180 lines
22 KiB
XML
180 lines
22 KiB
XML
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<mods:title id="31E393F126095722FBC64D023266D98A">Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover</mods:title>
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<mods:namePart id="717376FF55F81DD87B919E669A463265">Squires, Richard L.</mods:namePart>
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<mods:title id="381587B673E831649AC04420570D7954">PaleoBios</mods:title>
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<mods:part id="C878F8B5F21B0C632E078C4C9E686DDC">
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<mods:date id="845A6821C278D02A49E0A2C7388D05B3">2023</mods:date>
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<paragraph id="1A09685AFFCBFFB0FF79FDFDFD7FFD85" blockId="24.[219,693,582,644]" box="[219,693,582,608]" pageId="24" pageNumber="23">
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<heading id="4141DF36FFCBFFB0FF79FDFDFD7FFD85" allCaps="true" box="[219,693,582,608]" centered="true" fontSize="10" level="2" pageId="24" pageNumber="23" reason="2">
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<taxonomicName id="DDB613D9FFCBFFB0FF79FDFDFD7FFD85" ID-CoL="C5KVQ" authority="GROVES, 2011" authorityName="Groves" authorityYear="2011" box="[219,693,582,608]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="6" phylum="Mollusca" rank="species" species="batequensis">
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<emphasis id="28C2B448FFCBFFB0FF79FDFDFDC9FD85" box="[219,515,582,608]" italics="true" pageId="24" pageNumber="23">EOCYPRAEA BATEQUENSIS</emphasis>
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GROVES, 2011
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="1A09685AFFCBFFB0FE24FDD2FDC0FD61" blockId="24.[219,693,582,644]" box="[390,522,617,644]" pageId="24" pageNumber="23">
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<figureCitation id="828D74DFFFCBFFB0FE24FDD2FDC0FD61" box="[390,522,617,644]" captionStart="Figure 7" captionStartId="23.[112,199,200,226]" captionTargetId="figure-13@22.[139,1459,192,1920]" captionTargetPageId="22" captionText="Figure 7A–C. Cypraeidae, genus and species indeterminate, Lodo Formation (basal) (Paleocene, Seldanian Stage), Tumey Hills, Fresno County, central California. Basal (A), dorsal (B), and right (labral) (C) views, 1.3x, length 39 mm, width 25 mm, height 22 mm, of hypotype CASG 61666.06, CASG Locality 61666. D–F. Eocypraea batequensis (Groves, 2011), Bateque Formation (Eocene, “Capay Stage”), Mesa La Salina, Baja California Sur, México. Basal (D), dorsal (E), and right (labral) (F) views, 4.8x, length 9.4 mm, width 6.5 mm, height 5.1 mm, of holotype IGM 5174, LACMIP Locality 16951 [ex CSUN Locality1220b]. G–I. Eocypraea crescentensis (Groves, 2011), Crescent Formation (Eocene, “Capay Stage”), Larch Mountain, Black Hills area, Thurston County, Washington. Basal (G), dorsal (H), and left (columellar) (I) views, 3.9x, length 10.9 mm, width 7.6 mm, height 6.1 mm, of holotype LACMIP 16655.40, LACMIP Type 13646, LACMIP Locality 16655 [= LACMIP Locality 41563; ex CSUN Locality 1563]. J–L. Eocypraea jimgoederti (Groves, 2011), lower McIntosh Formation (middle Eocene, “Transition Stage” to “Tejon Stage”), Doty Hills, Lewis County, Washington. Basal (J), dorsal (K), and left (columellar) side (L) views, 3.5x, length 12.7 mm, width 8.2 mm, height 6.9 mm, of holotype LACMIP 41573.1, LACMIP Type 13647, LACMIP Locality 41573 [ex CSUN Locality 1573]. M–O. Eocypraea judithsmithae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (M), dorsal (N), and left (labral) (O) views, 1.5x, length 26.6 mm, width 18.4 mm, height 12.6 mm, of holotype LACMIP 40372.1, LACMIP Type 14940 (LACMIP Locality 40732)." figureDoi="http://doi.org/10.5281/zenodo.13869600" httpUri="https://zenodo.org/record/13869600/files/figure.png" pageId="24" pageNumber="23">FIGS. 7D–F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="52AC3BD1FFCBFFB0FF32FD0BFD3EFC78" pageId="24" pageNumber="23" type="description">
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<paragraph id="1A09685AFFCBFFB0FF32FD0BFF34FD08" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
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<taxonomicName id="DDB613D9FFCBFFB0FF32FD0BFEDBFD29" authorityName="Cossmann" authorityYear="1903" box="[144,273,688,716]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="23" phylum="Mollusca" rank="genus">
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<emphasis id="28C2B448FFCBFFB0FF32FD0BFEDBFD29" box="[144,273,688,716]" italics="true" pageId="24" pageNumber="23">Eocypraea</emphasis>
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</taxonomicName>
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? sp.
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<bibRefCitation id="7E2715ABFFCBFFB0FEF1FD0BFD00FD2F" author="Squires, R. L. & R. A. Demetrion" box="[339,714,688,714]" pageId="24" pageNumber="23" pagination="1 - 55" refId="ref43379" refString="Squires, R. L. and R. A. Demetrion. 1992. Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Contributions in Science, Natural History Museum of Los Angeles County 434: 1 - 55, figs. 1 - 144." type="journal article" year="1992">Squires and Demetrion, 1992</bibRefCitation>
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||
. p. 31, figs. 77–79.
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</paragraph>
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<paragraph id="1A09685AFFCBFFB0FF32FD4DFEEDFCD1" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
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<taxonomicName id="DDB613D9FFCBFFB0FF32FD4DFE64FCF4" box="[144,430,758,786]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="23" phylum="Mollusca" rank="subGenus" subGenus="Eocypraea">
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<emphasis id="28C2B448FFCBFFB0FF32FD4DFEDBFCF7" box="[144,273,758,786]" italics="true" pageId="24" pageNumber="23">Eocypraea</emphasis>
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(
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<emphasis id="28C2B448FFCBFFB0FE80FD4DFE69FCF7" box="[290,419,758,786]" italics="true" pageId="24" pageNumber="23">Eocypraea</emphasis>
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)
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</taxonomicName>
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n. sp. 1.
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<bibRefCitation id="7E2715ABFFCBFFB0FDBAFD4DFD71FCF5" author="Groves, L. T." box="[536,699,758,784]" pageId="24" pageNumber="23" pagination="7 - 10" refId="ref38049" refString="Groves, L. T. 1997. Fossil and recent species of eastern Pacific Cypraeacea (Cypraeidae and Eocypraeinae [Ovulidae]): An update. The Western Society of Malacology Annual Report 29: 7 - 10." type="book chapter" year="1997">Groves, 1997</bibRefCitation>
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||
. p. 8 [
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||
<emphasis id="28C2B448FFCBFFB0FCA7FD4EFF4DFCD1" italics="true" pageId="24" pageNumber="23">as E.</emphasis>
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(
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<emphasis id="28C2B448FFCBFFB0FF3AFCA3FF65FCD1" box="[152,175,792,820]" italics="true" pageId="24" pageNumber="23">E.</emphasis>
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) n. sp. 1].
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</paragraph>
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||
<paragraph id="1A09685AFFCBFFB0FF32FC86FE62FC9F" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
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||
<taxonomicName id="DDB613D9FFCBFFB0FF32FC86FD30FCB2" authority="Groves, 2011" authorityName="Groves" authorityYear="2011" box="[144,762,827,857]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="23" phylum="Mollusca" rank="species" species="batequensis" subGenus="Eocypraea">
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<emphasis id="28C2B448FFCBFFB0FF32FC86FEDBFCBC" box="[144,273,829,857]" italics="true" pageId="24" pageNumber="23">Eocypraea</emphasis>
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(
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<emphasis id="28C2B448FFCBFFB0FE87FC86FE6CFCBC" box="[293,422,829,857]" italics="true" pageId="24" pageNumber="23">Eocypraea</emphasis>
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)
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||
<emphasis id="28C2B448FFCBFFB0FE19FC80FD81FCB2" box="[443,587,827,855]" italics="true" pageId="24" pageNumber="23">batequensis</emphasis>
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||
Groves, 2011
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||
</taxonomicName>
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||
. p. 46 (table 1), 49, figs. 5, 6.
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||
</paragraph>
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||
<paragraph id="1A09685AFFCBFFB0FF32FC38FD3EFC78" blockId="24.[112,801,688,1770]" box="[144,756,897,927]" pageId="24" pageNumber="23">
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||
<taxonomicName id="DDB613D9FFCBFFB0FF32FC38FDCFFC78" authority="(Groves)" baseAuthorityName="Groves" box="[144,517,897,927]" class="Gastropoda" family="Eocypraeidae" genus="Grovesia" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="23" phylum="Mollusca" rank="species" species="batequensis">
|
||
<emphasis id="28C2B448FFCBFFB0FF32FC38FE45FC78" box="[144,399,897,927]" italics="true" pageId="24" pageNumber="23">Grovesia batequensis</emphasis>
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(Groves)
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||
</taxonomicName>
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||
.
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||
<bibRefCitation id="7E2715ABFFCBFFB0FDADFC38FD6EFC78" author="Fehse, D." box="[527,676,899,925]" pageId="24" pageNumber="23" pagination="1 - 12" refId="ref37236" refString="Fehse, D. 2021. Zur systemstischen stellung der Eocy- praeidae F. A. Schilder 1924. Part 2. (Mollusca: Gas- tropoda: Cypraeoidea). Privately published online, Berlin, Germany 46 p., pls. 1 - 12." type="book chapter" year="2021">Fehse, 2021</bibRefCitation>
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||
. p. 44.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="52AC3BD1FFCBFFB0FF32FC1DFE4FFB76" pageId="24" pageNumber="23" type="materials_examined">
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<paragraph id="1A09685AFFCBFFB0FF32FC1DFE30FBA8" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
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<materialsCitation id="AADE6207FFCBFFB0FF32FC1DFE30FBA8" collectionCode="LACMIP, CSUN" country="Mexico" county="Mesa La Salina" location="Locality" municipality="Bateque Formation" pageId="24" pageNumber="23" specimenCode="IGM 5174" specimenCount="1" stateProvince="Baja California Sur" typeStatus="holotype">
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<emphasis id="28C2B448FFCBFFB0FF32FC1DFDC4FC25" bold="true" box="[144,526,934,960]" pageId="24" pageNumber="23">
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<typeStatus id="C50DD6F8FFCBFFB0FF32FC1DFEC3FC25" box="[144,265,934,960]" pageId="24" pageNumber="23" type="holotype">Holotype</typeStatus>
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and Type Locality—
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</emphasis>
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<specimenCode id="4A10C021FFCBFFB0FDAFFC1DFD4EFC24" box="[525,644,934,961]" collectionCode="IGM" country="Mongolia" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="24" pageNumber="23">IGM 5174</specimenCode>
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(
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<figureCitation id="828D74DFFFCBFFB0FD32FC1DFCD9FC24" box="[656,787,934,961]" captionStart="Figure 7" captionStartId="23.[112,199,200,226]" captionTargetId="figure-13@22.[139,1459,192,1920]" captionTargetPageId="22" captionText="Figure 7A–C. Cypraeidae, genus and species indeterminate, Lodo Formation (basal) (Paleocene, Seldanian Stage), Tumey Hills, Fresno County, central California. Basal (A), dorsal (B), and right (labral) (C) views, 1.3x, length 39 mm, width 25 mm, height 22 mm, of hypotype CASG 61666.06, CASG Locality 61666. D–F. Eocypraea batequensis (Groves, 2011), Bateque Formation (Eocene, “Capay Stage”), Mesa La Salina, Baja California Sur, México. Basal (D), dorsal (E), and right (labral) (F) views, 4.8x, length 9.4 mm, width 6.5 mm, height 5.1 mm, of holotype IGM 5174, LACMIP Locality 16951 [ex CSUN Locality1220b]. G–I. Eocypraea crescentensis (Groves, 2011), Crescent Formation (Eocene, “Capay Stage”), Larch Mountain, Black Hills area, Thurston County, Washington. Basal (G), dorsal (H), and left (columellar) (I) views, 3.9x, length 10.9 mm, width 7.6 mm, height 6.1 mm, of holotype LACMIP 16655.40, LACMIP Type 13646, LACMIP Locality 16655 [= LACMIP Locality 41563; ex CSUN Locality 1563]. J–L. Eocypraea jimgoederti (Groves, 2011), lower McIntosh Formation (middle Eocene, “Transition Stage” to “Tejon Stage”), Doty Hills, Lewis County, Washington. Basal (J), dorsal (K), and left (columellar) side (L) views, 3.5x, length 12.7 mm, width 8.2 mm, height 6.9 mm, of holotype LACMIP 41573.1, LACMIP Type 13647, LACMIP Locality 41573 [ex CSUN Locality 1573]. M–O. Eocypraea judithsmithae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (M), dorsal (N), and left (labral) (O) views, 1.5x, length 26.6 mm, width 18.4 mm, height 12.6 mm, of holotype LACMIP 40372.1, LACMIP Type 14940 (LACMIP Locality 40732)." figureDoi="http://doi.org/10.5281/zenodo.13869600" httpUri="https://zenodo.org/record/13869600/files/figure.png" pageId="24" pageNumber="23">Figs.7D–F</figureCitation>
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), length
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<quantity id="DD4EC5BFFFCBFFB0FF65FC72FEE2FC01" box="[199,296,969,996]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="24" pageNumber="23" unit="mm" value="9.4">9.4 mm</quantity>
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, width
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, height
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.
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<collectionCode id="7CA7F09FFFCBFFB0FD1EFC72FCEAFC06" box="[700,800,969,995]" pageId="24" pageNumber="23">LACMIP</collectionCode>
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16951 [
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<emphasis id="28C2B448FFCBFFB0FEF3FC50FEA6FBE2" box="[337,364,1003,1031]" italics="true" pageId="24" pageNumber="23">ex</emphasis>
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<collectionCode id="7CA7F09FFFCBFFB0FEDAFC56FE0BFBE2" box="[376,449,1005,1031]" country="USA" httpUri="http://grbio.org/cool/sgeh-wrgd" name="California State University, Northridge" pageId="24" pageNumber="23" type="University or college">CSUN</collectionCode>
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Locality 1220b],
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<collectingMunicipality id="FA6DF220FFCBFFB0FD16FC56FF3CFBCF" pageId="24" pageNumber="23">Bateque Formation</collectingMunicipality>
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,
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<collectingCounty id="F36810D6FFCBFFB0FEA1FBABFE0BFBCF" box="[259,449,1040,1066]" pageId="24" pageNumber="23">Mesa La Salina</collectingCounty>
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,
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,
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(
|
||
<bibRefCitation id="7E2715ABFFCBFFB0FFD9FB88FE23FBA8" author="Squires, R. L. & R. A. Demetrion" box="[123,489,1075,1101]" pageId="24" pageNumber="23" pagination="1 - 55" refId="ref43379" refString="Squires, R. L. and R. A. Demetrion. 1992. Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Contributions in Science, Natural History Museum of Los Angeles County 434: 1 - 55, figs. 1 - 144." type="journal article" year="1992">Squires and Demetrion, 1992</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="1A09685AFFCBFFB0FF32FBEDFE4FFB76" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
|
||
<emphasis id="28C2B448FFCBFFB0FF32FBEDFE9AFB95" bold="true" box="[144,336,1110,1136]" pageId="24" pageNumber="23">Occurrence—</emphasis>
|
||
Lower Eocene. “Capay Stage,”
|
||
<collectingRegion id="D872A6B8FFCBFFB0FD45FBEDFED1FB76" country="Mexico" name="Baja California Sur" pageId="24" pageNumber="23">Baja California Sur</collectingRegion>
|
||
,
|
||
<collectingCountry id="62A128CAFFCBFFB0FE84FBC2FE4BFB76" box="[294,385,1145,1171]" name="Mexico" pageId="24" pageNumber="23">México</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFCBFFB0FF32FB26FD9AFB3F" pageId="24" pageNumber="23" type="etymology">
|
||
<paragraph id="1A09685AFFCBFFB0FF32FB26FD9AFB3F" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
|
||
<emphasis id="28C2B448FFCBFFB0FF32FB26FEF2FB52" bold="true" box="[144,312,1181,1207]" pageId="24" pageNumber="23">Etymology—</emphasis>
|
||
Originally named for the Eocene Bateque Formation,
|
||
<collectingRegion id="D872A6B8FFCBFFB0FF5DFB7BFE2CFB3F" box="[255,486,1216,1242]" country="Mexico" name="Baja California Sur" pageId="24" pageNumber="23">Baja California Sur</collectingRegion>
|
||
,
|
||
<collectingCountry id="62A128CAFFCBFFB0FE53FB7BFD86FB3F" box="[497,588,1216,1242]" name="Mexico" pageId="24" pageNumber="23">México</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFCBFFB0FF32FB58FCEAF946" pageId="24" pageNumber="23" type="description">
|
||
<paragraph id="1A09685AFFCBFFB0FF32FB58FCEAF946" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
|
||
<emphasis id="28C2B448FFCBFFB0FF32FB58FE80FB18" bold="true" box="[144,330,1251,1277]" pageId="24" pageNumber="23">Description—</emphasis>
|
||
Shell small in size. Shell ovate, smooth, dorsal moderately inflated with central area flat. Spire covered. Aperture gently curved and moderately wide, opening anteriorly; denticulation well developed on both sides of aperture. Labial lip (outer lip) with ap- proximately 30 teeth,strongest ones on central part of lip. Columellar lip (outer lip) with approximately 20 teeth, becoming smaller toward exhalant canal (posterior ca- nal). Labial lip teeth slighty stronger than columellar lip teeth. Terminal ridge present on left side of columellar lip. Anterior end of shell slightly flattened out and projecting short distance. Posterior end of shell not projecting. Bor- der on basal lateral margin of shell (Groves 2011: p. 49).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFCBFFB0FF32F916FEC0F90F" pageId="24" pageNumber="23" type="discussion">
|
||
<paragraph id="1A09685AFFCBFFB0FF32F916FEC0F90F" blockId="24.[112,801,688,1770]" pageId="24" pageNumber="23">
|
||
<emphasis id="28C2B448FFCBFFB0FF32F916FEEEF922" bold="true" box="[144,292,1709,1735]" pageId="24" pageNumber="23">Remarks—</emphasis>
|
||
<taxonomicName id="DDB613D9FFCBFFB0FE86F916FDFEF922" authorityName="Groves" authorityYear="2011" box="[292,564,1707,1737]" class="Gastropoda" family="Eocypraeidae" genus="Eocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="23" phylum="Mollusca" rank="species" species="batequensis">
|
||
<emphasis id="28C2B448FFCBFFB0FE86F916FDFEF922" box="[292,564,1707,1737]" italics="true" pageId="24" pageNumber="23">Eocypraea batequensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from its
|
||
<typeStatus id="C50DD6F8FFCBFFB0FF37F96BFECCF90F" box="[149,262,1744,1770]" pageId="24" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |