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<document id="A799623EA42E852D430B6ADF88B260E3" ID-CLB-Dataset="55811" ID-DOI="10.1080/00222933.2012.763059" ID-GBIF-Dataset="dfc793f4-7033-4586-9849-718e7e526360" ID-ISSN="1464-5262" ID-Zenodo-Dep="556127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628879433880" checkinUser="felipe" docAuthor="Dankittipakul, Pakawin, Tavano, Maria &amp; Singtripop, Tippawan" docDate="2013" docId="A676E712FFC4FF9606C1FCA4166F8DC4" docLanguage="en" docName="JNatHist.47.23-24.1539-1567.pdf" docOrigin="Journal of Natural History 47 (23 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2012.763059" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Jacaena angoonae Dankittipakul &amp; Tavano &amp; Singtripop 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="1565" masterDocId="5A4F9F6AFFDDFF8A071FFFF6123D8C6A" masterDocTitle="Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)" masterLastPageNumber="1567" masterPageNumber="1539" pageNumber="1562" updateTime="1699079059524" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="4FE0A389644FDA5BB3EDAE5BD5193C37" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="54205D18CEEBF3485EC9C9F8E08CAA6E">
<mods:title id="85ED7E67AC944A85F550965510642DCA">Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)</mods:title>
</mods:titleInfo>
<mods:name id="FB5E3C8CB1B56D3719A0D48CFD8D57D5" type="personal">
<mods:role id="D32C3E167997FC255C4CD38ADF32DFE3">
<mods:roleTerm id="45F0444EFF9E0A5614730A76EF6C1DBA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B4BACA98661A51F31531D134D18F9A7">Dankittipakul, Pakawin</mods:namePart>
<mods:affiliation id="FAFA929D7C953CB024151487929D283E">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand; &amp; Manukau Institute of Technology, Manukau, New Zealand;</mods:affiliation>
</mods:name>
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<mods:roleTerm id="C3F74F130630A8F994733A5A29C0149C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00B358B1BE478F3F0E97E33034E8B63A">Tavano, Maria</mods:namePart>
<mods:affiliation id="318C78F5E2F24D3695FDAA126A364C4B">Museo Civico di Storia Naturale ‘ Giacomo Doria’, Genova, Italy</mods:affiliation>
</mods:name>
<mods:name id="AA63643415C97EA29948D95C43553304" type="personal">
<mods:role id="AAF6BB33AB56D2D97CBF59950606BFE4">
<mods:roleTerm id="36E1D78868DEE5EEAB6A6102DE688798">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8311BF45135054C6E99A25867E38689">Singtripop, Tippawan</mods:namePart>
<mods:affiliation id="7509B78375406143E2F7D3630911F4BD">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand;</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E820205BF5FA0D5B5DA3DEBC14073205">text</mods:typeOfResource>
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<mods:title id="50E73511CEB6CFED785F11DDF3818888">Journal of Natural History</mods:title>
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<mods:part id="5B28B6E0D232A706B18DC7B71B069C03">
<mods:date id="010B90A6309200E894431D70FA32CCA8">2013</mods:date>
<mods:detail id="64D176A206965BE2A5086DCE99F85716" type="pubDate">
<mods:number id="2A12AE16AB8CB73AF92C5AC716C428FB">2013-04-29</mods:number>
</mods:detail>
<mods:detail id="73A1B8619818B95B089FA6BE1F2CAF94" type="volume">
<mods:number id="60A5F41DB3696860C7BE4A50909BDA42">47</mods:number>
</mods:detail>
<mods:detail id="49380FDA4932916AC4049848FFE1A09B" type="issue">
<mods:number id="FA14582A1C057338B1C313A200F417A9">23 - 24</mods:number>
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<mods:start id="1381C8B9724E19EFAD494522C56D34F0">1539</mods:start>
<mods:end id="42D19915F19BE4830AF14777EC51C90C">1567</mods:end>
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<mods:url id="4DEE39B8711F2660901AC2B59167C9D5">http://dx.doi.org/10.1080/00222933.2012.763059</mods:url>
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<mods:classification id="5DA220F7F80036A6EC91A279AFAB7278">journal article</mods:classification>
<mods:identifier id="72654752DF401DA38D9A12DCEF493881" type="CLB-Dataset">55811</mods:identifier>
<mods:identifier id="A5BCFCCC057C5C93F8EFFE80342D519E" type="DOI">10.1080/00222933.2012.763059</mods:identifier>
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<treatment id="A676E712FFC4FF9606C1FCA4166F8DC4" ID-DOI="http://doi.org/10.5281/zenodo.6502565" ID-GBIF-Taxon="194854863" ID-Zenodo-Dep="6502565" LSID="urn:lsid:plazi:treatment:A676E712FFC4FF9606C1FCA4166F8DC4" httpUri="http://treatment.plazi.org/id/A676E712FFC4FF9606C1FCA4166F8DC4" lastPageId="28" lastPageNumber="1565" pageId="25" pageNumber="1562">
<subSubSection id="66C5058FFFC4FF9306C1FCA4113C8F03" box="[478,769,850,873]" pageId="25" pageNumber="1562" type="nomenclature">
<paragraph id="2E605604FFC4FF9306C1FCA4113C8F03" blockId="25.[460,786,850,905]" box="[478,769,850,873]" pageId="25" pageNumber="1562">
<heading id="7528E168FFC4FF9306C1FCA4113C8F03" box="[478,769,850,873]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="1562" reason="2">
<taxonomicName id="E9DF2D87FFC4FF9306C1FCA4109A8F03" ID-CoL="3QHZB" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[478,679,850,873]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="1562" phylum="Arthropoda" rank="species" species="angoonae" status="sp. nov.">
<emphasis id="1CAB8A16FFC4FF9306C1FCA4109A8F03" bold="true" box="[478,679,850,873]" italics="true" pageId="25" pageNumber="1562">Jacaena angoonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC4FF9305B2FCA4113C8F03" box="[685,769,850,873]" pageId="25" pageNumber="1562" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC4FF9306D3FC87112F8FE3" box="[460,786,881,905]" pageId="25" pageNumber="1562" type="description">
<paragraph id="2E605604FFC4FF9306D3FC87112F8FE3" blockId="25.[460,786,850,905]" box="[460,786,881,905]" pageId="25" pageNumber="1562">
(
<figureCitation id="B6E44A81FFC4FF9306CBFC8710688FE3" box="[468,597,881,905]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="25" pageNumber="1562">Figures 2D</figureCitation>
,
<figureCitation id="B6E44A81FFC4FF93057FFC8410BC8FE3" box="[608,641,882,905]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="25" pageNumber="1562">7E</figureCitation>
,
<figureCitation id="B6E44A81FFC4FF930592FC8710938FE3" box="[653,686,881,905]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="25" pageNumber="1562">8E</figureCitation>
,
<figureCitation id="B6E44A81FFC4FF9305A5FC8711378FE3" box="[698,778,881,905]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="25" pageNumber="1562">14A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC4FF930790FC17119888C5" pageId="25" pageNumber="1562" type="materials_examined">
<paragraph id="2E605604FFC4FF930790FC1713158F92" blockId="25.[143,1104,992,1151]" box="[143,296,992,1016]" pageId="25" pageNumber="1562">
<heading id="7528E168FFC4FF930790FC1713158F92" box="[143,296,992,1016]" fontSize="10" level="3" pageId="25" pageNumber="1562" reason="5">
<emphasis id="1CAB8A16FFC4FF930790FC1713158F92" box="[143,296,992,1016]" italics="true" pageId="25" pageNumber="1562">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC4FF930790FBF113898815" blockId="25.[143,1104,992,1151]" pageId="25" pageNumber="1562">
<materialsCitation id="9EB75C59FFC4FF930790FBF113898815" ID-GBIF-Occurrence="3757642309" collectingDate="1983-06-04" collectingMethod="extraction of soil sample" collectionCode="MHNG" collectorName="P. J. Schwendinger" country="Thailand" county="Mae Taeng District" elevation="1930" location="Huay Nam Dang NP" municipality="Pai District" pageId="25" pageNumber="1562" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai" typeStatus="holotype">
<typeStatus id="F164E8A6FFC4FF930790FBF112C38875" box="[143,254,1031,1055]" pageId="25" pageNumber="1562" type="holotype">Holotype</typeStatus>
:
<specimenCount id="38D99D8DFFC4FF930618FBFE1328884A" box="[263,277,1032,1056]" pageId="25" pageNumber="1562" type="female">♀</specimenCount>
,
<collectingCountry id="56C81694FFC4FF93063DFBF113B18875" box="[290,396,1031,1055]" name="Thailand" pageId="25" pageNumber="1562">Thailand</collectingCountry>
:
<location id="2B0000DFFFC4FF93068AFBFE10BC8875" LSID="urn:lsid:plazi:treatment:A676E712FFC4FF9606C1FCA4166F8DC4:2B0000DFFFC4FF93068AFBFE10BC8875" box="[405,641,1032,1056]" country="Thailand" county="Mae Taeng District" municipality="Pai District" name="Huay Nam Dang NP" pageId="25" pageNumber="1562" stateProvince="Chiang Mai">Huay Nam Dang NP</location>
,
<location id="2B0000DFFFC4FF930595FBF111848855" LSID="urn:lsid:plazi:treatment:A676E712FFC4FF9606C1FCA4166F8DC4:2B0000DFFFC4FF930595FBF111848855" country="Thailand" county="Mae Taeng District" municipality="Pai District" name="joint border between Mae Taeng District of Chiang Mai Province and Pai District of Mae Hong Son Province" pageId="25" pageNumber="1562" stateProvince="Chiang Mai">joint border between Mae Taeng District of Chiang Mai Province and Pai District of Mae Hong Son Province</location>
,
<location id="2B0000DFFFC4FF9304D8FBD116768855" LSID="urn:lsid:plazi:treatment:A676E712FFC4FF9606C1FCA4166F8DC4:2B0000DFFFC4FF9304D8FBD116768855" box="[967,1099,1063,1087]" country="Thailand" county="Mae Taeng District" municipality="Pai District" name="Doi Chang" pageId="25" pageNumber="1562" stateProvince="Chiang Mai">Doi Chang</location>
,
<location id="2B0000DFFFC4FF930790FBBE13508835" LSID="urn:lsid:plazi:treatment:A676E712FFC4FF9606C1FCA4166F8DC4:2B0000DFFFC4FF930790FBBE13508835" box="[143,365,1095,1120]" country="Thailand" county="Mae Taeng District" municipality="Pai District" name="evergreen hill forest" pageId="25" pageNumber="1562" stateProvince="Chiang Mai">evergreen hill forest</location>
,
<quantity id="E927FBE1FFC4FF930667FBB113F08835" box="[376,461,1095,1119]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="25" pageNumber="1562" unit="m" value="1930.0">
<elevation id="A5F2B137FFC4FF930667FBB113F08835" box="[376,461,1095,1119]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="25" pageNumber="1562" unit="m" value="1930.0">1930 m</elevation>
</quantity>
,
<collectingMethod id="F79E2E13FFC4FF9306C8FBB110D18835" box="[471,748,1095,1119]" pageId="25" pageNumber="1562">extraction of soil sample</collectingMethod>
,
<date id="5A6170C4FFC4FF9305E8FBB111408835" box="[759,893,1095,1119]" pageId="25" pageNumber="1562" value="1983-06-04">
<collectingDate id="4A25892CFFC4FF9305E8FBB111408835" box="[759,893,1095,1119]" pageId="25" pageNumber="1562" value="1983-06-04">4 June 1983</collectingDate>
</date>
,
<collectorName id="832A33D2FFC4FF930497FBBE166D8835" box="[904,1104,1095,1119]" pageId="25" pageNumber="1562">P.J. Schwendinger</collectorName>
leg. (
<collectionCode id="48CECEC1FFC4FF9307DAFB91131E8815" box="[197,291,1127,1151]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="25" pageNumber="1562">MHNG</collectionCode>
, PDC 4650).
</materialsCitation>
</paragraph>
<paragraph id="2E605604FFC4FF930790FB6E119888C5" blockId="25.[143,933,1175,1200]" box="[143,933,1175,1200]" pageId="25" pageNumber="1562">
<materialsCitation id="9EB75C59FFC4FF930790FB6E119C88C5" ID-GBIF-Occurrence="3757642311" box="[143,929,1175,1200]" collectingDate="1983-06-04" collectorName="Huay Nam Dang NP &amp; P. J. Schwendinger" country="Thailand" county="Mae Taeng District" elevation="1930" location="Doi Chang" municipality="Pai District" pageId="25" pageNumber="1562" specimenCount="3" specimenCount-female="3" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="F164E8A6FFC4FF930790FB6E133C88DA" box="[143,257,1176,1200]" pageId="25" pageNumber="1562" type="paratype">Paratypes</typeStatus>
: Same data as holotype,
<specimenCount id="38D99D8DFFC4FF930509FB61100C88C5" box="[534,561,1175,1199]" pageId="25" pageNumber="1562" type="female">3♀</specimenCount>
(NHML &amp; THNM, PDC 4651)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC4FF930790FAF011D7898E" pageId="25" pageNumber="1562" type="diagnosis">
<paragraph id="2E605604FFC4FF930790FAF012C68977" blockId="25.[143,1103,1286,1508]" box="[143,251,1286,1309]" pageId="25" pageNumber="1562">
<heading id="7528E168FFC4FF930790FAF012C68977" box="[143,251,1286,1309]" fontSize="10" level="3" pageId="25" pageNumber="1562" reason="5">
<emphasis id="1CAB8A16FFC4FF930790FAF012C68977" box="[143,251,1286,1309]" italics="true" pageId="25" pageNumber="1562">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC4FF930790FADB11D7898E" blockId="25.[143,1103,1286,1508]" pageId="25" pageNumber="1562">
Females of this new species bear very close resemblance to those of
<taxonomicName id="E9DF2D87FFC4FF9304A4FAD81284890F" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="1562" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis id="1CAB8A16FFC4FF9304A4FAD81284890F" italics="true" pageId="25" pageNumber="1562">J. schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC4FF9307DAFABA1378890E" box="[197,325,1356,1380]" pageId="25" pageNumber="1562" rank="species">comb. nov.</taxonomicNameLabel>
in having large, medially situated copulatory orifices, and similar shape of spermathecae. However, in
<taxonomicName id="E9DF2D87FFC4FF93052AFA9B108489EE" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[565,697,1389,1412]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="1562" phylum="Arthropoda" rank="species" species="angoonae" status="sp. nov.">
<emphasis id="1CAB8A16FFC4FF93052AFA9B108489EE" box="[565,697,1389,1412]" italics="true" pageId="25" pageNumber="1562">J. angoonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC4FF9305DDFA9B112489EE" box="[706,793,1389,1412]" pageId="25" pageNumber="1562" rank="species">sp. nov.</taxonomicNameLabel>
the copulatory orifices are teardrop-shaped (
<figureCitation id="B6E44A81FFC4FF93064AFA7A13FB89CE" box="[341,454,1420,1444]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="25" pageNumber="1562">Figure 7E</figureCitation>
) and the insemination ducts are tubular without any modification (
<figureCitation id="B6E44A81FFC4FF9307E7FA5A135789AE" box="[248,362,1452,1476]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="25" pageNumber="1562">Figure 8E</figureCitation>
) whereas in
<taxonomicName id="E9DF2D87FFC4FF9306EBFA5B109989AE" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[500,676,1452,1476]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="1562" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis id="1CAB8A16FFC4FF9306EBFA5B109989AE" box="[500,676,1452,1476]" italics="true" pageId="25" pageNumber="1562">J. schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC4FF9305B4FA5A111989AE" box="[683,804,1452,1476]" pageId="25" pageNumber="1562" rank="species">comb. nov.</taxonomicNameLabel>
the orifices are subcircular (
<figureCitation id="B6E44A81FFC4FF930788FA3A1336898E" box="[151,267,1484,1508]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="25" pageNumber="1562">Figure 7A</figureCitation>
) and the median part of the ducts is greatly enlarged (
<figureCitation id="B6E44A81FFC4FF930476FA3A11E1898E" box="[873,988,1484,1508]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="25" pageNumber="1562">Figure 8F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC4FF930790F9CC10D18AF0" pageId="25" pageNumber="1562" type="etymology">
<paragraph id="2E605604FFC4FF930790F9CC133A8A38" blockId="25.[143,1103,1594,1690]" box="[143,263,1594,1618]" pageId="25" pageNumber="1562">
<heading id="7528E168FFC4FF930790F9CC133A8A38" box="[143,263,1594,1618]" fontSize="10" level="3" pageId="25" pageNumber="1562" reason="5">
<emphasis id="1CAB8A16FFC4FF930790F9CC133A8A38" box="[143,263,1594,1618]" italics="true" pageId="25" pageNumber="1562">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC4FF930790F99410D18AF0" blockId="25.[143,1103,1594,1690]" pageId="25" pageNumber="1562">The specific epithet is an eponymous patronym (genitive form of a noun) dedicated to Dr Angoon Lewvanich, a prolific Thai taxonomist.</paragraph>
</subSubSection>
<caption id="7AA0068CFFC7FF9007CCFBD213EF881E" ID-DOI="http://doi.org/10.5281/zenodo.6517678" ID-Zenodo-Dep="6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="26" pageNumber="1563" startId="26.[211,278,1060,1082]" targetBox="[213,1167,169,1027]" targetPageId="26">
<paragraph id="2E605604FFC7FF9007CCFBD213EF881E" blockId="26.[211,1172,1060,1140]" pageId="26" pageNumber="1563">
Figure 14.
<taxonomicName id="E9DF2D87FFC7FF900657FBD313C18850" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[328,508,1061,1082]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="1563" phylum="Arthropoda" rank="species" species="angoonae" status="sp. nov.">
<emphasis id="1CAB8A16FFC7FF900657FBD313C18850" box="[328,508,1061,1082]" italics="true" pageId="26" pageNumber="1563">Jacaena angoonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC7FF90051CFBD2106E8853" box="[515,595,1060,1081]" pageId="26" pageNumber="1563" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female (A–C);
<taxonomicName id="E9DF2D87FFC7FF900443FBD311EE8850" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[860,979,1060,1082]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="1563" phylum="Arthropoda" rank="species" species="peculiaris" status="sp. nov.">
<emphasis id="1CAB8A16FFC7FF900443FBD311EE8850" box="[860,979,1060,1082]" italics="true" pageId="26" pageNumber="1563">J. peculiaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC7FF9004C5FBD216178853" box="[986,1066,1060,1081]" pageId="26" pageNumber="1563" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C).
</paragraph>
</caption>
<subSubSection id="66C5058FFFC7FF9607CCFB4A130E8D4D" lastPageId="28" lastPageNumber="1565" pageId="26" pageNumber="1563" type="description">
<paragraph id="2E605604FFC7FF9007CCFB4A136F88B9" blockId="26.[211,1172,1212,1691]" box="[211,338,1212,1235]" pageId="26" pageNumber="1563">
<heading id="7528E168FFC7FF9007CCFB4A136F88B9" box="[211,338,1212,1235]" fontSize="10" level="3" pageId="26" pageNumber="1563" reason="5">
<emphasis id="1CAB8A16FFC7FF9007CCFB4A136F88B9" box="[211,338,1212,1235]" italics="true" pageId="26" pageNumber="1563">Description</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC7FF9007CCFB12119189B0" blockId="26.[211,1172,1212,1691]" pageId="26" pageNumber="1563">
<emphasis id="1CAB8A16FFC7FF9007CCFB12131F8896" box="[211,290,1252,1276]" italics="true" pageId="26" pageNumber="1563">Female</emphasis>
(
<typeStatus id="F164E8A6FFC7FF900626FB15139F8891" box="[313,418,1251,1275]" pageId="26" pageNumber="1563" type="holotype">holotype</typeStatus>
). Total length 7.5. Carapace 3.0 long, 2.4 wide. Opisthosoma 4.5 long, 2.6 wide. Eye sizes and interdistances: AME 0.10; ALE 0.13; PME 0.08; PLE 0.11; AME–AME 0.06; AME–ALE 0.05; PME–PME 0.13; PME–PLE 0.16. MOQ 0.36 long; front width 0.28; back width 0.33. Leg formula 4123. Spination. Leg I: femur 1–1pl; tibia 8pl 8rl; metatarsus 4pl 4rl. Leg II: tibia 7pl 6rl; metatarsus 4pl 4rl. Leg III: tibia 1–1–1v 1pl; metatarsus 1–1v 1pl. Leg IV: tibia 1–1–1–1–2v 1–1rl; metatarsus 1–1– 1v 1pl 1rl. Leg measurements: Leg I 8.6 (2.5, 1.0, 2.3, 1.85, 1.0); II 7.7 (2.2, 0.9, 1.9, 1.7, 1.0); III 6.1 (1.7, 0.8, 1.4, 0.85); IV 9.2 (2.5, 1.0, 2.1, 2.4, 1.2).
</paragraph>
<paragraph id="2E605604FFC7FF9007E1FA14105E8AF0" blockId="26.[211,1172,1212,1691]" pageId="26" pageNumber="1563">
Colouration and pattern (
<figureCitation id="B6E44A81FFC7FF900500FA1410A58990" box="[543,664,1506,1530]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="26" pageNumber="1563">Figure 2D</figureCitation>
): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown; coxae, trochanters and femora dark brown; patellae and tibiae brown, tibiae orange distally; metatarsi and tarsi I–II light brown, III–IV orange. Opisthosoma ovoid; dorsum with pattern consisting of paired, oblique patches, followed by three chevrons on dark grey background.
</paragraph>
<caption id="7AA0068CFFC6FF910790FAEF11E58926" ID-DOI="http://doi.org/10.5281/zenodo.6517680" ID-Zenodo-Dep="6517680" httpUri="https://zenodo.org/record/6517680/files/figure.png" pageId="27" pageNumber="1564" startId="27.[143,210,1305,1327]" targetBox="[145,1102,169,1272]" targetPageId="27">
<paragraph id="2E605604FFC6FF910790FAEF11E58926" blockId="27.[143,1104,1305,1356]" pageId="27" pageNumber="1564">
Figure 15.
<taxonomicName id="E9DF2D87FFC6FF91061BFAEC138D8945" authorityName="Thorell" authorityYear="1897" box="[260,432,1305,1327]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="1564" phylum="Arthropoda" rank="species" species="distincta">
<emphasis id="1CAB8A16FFC6FF91061BFAEC138D8945" box="[260,432,1305,1327]" italics="true" pageId="27" pageNumber="1564">Jacaena distincta</emphasis>
</taxonomicName>
, holotype, female (MSNG). Museum label (A). Original label by Thorell (B). Habitus, dorsal (C). Ditto, ventral (D). Epigyne, partially removed (E).
</paragraph>
</caption>
<paragraph id="2E605604FFC6FF9107A5FA7513EF8A30" blockId="27.[143,1104,1411,1626]" pageId="27" pageNumber="1564">
Copulatory organ (
<figureCitation id="B6E44A81FFC6FF91068BFA75102E89F1" box="[404,531,1411,1435]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="27" pageNumber="1564">Figures 7E</figureCitation>
,
<figureCitation id="B6E44A81FFC6FF91053FFA75107C89F1" box="[544,577,1411,1435]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="27" pageNumber="1564">8E</figureCitation>
,
<figureCitation id="B6E44A81FFC6FF910551FA75109F89F1" box="[590,674,1411,1435]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="27" pageNumber="1564">14A–C</figureCitation>
): Copulatory orifices teardrop-shaped (
<figureCitation id="B6E44A81FFC6FF910788FA55131C89D1" box="[151,289,1443,1467]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="27" pageNumber="1564">Figure 14A</figureCitation>
). Insemination ducts elongated, loosely coiled (
<figureCitation id="B6E44A81FFC6FF910453FA5511EE89D1" box="[844,979,1443,1467]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="27" pageNumber="1564">Figure 14B</figureCitation>
); proximal duct connected to copulatory orifice semi-transparent (
<figureCitation id="B6E44A81FFC6FF910419FA34114189B0" box="[774,892,1474,1498]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="27" pageNumber="1564">Figure 8E</figureCitation>
), forming sigmoid arch; secretory ampullae situated on posterior surface of proximal ducts; distal half of duct ascending anteriorly then descending posteriorly to connect with spermathecae. Spermathecae elliptical, anterior part distinctly enlarged, more or less spherical with glandular pore (
<figureCitation id="B6E44A81FFC6FF91065CF9B413FB8A30" box="[323,454,1602,1626]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="27" pageNumber="1564">Figure 14B</figureCitation>
).
</paragraph>
<paragraph id="2E605604FFC6FF910790F974136C8AF0" blockId="27.[143,337,1666,1690]" box="[143,337,1666,1690]" pageId="27" pageNumber="1564">
<heading id="7528E168FFC6FF910790F974136C8AF0" box="[143,337,1666,1690]" fontSize="10" level="3" pageId="27" pageNumber="1564" reason="5">
<emphasis id="1CAB8A16FFC6FF910790F97412F48AF0" box="[143,201,1666,1690]" italics="true" pageId="27" pageNumber="1564">Male</emphasis>
. Unknown.
</heading>
</paragraph>
<paragraph id="2E605604FFC1FF9607CCFF5E13418CAA" blockId="28.[211,1172,168,295]" box="[211,380,168,192]" pageId="28" pageNumber="1565">
<heading id="7528E168FFC1FF9607CCFF5E13418CAA" box="[211,380,168,192]" fontSize="10" level="3" pageId="28" pageNumber="1565" reason="5">
<emphasis id="1CAB8A16FFC1FF9607CCFF5E13418CAA" box="[211,380,168,192]" italics="true" pageId="28" pageNumber="1565">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC1FF9607CCFF26130E8D4D" blockId="28.[211,1172,168,295]" pageId="28" pageNumber="1565">
<taxonomicName id="E9DF2D87FFC1FF9607CCFF26139D8C8D" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[211,416,208,231]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="1565" phylum="Arthropoda" rank="species" species="angoonae" status="sp. nov.">
<emphasis id="1CAB8A16FFC1FF9607CCFF26139D8C8D" box="[211,416,208,231]" italics="true" pageId="28" pageNumber="1565">Jacaena angoonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC1FF9606B2FF26103A8C8D" box="[429,519,208,231]" pageId="28" pageNumber="1565" rank="species">sp. nov.</taxonomicNameLabel>
inhabits evergreen hill forest of Doi Chang (
<quantity id="E927FBE1FFC1FF960335FF3916BA8C8D" box="[1066,1159,207,231]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="28" pageNumber="1565" unit="m" value="1930.0">1930 m</quantity>
). This new species marks the highest altitudinal record among members of the genus
<taxonomicName id="E9DF2D87FFC1FF9607CCFEE613168D4D" authorityName="Thorell" authorityYear="1897" box="[211,299,272,295]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="1565" phylum="Arthropoda" rank="genus">
<emphasis id="1CAB8A16FFC1FF9607CCFEE613168D4D" box="[211,299,272,295]" italics="true" pageId="28" pageNumber="1565">Jacaena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC1FF9607CCFE99166F8DC4" box="[211,1106,367,431]" pageId="28" pageNumber="1565" type="distribution">
<paragraph id="2E605604FFC1FF9607CCFE9913658DED" blockId="28.[211,1106,367,431]" box="[211,344,367,391]" pageId="28" pageNumber="1565">
<heading id="7528E168FFC1FF9607CCFE9913658DED" box="[211,344,367,391]" fontSize="10" level="3" pageId="28" pageNumber="1565" reason="5">
<emphasis id="1CAB8A16FFC1FF9607CCFE9913658DED" box="[211,344,367,391]" italics="true" pageId="28" pageNumber="1565">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC1FF9607CCFE61166F8DC4" blockId="28.[211,1106,367,431]" box="[211,1106,406,431]" pageId="28" pageNumber="1565">
Known only from the
<typeStatus id="F164E8A6FFC1FF9606D2FE6113C08DC5" box="[461,509,407,431]" pageId="28" pageNumber="1565">type</typeStatus>
locality,
<location id="2B0000DFFFC1FF96057CFE6111528DC4" LSID="urn:lsid:plazi:treatment:A676E712FFC4FF9606C1FCA4166F8DC4:2B0000DFFFC1FF96057CFE6111528DC4" box="[611,879,406,431]" country="Thailand" name="Mae Hongson Province" pageId="28" pageNumber="1565">Mae Hongson Province</location>
, northern
<collectingCountry id="56C81694FFC1FF9604FBFE6016738DC4" box="[996,1102,406,430]" name="Thailand" pageId="28" pageNumber="1565">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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