figureCitation>
, 
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Shell of Rhiostoma morleti A specimen CUMZ 10002 / 2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos B, C specimens CUMZ 10002 / 1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos D specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos E, F specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos." figureDoi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1">47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
The previous record of this species was from Luang Phrabang, Laos, and Ha Giang and Lang Son, North Vietnam (
<bibRefCitation author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="343 - 471" refId="B32" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M.H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles. Journal de Conchyliologie 53 (1905): 343 - 471" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles." volume="53" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
). Later records were reported from Ha Giang, Yen Bai, Son La, Lai Chau, and Dien Bien provinces, north Vietnam (
<bibRefCitation DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
). This study recorded specimens from several localities in Houaphanh Province, northern Laos (Fig. 
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Distribution map of Rhiostoma morleti - species group." figureDoi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1">44</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The identification of this species without the operculum might easily (and wrongly) place it into the genus 
<taxonomicName authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos." kingdom="Animalia" lsidName="Pterocyclos." order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos.</emphasis>
</taxonomicName>
Nevertheless, the unique calcareous cup-shaped operculum with elevated lamellae of the type specimen clearly indicates that it is 
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The two specimens figured in 
<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. (2015</bibRefCitation>
: fig. 4c, d) as 
<taxonomicName lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
sp. have the tubular-shaped breathing device that is curved, and with tip attached to the preceding whorl. This character clearly differs from the typical 
<taxonomicName lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
</taxonomicName>
, but rather should be referred to as 
<taxonomicName lsidName="R. simplicilabre" pageId="0" pageNumber="1" rank="species" species="simplicilabre">
<emphasis italics="true" pageId="0" pageNumber="1">R. simplicilabre</emphasis>
</taxonomicName>
. The specimen figured in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B65" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. (2019)</bibRefCitation>
as 
<taxonomicName lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
</taxonomicName>
from Bolikhamxay, Laos, shows considerable difference from the type specimen and from the newly collected specimens from Houaphanh Province, northern Laos. Therefore, this study recognises it as 
<taxonomicName lsidName="R. laosense" pageId="0" pageNumber="1" rank="species" species="laosense">
<emphasis italics="true" pageId="0" pageNumber="1">R. laosense</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<caption doi="10.3897/zookeys.1142.90097.figure43" httpUri="https://binary.pensoft.net/fig/799901" pageId="0" pageNumber="1" start="Figure 43" startId="F43">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 43.</emphasis>
Live snails of the species group IV 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma morleti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma morleti</emphasis>
</taxonomicName>
, from Vieng Xai, Houaphanh, Laos 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName authorityName="Tongkerd &amp; Inkhavilay" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma laosense" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="laosense">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma laosense</emphasis>
</taxonomicName>
sp. nov. from Hineboun, Khammouane, Laos. Photographs: K. Inkhavilay.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1" start="Figure 44" startId="F44">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 44.</emphasis>
Distribution map of 
<taxonomicName authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma morleti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma morleti</emphasis>
</taxonomicName>
-species group.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1142.90097.figure45" httpUri="https://binary.pensoft.net/fig/799903" pageId="0" pageNumber="1" start="Figure 45" startId="F45">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 45.</emphasis>
Shell of 
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
spp. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1857" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma cochinchinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cochinchinensis">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma cochinchinensis</emphasis>
</taxonomicName>
probable syntype NHMUK 20170354 from Cochinchina 
<emphasis bold="true" pageId="0" pageNumber="1">B-F</emphasis>
<taxonomicName baseAuthorityName="Morelet" baseAuthorityYear="1875" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma cambodjense" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambodjense">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma cambodjense</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
syntype NHMUK 1893.2.4.776 from Battambang, Cambodje 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
syntype of &quot; 
<taxonomicName authorityName="Morlet" authorityYear="1889" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus pulchellus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pulchellus">
<emphasis italics="true" pageId="0" pageNumber="1">Opisthoporus pulchellus</emphasis>
</taxonomicName>
Morlet, 1889&quot; MNHN-IM-2000-9828 from Mount. Sisophon 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
specimen CUMZ 10203/2 from Thammaban Khiri, Serei Saophoan (Sisophon), Cambodia 
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
specimens CUMZ 4714 from Khao Chakan, Srakeo. Photograph: A Robin and V Heros, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1142.90097.figure46" httpUri="https://binary.pensoft.net/fig/799904" pageId="0" pageNumber="1" start="Figure 46" startId="F46">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
Shell of 
<taxonomicName authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma morleti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma morleti</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype MNHN-IM-2000-20961 from chaine de partage entre le 
<normalizedToken originalValue="Mékong">Mekong</normalizedToken>
et le 
<normalizedToken originalValue="Ménam">Menam</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
syntype MNHN-IM-2000-33837 from Ha-Giang 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
syntype MNHN-IM-2000-33838 from Ha-Giang 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
syntype RBINS-MT-658845 from Not-Son, Tonkin 
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
specimens CUMZ 10004/2 from Ban Na Wid, Vieng Xai, Houaphanh, Laos. Photograph: M Caballer and V Heros, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1142.90097.figure47" httpUri="https://binary.pensoft.net/fig/799905" pageId="0" pageNumber="1" start="Figure 47" startId="F47">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 47.</emphasis>
Shell of 
<taxonomicName authorityName="Dautzenberg &amp; Fischer" authorityYear="1906" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma morleti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma morleti</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
specimen CUMZ 10002/2 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos 
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
specimens CUMZ 10002/1 from Kraisorn Cave, Vieng Xai, Houaphanh, Laos 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
specimen CUMZ 10003 from Km 31, Vieng Xai, Houaphanh, Laos 
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
specimen CUMZ 10209 from Tam Xang, Mueng Kham, Xiangkhouang, Laos.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>