164 lines
14 KiB
XML
164 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.732.21677" ID-GBIF-Dataset="05db50e8-b4e5-422a-b78f-9676d74ac319" ID-PMC="PMC5799741" ID-Pensoft-Pub="1313-2970-732-1" ID-PubMed="29416404" ID-ZBK="9995702B61464BA1BB5323DC9BA9650F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-732-1" ModsDocOrigin="ZooKeys 732" ModsDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" checkinTime="1516831538731" checkinUser="pensoft" docAuthor="Mattia, Willy De, Neiber, Marco T. & Groh, Klaus" docDate="2018" docId="90F383DC69C7D5C482060EEB77029756" docLanguage="en" docName="ZooKeys 732: 1-125" docOrigin="ZooKeys 732" docSource="http://dx.doi.org/10.3897/zookeys.732.21677" docTitle="Wollastonia inexpectata De Mattia & Groh, sp. n." docType="treatment" docUuid="AD8D733D-8248-47EF-B9D8-CF9A3F831437" docUuidSource="ZooBank" docVersion="6" lastPageNumber="62" masterDocId="FFA3FF812B32FFA8FFBAFFCDFF9AFF92" masterDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" masterLastPageNumber="125" masterPageNumber="1" pageNumber="61" updateTime="1668165370779" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mattia, Willy De</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neiber, Marco T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Groh, Klaus</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>732</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>125</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.732.21677</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.732.21677</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-732-1</mods:identifier>
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<mods:identifier type="ZBK">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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<mods:identifier type="ZooBank">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="140417769" LSID="urn:lsid:zoobank.org:act:AD8D733D-8248-47EF-B9D8-CF9A3F831437" httpUri="http://treatment.plazi.org/id/90F383DC69C7D5C482060EEB77029756" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
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<subSubSection pageId="60" pageNumber="61" type="nomenclature">
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<paragraph pageId="60" pageNumber="61">
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†
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<taxonomicName LSID="http://zoobank.org/AD8D733D-8248-47EF-B9D8-CF9A3F831437" authority="De Mattia & Groh" class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia inexpectata" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="inexpectata">Wollastonia inexpectata De Mattia & Groh</taxonomicName>
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<taxonomicNameLabel pageId="60" pageNumber="61">sp. n.</taxonomicNameLabel>
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Figs 200-202, 203
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="type material">
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<paragraph pageId="60" pageNumber="61">Type material.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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SMF 348930, holotype, from loc. typ., leg. W. De Mattia, May 23 2015 (see Fig. 200); NMWC 80.202 Acc. 55 158/1 PT as "
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Geomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geomitra oxytropis" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="oxytropis">Geomitra oxytropis</taxonomicName>
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Lowe var.
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<taxonomicName class="Gastropoda" family="Helicinidae" genus="Helicoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helicoidea var. subcarinulata" order="Cycloneritida" pageId="60" pageNumber="61" phylum="Mollusca" rank="variety" variety="subcarinulata">subcarinulata</taxonomicName>
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Woll. Porto Santo" (Figs 201-202).
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</paragraph>
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<caption pageId="60" pageNumber="61">
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<paragraph pageId="60" pageNumber="61">
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Figures 200-202. Shells of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia inexpectata" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="inexpectata">Wollastonia inexpectata</taxonomicName>
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sp. n. 200 holotype, SMF 348930 201 paratype NMWC 80.202 Acc. 55 158 202 label of the lot NMWC 80.202 Acc. 55 158. Scale bar 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="locus typicus">
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<paragraph pageId="60" pageNumber="61">Locus typicus.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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N of airport, end of the runway towards Fonte de Areia, Quaternary calcareous sand deposit,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.090275">33°05'25"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-17.349445">17°20'58"W</geoCoordinate>
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, 99 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="etymology">
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<paragraph pageId="60" pageNumber="61">Etymology.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia inexpectata" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="inexpectata">Wollastonia inexpectata</taxonomicName>
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sp. n. was unexpectedly found during intensive field researches at Fonte da Areia when looking for
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<taxonomicName lsidName="H. echinoderma" pageId="60" pageNumber="61" rank="species" species="echinoderma">H. echinoderma</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="60" pageNumber="61" type="diagnosis">
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<paragraph pageId="60" pageNumber="61">Diagnosis.</paragraph>
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<paragraph pageId="60" pageNumber="61">
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Shell large for the genus, solid and conical. Whorls rounded, with deep sutures. The
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<normalizedToken originalValue="shell’s">shell's</normalizedToken>
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surface covered with small and densely set tubercles. Body whorl with a peripheral, rounded keel. Other teleoconch whorls without visible keel. Umbilicus narrow but open. Last whorl descending toward the aperture. Aperture oval with continuous peristome.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="description">
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<paragraph pageId="60" pageNumber="61">Description of the shell.</paragraph>
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<paragraph lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
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The shell is large for the genus and rather conical, with 6.3 regularly increasing whorls. The protoconch has 1.7 smooth whorls. The teleoconch whorls are distinctly convex and separated by a slightly impressed but simple suture. The body whorl measures 64% and the penultimate whorl 14% of total shell height and is descending towards the aperture in its last 5% in an angle of 36°. It has a rounded angulation in its upper third (in relation to total shell height) that is emphasised by a narrow, only slightly concave impression below the periphery. The lower part of the body whorl (beneath the periphery in frontal view) is otherwise rather convex. The aperture is regular elliptical, measuring 49% of the total shell width and 29% of the total shell height. It is inclined to the vertical axis of the shell in an angle of 57°. The peristome is completely detached from the body whorl, expanded and distinctly reflected, especially in its basal and columellar part. The umbilicus is eccentric, closed in the early whorls, but pinhole-like in the body whorl, measuring 5% of the maximum shell diameter. The sculpture of the teleoconch consists of oblique radial ribs (10 in the penultimate quadrant of the body whorl) and numerous, small roundish tubercles (132 in the standard square basal surface of the shell). The colour
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<pageBreakToken pageId="61" pageNumber="62" start="start">is</pageBreakToken>
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only preserved as a yellowish hue on the teleoconch, a slightly darker yellowish, narrow spiral band in the middle of the base and a lighter marked keel. See Figs 200-202.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="61" pageNumber="62" type="description">
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<paragraph pageId="61" pageNumber="62">Measurements.</paragraph>
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<paragraph pageId="61" pageNumber="62">D 8.6 mm; H 6.7 mm; FW 4.6 mm; PA 41.2°; DU 0.6 mm; NT 132; NW 6.6 (n = 1). Ratio D/H 1.3; ratio FW/H 0.7.</paragraph>
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</subSubSection>
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<subSubSection pageId="61" pageNumber="62" type="distribution">
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<paragraph pageId="61" pageNumber="62">Distribution.</paragraph>
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<paragraph pageId="61" pageNumber="62">The species is only known from the type locality. A second specimen is known from another unidentified locality in Porto Santo and housed at the NMW. The distribution is shown in Fig. 203.</paragraph>
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<caption pageId="61" pageNumber="62">
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<paragraph pageId="61" pageNumber="62">
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Figure 203. Distribution of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia inexpectata" order="Stylommatophora" pageId="61" pageNumber="62" phylum="Mollusca" rank="species" species="inexpectata">Wollastonia inexpectata</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="61" pageNumber="62" type="comparisons">
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<paragraph pageId="61" pageNumber="62">Comparison and comments.</paragraph>
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<paragraph pageId="61" pageNumber="62">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia inexpectata" order="Stylommatophora" pageId="61" pageNumber="62" phylum="Mollusca" rank="species" species="inexpectata">Wollastonia inexpectata</taxonomicName>
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sp. n. is superficially similar to
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<taxonomicName lsidName="H. echinoderma" pageId="61" pageNumber="62" rank="species" species="echinoderma">H. echinoderma</taxonomicName>
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, or
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<taxonomicName lsidName="W. vermetiformis" pageId="61" pageNumber="62" rank="species" species="vermetiformis">W. vermetiformis</taxonomicName>
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and
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<taxonomicName lsidName="W. falknerorum" pageId="61" pageNumber="62" rank="species" species="falknerorum">W. falknerorum</taxonomicName>
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sp. n. but is readily distinguishable from these species by the lack of a second keel, its much finer sculpture, narrower umbilicus and regular conical form. From
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<taxonomicName lsidName="H. echinoderma" pageId="61" pageNumber="62" rank="species" species="echinoderma">H. echinoderma</taxonomicName>
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it can also be distinguished by its regular convex and not stepped whorls and the regular elliptical rather than oblique ovate aperture. From the similarly sized
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<taxonomicName lsidName="W. subcarinulata" pageId="61" pageNumber="62" rank="species" species="subcarinulata">W. subcarinulata</taxonomicName>
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it differs in the narrower umbilicus, the presence of a rounded keel, the finer granulation and the higher shell in relation to its width.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="61" pageNumber="62" type="taxonomic remarks">
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<paragraph pageId="61" pageNumber="62">Taxonomic remarks.</paragraph>
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<paragraph pageId="61" pageNumber="62">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia inexpectata" order="Stylommatophora" pageId="61" pageNumber="62" phylum="Mollusca" rank="species" species="inexpectata">Wollastonia inexpectata</taxonomicName>
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sp. n. is included in the genus
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia" order="Stylommatophora" pageId="61" pageNumber="62" phylum="Mollusca" rank="genus">Wollastonia</taxonomicName>
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because it is similar to
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<taxonomicName lsidName="W. oxytropis" pageId="61" pageNumber="62" rank="species" species="oxytropis">W. oxytropis</taxonomicName>
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in size and surface sculpture. It is noteworthy that the similarly large-sized
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella echinoderma" order="Stylommatophora" pageId="61" pageNumber="62" phylum="Mollusca" rank="species" species="echinoderma">Hystricella echinoderma</taxonomicName>
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most probably also originates from the Quaternary deposits in the north of the island, and likewise is extremely rare there.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="61" pageNumber="62" type="status and conservation">
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<paragraph pageId="61" pageNumber="62">Status and conservation.</paragraph>
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<paragraph pageId="61" pageNumber="62">
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Extinct before the
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<normalizedToken originalValue="islands’">islands'</normalizedToken>
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scientific exploration in the 19th century, possibly already before human settlement.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |