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<document id="4FA6A2A690E9226D2168AC28A8059D5D" ID-CLB-Dataset="189446" ID-DOI="10.11646/zootaxa.5209.4.4" ID-GBIF-Dataset="fbc4017f-ae71-4f97-a916-a9ea25d41548" ID-ISSN="1175-5326" ID-Zenodo-Dep="7333850" ID-ZooBank="A151ED2E-9A3E-46B2-A94D-202C4B06A55F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668767965713" checkinUser="plazi" docAuthor="Gupta, Ankita, Achterberg, Cornelis Van, Pattar, Rohit &amp; Minz, Kriti Arpana" docDate="2022" docId="904287A89A07FF8146D5F8A9FEF2FD7A" docLanguage="en" docName="zootaxa.5209.4.4.pdf" docOrigin="Zootaxa 5209 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leiophron crassivena Gupta &amp; van Achterberg 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="460" masterDocId="6C7BFFD09A04FF844642FE47FFE8FF8A" masterDocTitle="On the occurrence of a Palaearctic species of the genus Aphaereta Foerster (Braconidae, Alysiinae) and description of a new species of the genus Leiophron Nees (Braconidae, Euphorinae) from central India" masterLastPageNumber="462" masterPageNumber="455" pageNumber="458" updateTime="1699411788882" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="693807E07A3104988AC590EF755679B1">On the occurrence of a Palaearctic species of the genus Aphaereta Foerster (Braconidae, Alysiinae) and description of a new species of the genus Leiophron Nees (Braconidae, Euphorinae) from central India</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="B1A0CA371DE95FEEF3FD23136193B421">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4C6CA20D4EEFA69A315C82BF67AD645">Gupta, Ankita</mods:namePart>
</mods:name>
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<mods:roleTerm id="E79BE935FDACBD8067807DE4C5901289">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39B5B5ED47F8793B1CD49685AD1C3EFF">Achterberg, Cornelis Van</mods:namePart>
<mods:nameIdentifier id="D75A1E8D266DD0D4686D714BD4E889DF" type="ORCID">0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation id="037F42B651129BB8938B342045373D81">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. https: // orcid. org / 0000 - 0002 - 6495 - 4853</mods:affiliation>
</mods:name>
<mods:name id="14CDFFF6B372655B2E3B092D1DCC921C" type="personal">
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<mods:roleTerm id="47BD245F205C8B1BCD405CE8E69E8D3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B61C42C95ACD81940F7D3A7665CF187">Pattar, Rohit</mods:namePart>
<mods:nameIdentifier id="A86FDCEB53752D41BE211E25963B15AD" type="ORCID">0000-0002-2243-3777</mods:nameIdentifier>
<mods:affiliation id="CDCA6AEF24F6FB9690673729BD333339">https: // orcid. org / 0000 - 0002 - 2243 - 3777</mods:affiliation>
</mods:name>
<mods:name id="A9DB9A08B4208170779F44B1B6E2D30C" type="personal">
<mods:role id="F533E2F9F232C61D3B05BB9FA9287FC6">
<mods:roleTerm id="E68F0860FE8B24FC51479B62C6A8683D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="85BE4787521285112912C1A2B791B391">Minz, Kriti Arpana</mods:namePart>
<mods:nameIdentifier id="412B8294ADAC4D27B0019B193E662FA2" type="ORCID">0000-0002-9687-1251</mods:nameIdentifier>
<mods:affiliation id="20118311CAB64005CFFB242AF2C091FD">https: // orcid. org / 0000 - 0002 - 9687 - 1251</mods:affiliation>
</mods:name>
<mods:typeOfResource id="698FB4D41478AF44BD8B40D2782DBA83">text</mods:typeOfResource>
<mods:relatedItem id="9F30E2A8C1E9DEC21BA2F10C6BA24651" type="host">
<mods:titleInfo id="379B8F4A7F6805D82BD3CBCA5E4148B8">
<mods:title id="9C757EA7782016BA7C0B780898502F53">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8CA9E454F73E57AAC53590B70B9AAD78">
<mods:date id="2AB375BE0943E4F5F86F7D6CD0B80A9C">2022</mods:date>
<mods:detail id="AC924F927E0865C9CAA04F1AC6DF566A" type="pubDate">
<mods:number id="D97937476C3A997368AF9D4E723BD49C">2022-11-18</mods:number>
</mods:detail>
<mods:detail id="3F7042D3DF73E585609AB59F00F269DF" type="volume">
<mods:number id="517B59FBAA35B8FA362E2C577358F24D">5209</mods:number>
</mods:detail>
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<mods:number id="1B1CBBF0C6C73A64450F2322705379A8">4</mods:number>
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<mods:classification id="B980042DDB93D068F765D4742F08038D">journal article</mods:classification>
<mods:identifier id="CC219F424449163DCAE4F90AC0FB7143" type="CLB-Dataset">189446</mods:identifier>
<mods:identifier id="5A60B22EC62858DE85755266EA73448F" type="DOI">10.11646/zootaxa.5209.4.4</mods:identifier>
<mods:identifier id="55D71C33DCB3E8D26B9E5F934D5D4215" type="GBIF-Dataset">fbc4017f-ae71-4f97-a916-a9ea25d41548</mods:identifier>
<mods:identifier id="292EC0322987F5A7A7572FB90D96A3EE" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="904287A89A07FF8146D5F8A9FEF2FD7A" ID-DOI="http://doi.org/10.5281/zenodo.7333852" ID-GBIF-Taxon="205097994" ID-Zenodo-Dep="7333852" LSID="urn:lsid:plazi:treatment:904287A89A07FF8146D5F8A9FEF2FD7A" httpUri="http://treatment.plazi.org/id/904287A89A07FF8146D5F8A9FEF2FD7A" lastPageId="5" lastPageNumber="460" pageId="3" pageNumber="458">
<subSubSection id="50F165359A07FF8746D5F8A9FCABF882" box="[151,835,1774,1801]" pageId="3" pageNumber="458" type="nomenclature">
<paragraph id="185436BE9A07FF8746D5F8A9FCABF882" blockId="3.[151,835,1774,1837]" box="[151,835,1774,1801]" pageId="3" pageNumber="458">
<heading id="431C81D29A07FF8746D5F8A9FCABF882" bold="true" box="[151,835,1774,1801]" fontSize="11" level="1" pageId="3" pageNumber="458" reason="1">
<emphasis id="2A9FEAAC9A07FF8746D5F8A9FCABF882" bold="true" box="[151,835,1774,1801]" pageId="3" pageNumber="458">
<taxonomicName id="DFEB4D3D9A07FF8746D5F8A9FD35F882" authority="Gupta &amp; van Achterberg" authorityName="Gupta &amp; van Achterberg" authorityYear="2022" box="[151,733,1774,1801]" class="Insecta" family="Braconidae" genus="Leiophron" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="458" phylum="Arthropoda" rank="species" species="crassivena" status="sp. nov.">
<emphasis id="2A9FEAAC9A07FF8746D5F8A9FE75F882" bold="true" box="[151,413,1774,1801]" italics="true" pageId="3" pageNumber="458">Leiophron crassivena</emphasis>
Gupta &amp; van Achterberg
</taxonomicName>
<taxonomicNameLabel id="31AC57D79A07FF8744A7F8A9FCABF882" box="[741,835,1774,1800]" pageId="3" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="50F165359A07FF8746D5F955FF10F8A7" box="[151,248,1810,1837]" pageId="3" pageNumber="458" type="description">
<paragraph id="185436BE9A07FF8746D5F955FF10F8A7" blockId="3.[151,835,1774,1837]" box="[151,248,1810,1837]" pageId="3" pageNumber="458">
<figureCitation id="80D02A3B9A07FF8746D5F955FF10F8A7" box="[151,248,1810,1837]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="4.[152,266,1557,1583]" captionStartId-1="5.[152,266,2006,2032]" captionStartId-2="6.[152,266,1866,1892]" captionTargetBox-0="[187,1415,214,1463]" captionTargetBox-1="[151,1436,774,1982]" captionTargetBox-2="[239,1378,227,1765]" captionTargetId-0="figure-23@4.[153,1435,179,1533]" captionTargetId-1="figure-319@5.[151,1436,774,1982]" captionTargetId-2="figure-23@6.[151,1436,179,1842]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 3. Leiophron crassivena Gupta &amp; van Achterberg sp. nov., holotype (female)." captionText-1="FIGURE 4. Leiophron crassivena Gupta &amp; van Achterberg sp. nov., male." captionText-2="FIGURE 5. Leiophron crassivena Gupta &amp; van Achterberg sp. nov. A—Head in frontal aspect; B—Vertex; C— Mesosoma; D—Metasoma; E—Wings." figureDoi-0="http://doi.org/10.5281/zenodo.7333860" figureDoi-1="http://doi.org/10.5281/zenodo.7333862" figureDoi-2="http://doi.org/10.5281/zenodo.7333864" httpUri-0="https://zenodo.org/record/7333860/files/figure.png" httpUri-1="https://zenodo.org/record/7333862/files/figure.png" httpUri-2="https://zenodo.org/record/7333864/files/figure.png" pageId="3" pageNumber="458">Figs 3–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="50F165359A07FF8146D5F91DFB68FE26" lastPageId="5" lastPageNumber="460" pageId="3" pageNumber="458" type="materials_examined">
<paragraph id="185436BE9A07FF8746D5F91DFA9DF836" blockId="3.[151,1436,1882,2017]" pageId="3" pageNumber="458">
<emphasis id="2A9FEAAC9A07FF8746D5F91DFEABF8FE" bold="true" box="[151,323,1882,1908]" pageId="3" pageNumber="458">Type material.</emphasis>
<materialsCitation id="A8833CE39A07FF87470AF91DFBECF812" ID-GBIF-Occurrence="3967670303" collectingDate="2022-03-25" collectingDateMax="2022-03-30" collectingDateMin="2022-03-25" collectionCode="NBAIR" collectorName="Kriti A. Minz." country="India" latitude="22.809206" location="Mainpat" longLatPrecision="1" longitude="83.29414" pageId="3" pageNumber="458" specimenCount="1" specimenCount-female="1" stateProvince="Chhattisgarh" typeStatus="holotype">
<emphasis id="2A9FEAAC9A07FF87470AF91DFE54F8FE" bold="true" box="[328,444,1882,1908]" pageId="3" pageNumber="458">
<typeStatus id="C750881C9A07FF87470AF91DFE5FF8FE" box="[328,439,1882,1908]" pageId="3" pageNumber="458" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="0EEDFD379A07FF874783F91CFE3FF8FF" box="[449,471,1883,1909]" pageId="3" pageNumber="458" type="female">♀</specimenCount>
;
<collectingCountry id="60FC762E9A07FF8747A1F91DFDF7F8FE" box="[483,543,1882,1908]" name="India" pageId="3" pageNumber="458">India</collectingCountry>
,
<collectingRegion id="DA2FF85C9A07FF87446BF91DFD53F8FE" box="[553,699,1882,1908]" country="India" name="Chhattisgarh" pageId="3" pageNumber="458">Chhattisgarh</collectingRegion>
,
<location id="1D3460659A07FF874486F91DFCCCF8FE" LSID="urn:lsid:plazi:treatment:904287A89A07FF8146D5F8A9FEF2FD7A:1D3460659A07FF874486F91DFCCCF8FE" box="[708,804,1882,1908]" country="India" latitude="22.809206" longLatPrecision="1" longitude="83.29414" name="Mainpat" pageId="3" pageNumber="458" stateProvince="Chhattisgarh">Mainpat</location>
,
<geoCoordinate id="7DDF50799A07FF87456CF91BFC05F8FF" box="[814,1005,1883,1911]" degrees="22" direction="north" minutes="48" orientation="latitude" pageId="3" pageNumber="458" precision="1" seconds="33.146" value="22.809206">22°48′33.146′′ N</geoCoordinate>
,
<geoCoordinate id="7DDF50799A07FF8745B4F91DFB59F8FF" box="[1014,1201,1882,1911]" degrees="83" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="458" precision="1" seconds="38.919" value="83.29414">83°17′38.919′′ E</geoCoordinate>
,
<collectingMethod id="C1AA4EA99A07FF8742F9F91DFA80F8FF" box="[1211,1384,1882,1909]" pageId="3" pageNumber="458">yellow pan trap</collectingMethod>
,
<date id="6C55107E9A07FF874330F91DFEFCF812" pageId="3" pageNumber="458" value="2022-03-25" valueMax="2022-03-30" valueMin="2022-03-25">
<collectingDate id="7C11E9969A07FF874330F91DFEFCF812" pageId="3" pageNumber="458" value="2022-03-25" valueMax="2022-03-30" valueMin="2022-03-25">25– 30.iii.2022</collectingDate>
</date>
; leg
<collectorName id="B51E53689A07FF87470AF939FE0FF812" box="[328,487,1918,1945]" pageId="3" pageNumber="458">Kriti A. Minz.</collectorName>
code—
<collectionCode id="7EFAAE7B9A07FF87447CF939FD7DF813" box="[574,661,1918,1945]" pageId="3" pageNumber="458">NBAIR</collectionCode>
/
<collectorName id="B51E53689A07FF8744DFF939FD39F813" box="[669,721,1918,1945]" pageId="3" pageNumber="458">Brac</collectorName>
/
<collectorName id="B51E53689A07FF87449BF939FCFCF813" box="[729,788,1918,1945]" pageId="3" pageNumber="458">Euph</collectorName>
/
<collectorName id="B51E53689A07FF87455FF939FCB8F813" box="[797,848,1918,1945]" pageId="3" pageNumber="458">Leio</collectorName>
/25322H (NIM)
</materialsCitation>
.
<materialsCitation id="A8833CE39A07FF87424DF938FA99F836" ID-GBIF-Occurrence="3967670301" collectingDate="2022-03-25" collectingDateMax="2022-03-30" collectingDateMin="2022-03-25" collectionCode="NBAIR" collectorName="Kriti A. Minz." country="India" latitude="22.809206" location="Mainpat" longLatPrecision="1" longitude="83.29414" pageId="3" pageNumber="458" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chhattisgarh" typeStatus="paratype">
<typeStatus id="C750881C9A07FF87424DF938FB9CF813" box="[1039,1140,1919,1945]" pageId="3" pageNumber="458" type="paratype">Paratype</typeStatus>
,
<specimenCount id="0EEDFD379A07FF87423CF938FB12F812" box="[1150,1274,1918,1945]" pageId="3" pageNumber="458" type="female">one female</specimenCount>
and
<specimenCount id="0EEDFD379A07FF874372F938FA70F812" box="[1328,1432,1918,1945]" pageId="3" pageNumber="458" type="male">one male</specimenCount>
, same data as holotype, code—
<collectionCode id="7EFAAE7B9A07FF8747ACF9E5FDAEF837" box="[494,582,1954,1981]" pageId="3" pageNumber="458">NBAIR</collectionCode>
/Brac/Euph/Leio/25322P1 and
<collectionCode id="7EFAAE7B9A07FF8745E2F9E5FC1FF837" box="[928,1015,1954,1981]" pageId="3" pageNumber="458">NBAIR</collectionCode>
/Brac/Euph/Leio/25322P2 (NIM)
</materialsCitation>
.
</paragraph>
<paragraph id="185436BE9A07FF874685F981FB1CF86B" blockId="3.[151,1436,1882,2017]" box="[199,1268,1990,2017]" pageId="3" pageNumber="458">
<emphasis id="2A9FEAAC9A07FF874685F981FED3F86A" bold="true" box="[199,315,1990,2016]" pageId="3" pageNumber="458">
<typeStatus id="C750881C9A07FF874685F981FEDEF86A" box="[199,310,1990,2016]" pageId="3" pageNumber="458" type="holotype">Holotype</typeStatus>
.
</emphasis>
Body length
<quantity id="DF139B5B9A07FF874791F981FDC4F86B" box="[467,556,1990,2017]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="458" unit="mm" value="3.2">3.2 mm</quantity>
; fore wing length
<quantity id="DF139B5B9A07FF8744B5F981FCB8F86B" box="[759,848,1990,2017]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="458" unit="mm" value="2.3">2.3 mm</quantity>
; exserted part of ovipositor
<quantity id="DF139B5B9A07FF8742CBF981FB18F86B" box="[1161,1264,1990,2017]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="458" unit="mm" value="0.15">0.15 mm</quantity>
.
</paragraph>
<caption id="4C9466369A00FF8046DAF852FB90F9A5" ID-DOI="http://doi.org/10.5281/zenodo.7333860" ID-Zenodo-Dep="7333860" box="[152,1144,1557,1584]" httpUri="https://zenodo.org/record/7333860/files/figure.png" pageId="4" pageNumber="459" startId="4.[152,266,1557,1583]" targetBox="[187,1415,214,1463]" targetPageId="4">
<paragraph id="185436BE9A00FF8046DAF852FB90F9A5" blockId="4.[152,1144,1557,1584]" box="[152,1144,1557,1584]" pageId="4" pageNumber="459">
<emphasis id="2A9FEAAC9A00FF8046DAF852FECEF9A5" bold="true" box="[152,294,1557,1583]" pageId="4" pageNumber="459">FIGURE 3.</emphasis>
<taxonomicName id="DFEB4D3D9A00FF80476FF852FCD1F9A5" authority="Gupta &amp; van Achterberg" authorityName="Gupta &amp; van Achterberg" authorityYear="2022" box="[301,825,1557,1584]" class="Insecta" family="Braconidae" genus="Leiophron" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="459" phylum="Arthropoda" rank="species" species="crassivena" status="sp. nov.">
<emphasis id="2A9FEAAC9A00FF80476FF852FDC8F9A5" box="[301,544,1557,1583]" italics="true" pageId="4" pageNumber="459">Leiophron crassivena</emphasis>
Gupta &amp; van Achterberg
</taxonomicName>
<emphasis id="2A9FEAAC9A00FF804503F852FC72F9A5" bold="true" box="[833,922,1557,1583]" pageId="4" pageNumber="459">
<taxonomicNameLabel id="31AC57D79A00FF804503F852FC72F9A5" box="[833,922,1557,1583]" pageId="4" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (female).
</paragraph>
</caption>
<paragraph id="185436BE9A00FF804685F81AFD28F969" blockId="4.[151,1436,1629,1980]" pageId="4" pageNumber="459">
<emphasis id="2A9FEAAC9A00FF804685F81AFEC8F9FD" bold="true" box="[199,288,1629,1655]" pageId="4" pageNumber="459">Colour.</emphasis>
Body mainly brownish (
<figureCitation id="80D02A3B9A00FF804470F81AFD9FF9FD" box="[562,631,1629,1655]" captionStart="FIGURE 3" captionStartId="4.[152,266,1557,1583]" captionTargetBox="[187,1415,214,1463]" captionTargetId="figure-23@4.[153,1435,179,1533]" captionTargetPageId="4" captionText="FIGURE 3. Leiophron crassivena Gupta &amp; van Achterberg sp. nov., holotype (female)." figureDoi="http://doi.org/10.5281/zenodo.7333860" httpUri="https://zenodo.org/record/7333860/files/figure.png" pageId="4" pageNumber="459">Figs 3</figureCitation>
&amp;
<figureCitation id="80D02A3B9A00FF8044DBF81AFD42F9FD" box="[665,682,1629,1655]" captionStart="FIGURE 4" captionStartId="5.[152,266,2006,2032]" captionTargetBox="[151,1436,774,1982]" captionTargetId="figure-319@5.[151,1436,774,1982]" captionTargetPageId="5" captionText="FIGURE 4. Leiophron crassivena Gupta &amp; van Achterberg sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.7333862" httpUri="https://zenodo.org/record/7333862/files/figure.png" pageId="4" pageNumber="459">4</figureCitation>
); antennomeres yellowish but 3–4 apical antennomeres dark brown; legs in general yellowish with comparatively paler fore and mid legs; fore wings prominently infuscated, fore wing with pterostigma dark brown in apical half and remainder pale yellowish; first tergite and majority of the second tergite yellowish brown, following tergites black.
</paragraph>
<paragraph id="185436BE9A00FF804685F8AAFCEAF831" blockId="4.[151,1436,1629,1980]" pageId="4" pageNumber="459">
<emphasis id="2A9FEAAC9A00FF804685F8AAFEE6F88D" bold="true" box="[199,270,1773,1799]" pageId="4" pageNumber="459">Head.</emphasis>
Antenna with 15 antennomeres; F1 3.8 × as long as wide and 1.1 × F2; penultimate antennomere 1.4 × as long as wide; head rather square (
<figureCitation id="80D02A3B9A00FF804453F956FD83F8A6" box="[529,619,1809,1836]" captionStart="FIGURE 5" captionStartId="6.[152,266,1866,1892]" captionTargetBox="[239,1378,227,1765]" captionTargetId="figure-23@6.[151,1436,179,1842]" captionTargetPageId="6" captionText="FIGURE 5. Leiophron crassivena Gupta &amp; van Achterberg sp. nov. A—Head in frontal aspect; B—Vertex; C— Mesosoma; D—Metasoma; E—Wings." figureDoi="http://doi.org/10.5281/zenodo.7333864" httpUri="https://zenodo.org/record/7333864/files/figure.png" pageId="4" pageNumber="459">Fig. 5A</figureCitation>
), ca. 1.4 × as wide as long in dorsal view, 1.3× as wide as long in frontal view; temple smooth; eye length 1.6 × temple in dorsal view (
<figureCitation id="80D02A3B9A00FF804507F972FC75F8DA" box="[837,925,1845,1872]" captionStart="FIGURE 5" captionStartId="6.[152,266,1866,1892]" captionTargetBox="[239,1378,227,1765]" captionTargetId="figure-23@6.[151,1436,179,1842]" captionTargetPageId="6" captionText="FIGURE 5. Leiophron crassivena Gupta &amp; van Achterberg sp. nov. A—Head in frontal aspect; B—Vertex; C— Mesosoma; D—Metasoma; E—Wings." figureDoi="http://doi.org/10.5281/zenodo.7333864" httpUri="https://zenodo.org/record/7333864/files/figure.png" pageId="4" pageNumber="459">Fig. 5B</figureCitation>
); POL:OOL: OD = 14.3: 12.4: 4.5; POL 1.1 × OOL; malar suture present, malar space subequal to 0.7 × basal width of mandible; occipital carina ventrally present; face smooth and setose, clypeus smooth and intertentorial line 3.2 × tentorio-ocular line; vertex smooth around ocelli, with shallow punctures near temple and occiput.
</paragraph>
<paragraph id="185436BE9A01FF814685FED1FAA5FECA" blockId="5.[151,1436,150,752]" pageId="5" pageNumber="460">
<emphasis id="2A9FEAAC9A01FF814685FED1FEA4FF3A" bold="true" box="[199,332,150,176]" pageId="5" pageNumber="460">Mesosoma.</emphasis>
Mesosoma 1.8 × as long as wide in dorsal view, notauli entirely absent, median and lateral lobes of mesoscutum smooth (
<figureCitation id="80D02A3B9A01FF8147CFFEFDFE0EFF5F" box="[397,486,186,213]" captionStart="FIGURE 5" captionStartId="6.[152,266,1866,1892]" captionTargetBox="[239,1378,227,1765]" captionTargetId="figure-23@6.[151,1436,179,1842]" captionTargetPageId="6" captionText="FIGURE 5. Leiophron crassivena Gupta &amp; van Achterberg sp. nov. A—Head in frontal aspect; B—Vertex; C— Mesosoma; D—Metasoma; E—Wings." figureDoi="http://doi.org/10.5281/zenodo.7333864" httpUri="https://zenodo.org/record/7333864/files/figure.png" pageId="5" pageNumber="460">Fig. 5C</figureCitation>
). Propodeum irregularly reticulate rugose with no median carina. Fore wing 3.3 × as long as wide (
<figureCitation id="80D02A3B9A01FF814776FE99FE62FF73" box="[308,394,222,249]" captionStart="FIGURE 5" captionStartId="6.[152,266,1866,1892]" captionTargetBox="[239,1378,227,1765]" captionTargetId="figure-23@6.[151,1436,179,1842]" captionTargetPageId="6" captionText="FIGURE 5. Leiophron crassivena Gupta &amp; van Achterberg sp. nov. A—Head in frontal aspect; B—Vertex; C— Mesosoma; D—Metasoma; E—Wings." figureDoi="http://doi.org/10.5281/zenodo.7333864" httpUri="https://zenodo.org/record/7333864/files/figure.png" pageId="5" pageNumber="460">Fig. 5E</figureCitation>
); pterostigma 2.8 × as long as wide; 1-
<collectionCode id="7EFAAE7B9A01FF814578FE99FCBBFF72" box="[826,851,222,248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="460">R</collectionCode>
1 0.35–0.37 × length of pterostigma; 1-M thickened; 1-CU1, 2-CU1 present; r+3-SR+SR1 well developed; 1-SR+M weakly developed; basal cell glabrous and subbasal cell largely so. Hind femur, hind tibia and hind basitarsus 4.9, 8.0, 8.9 × as long as their width, respectively.
</paragraph>
<paragraph id="185436BE9A01FF814685FF0DFB68FE26" blockId="5.[151,1436,150,752]" pageId="5" pageNumber="460">
<emphasis id="2A9FEAAC9A01FF814685FF0DFEA3FEEE" bold="true" box="[199,331,330,356]" pageId="5" pageNumber="460">Metasoma.</emphasis>
Metasoma 2.5 × its width in dorsal view and 2.7 × its height in lateral view; first tergite longitudinally striate (
<figureCitation id="80D02A3B9A01FF81476BFF29FE6AFE03" box="[297,386,366,393]" captionStart="FIGURE 5" captionStartId="6.[152,266,1866,1892]" captionTargetBox="[239,1378,227,1765]" captionTargetId="figure-23@6.[151,1436,179,1842]" captionTargetPageId="6" captionText="FIGURE 5. Leiophron crassivena Gupta &amp; van Achterberg sp. nov. A—Head in frontal aspect; B—Vertex; C— Mesosoma; D—Metasoma; E—Wings." figureDoi="http://doi.org/10.5281/zenodo.7333864" httpUri="https://zenodo.org/record/7333864/files/figure.png" pageId="5" pageNumber="460">Fig. 5D</figureCitation>
), 3.9 × longer than its apical width; spiracles situated at middle of tergite and ventral one third of
<collectionCode id="7EFAAE7B9A01FF8146F7FFD5FF22FE26" box="[181,202,402,428]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="460">T</collectionCode>
1 closed in middle;
<collectionCode id="7EFAAE7B9A01FF8147E0FFD5FE5CFE26" box="[418,436,402,428]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="460">T</collectionCode>
2 smooth, 0.4 × length of metasoma, remaining tergites smooth.
</paragraph>
</subSubSection>
<subSubSection id="50F165359A01FF814685FFF1FA96FE5B" box="[199,1406,438,465]" pageId="5" pageNumber="460" type="description">
<paragraph id="185436BE9A01FF814685FFF1FA96FE5B" blockId="5.[151,1436,150,752]" box="[199,1406,438,465]" pageId="5" pageNumber="460">
<emphasis id="2A9FEAAC9A01FF814685FFF1FED7FE5A" bold="true" box="[199,319,438,464]" pageId="5" pageNumber="460">Variation.</emphasis>
The male
<typeStatus id="C750881C9A01FF8147F7FFF0FDFDFE5B" box="[437,533,439,465]" pageId="5" pageNumber="460" type="paratype">paratype</typeStatus>
(
<figureCitation id="80D02A3B9A01FF814467FFF1FD80FE5A" box="[549,616,438,464]" captionStart="FIGURE 4" captionStartId="5.[152,266,2006,2032]" captionTargetBox="[151,1436,774,1982]" captionTargetId="figure-319@5.[151,1436,774,1982]" captionTargetPageId="5" captionText="FIGURE 4. Leiophron crassivena Gupta &amp; van Achterberg sp. nov., male." figureDoi="http://doi.org/10.5281/zenodo.7333862" httpUri="https://zenodo.org/record/7333862/files/figure.png" pageId="5" pageNumber="460">Fig. 4</figureCitation>
) is very similar to the
<typeStatus id="C750881C9A01FF814527FFF1FC2FFE5A" box="[869,967,438,464]" pageId="5" pageNumber="460" type="holotype">holotype</typeStatus>
but differs in having 16 antennomeres.
</paragraph>
</subSubSection>
<subSubSection id="50F165359A01FF814685FF9DFEBAFD92" pageId="5" pageNumber="460" type="etymology">
<paragraph id="185436BE9A01FF814685FF9DFEBAFD92" blockId="5.[151,1436,150,752]" pageId="5" pageNumber="460">
<emphasis id="2A9FEAAC9A01FF814685FF9DFEA6FE7E" bold="true" box="[199,334,474,500]" pageId="5" pageNumber="460">Etymology.</emphasis>
Named after the thickened vein 1-M of the fore wing: “crassus” is Latin for thickened and “vena” is Latin for vein.
</paragraph>
</subSubSection>
<subSubSection id="50F165359A01FF814685FC65FEF2FD7A" pageId="5" pageNumber="460" type="discussion">
<paragraph id="185436BE9A01FF814685FC65FEF2FD7A" blockId="5.[151,1436,150,752]" pageId="5" pageNumber="460">
<emphasis id="2A9FEAAC9A01FF814685FC65FED3FDB6" bold="true" box="[199,315,546,572]" pageId="5" pageNumber="460">Remarks.</emphasis>
This species comes close to
<taxonomicName id="DFEB4D3D9A01FF8144C4FC64FD03FDB6" box="[646,747,546,572]" class="Insecta" family="Braconidae" genus="Leiophron" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="460" phylum="Arthropoda" rank="species" species="hayati">
<emphasis id="2A9FEAAC9A01FF8144C4FC64FD03FDB6" box="[646,747,546,572]" italics="true" pageId="5" pageNumber="460">L. hayati</emphasis>
</taxonomicName>
by having the length of the malar space and the basal width of the mandible subequal, the face smooth, the colour pattern similar and the notauli entirely absent (
<bibRefCitation id="7C7A4B4F9A01FF8142BCFC01FF3CFD0E" author="Shamim, M. &amp; Ahmad, Z. &amp; Samiuddin, A." pageId="5" pageNumber="460" pagination="5 - 22" refId="ref2661" refString="Shamim, M., Ahmad, Z. &amp; Samiuddin, A. (2009) Taxonomic studies of Indian Leiophron subgen. Leiophron (Hymenoptera: Braconidae: Euphorinae), with the description of seven new species. Folia Heyrovskyana, Series A, 17, 1, 5 - 22." type="journal article" year="2009">
Shamim
<emphasis id="2A9FEAAC9A01FF814326FC00FA7DFDEA" box="[1380,1429,582,608]" italics="true" pageId="5" pageNumber="460">et al</emphasis>
. 2009
</bibRefCitation>
). However, it can be differentiated with the following set of characters: length of metasoma 2.6 × its height (4.2 × its height in
<taxonomicName id="DFEB4D3D9A01FF814736FCC8FE32FD22" box="[372,474,654,680]" class="Insecta" family="Braconidae" genus="Leiophron" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="460" phylum="Arthropoda" rank="species" species="hayati">
<emphasis id="2A9FEAAC9A01FF814736FCC8FE32FD22" box="[372,474,654,680]" italics="true" pageId="5" pageNumber="460">L. hayati</emphasis>
</taxonomicName>
); hind femur 4.9 × as long as wide (3.0 × as long as wide); T1 3.9 × longer than its apical width (3.3 × longer than its apical width) and T2 0.4 × as long as metasoma (more than 0.5 × as long as metasoma).
</paragraph>
</subSubSection>
</treatment>
</document>
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