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<document id="0441F7F38F7D2FD5A041F0B2C2CFC9CF" ID-CLB-Dataset="50297" ID-DOI="10.5281/zenodo.174029" ID-GBIF-Dataset="d9d8d147-e8fc-4014-8af2-aa2fd28c20df" ID-ISSN="1175­5326" ID-Zenodo-Dep="174029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790304226" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2006" docId="7F54080DFFE6FF89D56AD845FA80FE5B" docLanguage="en" docName="zt01325p384.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Laena moguntia Schawaller, 2006, n. sp." docType="treatment" docVersion="11" lastPageNumber="381" masterDocId="836D7075FFEDFF84D462DD29FF82FFF5" masterDocTitle="New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *" masterLastPageNumber="384" masterPageNumber="369" pageNumber="380" updateTime="1732608391595" updateUser="ExternalLinkService">
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<mods:titleInfo id="C5CF8BBB33F8D2D77D270918B99BABAC">
<mods:title id="DD9E48BE4B668CBB2B223CD0900C2055">New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *</mods:title>
</mods:titleInfo>
<mods:name id="A905271D72620617410178D7907D5DD3" type="personal">
<mods:role id="C33E2A2436DE31963B57CACB7DA67C06">
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<mods:namePart id="F73E427347B05ADF6138EC29FE9F42AF">Schawaller, Wolfgang</mods:namePart>
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<mods:title id="7850F7A0883CEC083DA5D88DAEE7F24B">Zootaxa</mods:title>
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<mods:part id="A7A8596C5295642818215B35D7D45B37">
<mods:date id="CD2667B6A12511FEC6F64367774CBBEF">2006</mods:date>
<mods:detail id="2B5A05C785D13CFD22608528281807EB" type="volume">
<mods:number id="85349B832552432AEE671E3960F2526F">1325</mods:number>
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<mods:classification id="D52D33550165A513AB060D89939A9D40">journal article</mods:classification>
<mods:identifier id="5358EDB4DB5561C1792744C11EE0AC04" type="CLB-Dataset">50297</mods:identifier>
<mods:identifier id="4DB479327999924CA709E758AD4C2A32" type="DOI">10.5281/zenodo.174029</mods:identifier>
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<treatment id="7F54080DFFE6FF89D56AD845FA80FE5B" ID-DOI="http://doi.org/10.5281/zenodo.6258221" ID-GBIF-Taxon="119350254" ID-Zenodo-Dep="6258221" LSID="urn:lsid:plazi:treatment:7F54080DFFE6FF89D56AD845FA80FE5B" httpUri="http://treatment.plazi.org/id/7F54080DFFE6FF89D56AD845FA80FE5B" lastPageId="13" lastPageNumber="381" pageId="11" pageNumber="380" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="BFE7EA90FFE6FF8FD56AD845FD2BFA73" box="[264,681,1388,1414]" pageId="11" pageNumber="380" type="nomenclature">
<paragraph id="F742B91BFFE6FF8FD56AD845FD2BFA73" blockId="11.[264,681,1388,1414]" box="[264,681,1388,1414]" pageId="11" pageNumber="380">
<heading id="AC0A0E77FFE6FF8FD56AD845FD2BFA73" bold="true" box="[264,681,1388,1414]" fontSize="11" level="1" pageId="11" pageNumber="380" reason="1">
<emphasis id="C5896509FFE6FF8FD56AD845FD2BFA73" bold="true" box="[264,681,1388,1414]" pageId="11" pageNumber="380">
<taxonomicName id="30FDC298FFE6FF8FD56AD845FE4DFA73" box="[264,463,1388,1414]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="380" phylum="Arthropoda" rank="species" species="moguntia" status="sp. nov.">
<emphasis id="C5896509FFE6FF8FD56AD845FE4DFA73" bold="true" box="[264,463,1388,1414]" italics="true" pageId="11" pageNumber="380">Laena moguntia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE6FF8FD5B5D845FD9BFA73" box="[471,537,1388,1414]" pageId="11" pageNumber="380" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="6FC6A59EFFE6FF8FD64AD845FDF6FA73" box="[552,628,1388,1414]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="11" pageNumber="380">Figs 9</figureCitation>
,
<figureCitation id="6FC6A59EFFE6FF8FD6E1D845FD20FA73" box="[643,674,1388,1414]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="11" pageNumber="380">22</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFE6FF8FD56AD89FFB00FA24" box="[264,1154,1462,1489]" pageId="11" pageNumber="380" type="materials_examined">
<paragraph id="F742B91BFFE6FF8FD56AD89FFB00FA24" blockId="11.[264,1323,1462,1889]" box="[264,1154,1462,1489]" pageId="11" pageNumber="380">
<typeStatus id="284607B9FFE6FF8FD56AD89FFEF8FA25" box="[264,378,1462,1488]" pageId="11" pageNumber="380" type="holotype">
<emphasis id="C5896509FFE6FF8FD56AD89FFEF8FA25" bold="true" box="[264,378,1462,1488]" pageId="11" pageNumber="380">Holotype</emphasis>
</typeStatus>
(♂): N
<collectingCountry id="8FEAF98BFFE6FF8FD5B0D89EFDBAFA24" box="[466,568,1463,1489]" name="Vietnam" pageId="11" pageNumber="380">Vietnam</collectingCountry>
, Tam
<collectingCountry id="8FEAF98BFFE6FF8FD61DD89EFD31FA24" box="[639,691,1463,1489]" name="Iceland" pageId="11" pageNumber="380">Dao</collectingCountry>
,
<date id="83439FDBFFE6FF8FD6DDD89EFCF2FA24" box="[703,880,1463,1489]" pageId="11" pageNumber="380" value="1998-08-01" valueMax="1998-08-08" valueMin="1998-08-01">1.8.VIII.1998</date>
, leg. N. Kanie,
<collectionCode id="91EC21DEFFE6FF8FD045D89EFBFCFA24" box="[1063,1150,1463,1489]" pageId="11" pageNumber="380">CKAO</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFE6FF8FD55AD8F7FDD7F9BC" pageId="11" pageNumber="380" type="etymology">
<paragraph id="F742B91BFFE6FF8FD55AD8F7FDD7F9BC" blockId="11.[264,1323,1462,1889]" pageId="11" pageNumber="380">
<emphasis id="C5896509FFE6FF8FD55AD8F7FE4AFA0D" bold="true" box="[312,456,1502,1528]" pageId="11" pageNumber="380">Etymology:</emphasis>
Dedicated to the city of Mainz, the Roman Moguntium, where the author received his diploma and PhD at the Johannes Gutenberg University under the supervision of Prof. Dr. Jochen Martens.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFE6FF8FD55ADB7FFDC1F894" pageId="11" pageNumber="380" type="diagnosis">
<paragraph id="F742B91BFFE6FF8FD55ADB7FFDC1F894" blockId="11.[264,1323,1462,1889]" pageId="11" pageNumber="380">
<emphasis id="C5896509FFE6FF8FD55ADB7FFE38F985" bold="true" box="[312,442,1622,1648]" pageId="11" pageNumber="380">Diagnosis:</emphasis>
<taxonomicName id="30FDC298FFE6FF8FD5A0DB7EFD01F985" box="[450,643,1623,1648]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="380" phylum="Arthropoda" rank="species" species="moguntia" status="sp. nov.">
<emphasis id="C5896509FFE6FF8FD5A0DB7EFD01F985" box="[450,643,1623,1648]" italics="true" pageId="11" pageNumber="380">Laena moguntia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE6FF8FD6E9DB7FFD4CF985" box="[651,718,1622,1648]" pageId="11" pageNumber="380" rank="species">
<emphasis id="C5896509FFE6FF8FD6E9DB7FFD4CF985" bold="true" box="[651,718,1622,1648]" pageId="11" pageNumber="380">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="30FDC298FFE6FF8FD76BDB7EFB3FF984" authority="Masumoto, 1996" authorityName="Masumoto" authorityYear="1996" box="[777,1213,1623,1649]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="380" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="C5896509FFE6FF8FD76BDB7EFC6FF985" box="[777,1005,1623,1648]" italics="true" pageId="11" pageNumber="380">Laena tamdaoensis</emphasis>
Masumoto, 1996
</taxonomicName>
from the same locality in
<collectingCountry id="8FEAF98BFFE6FF8FD5B4DB56FDBBF96C" box="[470,569,1663,1689]" name="Vietnam" pageId="11" pageNumber="380">Vietnam</collectingCountry>
share the bigger body size, prominent eyes, crenulate lateral pronotal margins and dorsal structure and setation of the elytra, but can be separated by a rougher dorsal punctation with distinct impressions on the pronotum (compare
<figureCitation id="6FC6A59EFFE6FF8FD0C1DBE6FA98F91C" box="[1187,1306,1743,1769]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="11" pageNumber="380">Figs 9, 14</figureCitation>
), rounder shape of the elytra, the alternate intervals nearly keel­like (
<figureCitation id="6FC6A59EFFE6FF8FD07CDBDEFB1AF8E4" box="[1054,1176,1783,1809]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="11" pageNumber="380">Figs 9, 14</figureCitation>
), the pair of distinct teeth on each femur (only a single tooth in
<taxonomicName id="30FDC298FFE6FF8FD708DA36FBCFF8CD" box="[874,1101,1823,1848]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="380" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="C5896509FFE6FF8FD708DA36FBCFF8CD" box="[874,1101,1823,1848]" italics="true" pageId="11" pageNumber="380">Laena tamdaoensis</emphasis>
</taxonomicName>
), and the shape of the aedeagus (
<figureCitation id="6FC6A59EFFE6FF8FD5CCDA6EFDB4F894" box="[430,566,1863,1889]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="11" pageNumber="380">Figs 22, 25</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="A382E993FFE1FF88D56AD84FFEEAF9BB" httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="12" pageNumber="381" targetBox="[346,1246,319,1358]" targetPageId="12">
<paragraph id="F742B91BFFE1FF88D56AD84FFEEAF9BB" blockId="12.[264,1323,1382,1614]" pageId="12" pageNumber="381">
<emphasis id="C5896509FFE1FF88D56AD84FFE52FA8B" bold="true" box="[264,464,1382,1406]" pageId="12" pageNumber="381">FIGURES 1726.</emphasis>
Aedeagus of
<taxonomicName id="30FDC298FFE1FF88D60BD84EFD29FA8B" box="[617,683,1383,1406]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="C5896509FFE1FF88D60BD84EFD29FA8B" box="[617,683,1383,1406]" italics="true" pageId="12" pageNumber="381">Laena</emphasis>
</taxonomicName>
species. 17
<taxonomicName id="30FDC298FFE1FF88D756D84EFB82FA8B" box="[820,1024,1383,1406]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="inthanonica" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D756D84EFB82FA8B" box="[820,1024,1383,1406]" italics="true" pageId="12" pageNumber="381">Laena inthanonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D06AD84FFBC4FA8B" box="[1032,1094,1382,1406]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D06AD84FFBC4FA8B" bold="true" box="[1032,1094,1382,1406]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype MHNG, Thailand. 18
<taxonomicName id="30FDC298FFE1FF88D5F4D8A3FDADFA54" box="[406,559,1418,1441]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="jocheni" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D5F4D8A3FDADFA54" box="[406,559,1418,1441]" italics="true" pageId="12" pageNumber="381">Laena jocheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D657D8A0FDF0FA54" box="[565,626,1417,1441]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D657D8A0FDF0FA54" bold="true" box="[565,626,1417,1441]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype SMNS, Laos. 19
<taxonomicName id="30FDC298FFE1FF88D7D4D8A3FBE1FA54" box="[950,1123,1418,1441]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="kenyirica" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D7D4D8A3FBE1FA54" box="[950,1123,1418,1441]" italics="true" pageId="12" pageNumber="381">Laena kenyirica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D008D8A0FB25FA54" box="[1130,1191,1417,1441]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D008D8A0FB25FA54" bold="true" box="[1130,1191,1417,1441]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype SMNS, Malaysia. 20
<taxonomicName id="30FDC298FFE1FF88D58FD885FD1AFA36" box="[493,664,1452,1475]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="khaolaka" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D58FD885FD1AFA36" box="[493,664,1452,1475]" italics="true" pageId="12" pageNumber="381">Laena khaolaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D6FDD882FD58FA36" box="[671,730,1451,1475]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D6FDD882FD58FA36" bold="true" box="[671,730,1451,1475]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype SMNS, Thailand. 21
<taxonomicName id="30FDC298FFE1FF88D024D885FB60FA36" box="[1094,1250,1452,1475]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="kiyoshii" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D024D885FB60FA36" box="[1094,1250,1452,1475]" italics="true" pageId="12" pageNumber="381">Laena kiyoshii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D08AD882FAA6FA36" box="[1256,1316,1451,1475]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D08AD882FAA6FA36" bold="true" box="[1256,1316,1451,1475]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype SMNS, Laos. 22
<taxonomicName id="30FDC298FFE1FF88D65ED8E6FD68FA13" box="[572,746,1487,1510]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="moguntia" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D65ED8E6FD68FA13" box="[572,746,1487,1510]" italics="true" pageId="12" pageNumber="381">Laena moguntia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D693D8E7FCAEFA13" box="[753,812,1486,1510]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D693D8E7FCAEFA13" bold="true" box="[753,812,1486,1510]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype CKAO, Vietnam. 23
<taxonomicName id="30FDC298FFE1FF88D0F5D8E6FAA8FA13" box="[1175,1322,1487,1510]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D0F5D8E6FAA8FA13" box="[1175,1322,1487,1510]" italics="true" pageId="12" pageNumber="381">Laena schulzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D56AD8D8FECAF9FC" box="[264,328,1521,1545]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D56AD8D8FECAF9FC" bold="true" box="[264,328,1521,1545]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype SMNS, Malaysia. 24
<taxonomicName id="30FDC298FFE1FF88D6B2D8DBFCE9F9FC" box="[720,875,1522,1545]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="rolandi" status="sp. nov.">
<emphasis id="C5896509FFE1FF88D6B2D8DBFCE9F9FC" box="[720,875,1522,1545]" italics="true" pageId="12" pageNumber="381">Laena rolandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE1FF88D714D8D8FC34F9FC" box="[886,950,1521,1545]" pageId="12" pageNumber="381" rank="species">
<emphasis id="C5896509FFE1FF88D714D8D8FC34F9FC" bold="true" box="[886,950,1521,1545]" pageId="12" pageNumber="381">n. sp.</emphasis>
</taxonomicNameLabel>
, ♂ holotype CRGT, Thailand. 25
<taxonomicName id="30FDC298FFE1FF88D56ADB3DFE5FF9DE" box="[264,477,1556,1579]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="C5896509FFE1FF88D56ADB3DFE5FF9DE" box="[264,477,1556,1579]" italics="true" pageId="12" pageNumber="381">Laena tamdaoensis</emphasis>
</taxonomicName>
, ♂ holotype NSMT, Vietnam. 26
<taxonomicName id="30FDC298FFE1FF88D70CDB3DFBB1F9DE" box="[878,1075,1556,1579]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="C5896509FFE1FF88D70CDB3DFBB1F9DE" box="[878,1075,1556,1579]" italics="true" pageId="12" pageNumber="381">Laena vietnamica</emphasis>
</taxonomicName>
, non­type ♂ HNHM, Vietnam.
</paragraph>
</caption>
<subSubSection id="BFE7EA90FFE1FF89D55ADBA8FA80FE5B" lastPageId="13" lastPageNumber="382" pageId="12" pageNumber="381" type="description">
<paragraph id="F742B91BFFE1FF89D55ADBA8FA80FE5B" blockId="12.[264,1323,1665,1892]" lastBlockId="13.[264,1322,284,430]" lastPageId="13" lastPageNumber="382" pageId="12" pageNumber="381">
<emphasis id="C5896509FFE1FF88D55ADBA8FE50F96E" bold="true" box="[312,466,1665,1691]" pageId="12" pageNumber="381">Description:</emphasis>
Body length 7.0 mm. Eyes prominent. Pronotum (
<figureCitation id="6FC6A59EFFE1FF88D02ADBABFB10F969" box="[1096,1170,1666,1692]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="12" pageNumber="381">Fig. 9</figureCitation>
) with larger punctures, distance between 0.51 diameters, most punctures bearing long erect seta; surface with distinct mediobasal impression and a pair of distinct discal impressions, surface dull; lateral margin distinctly bordered, crenulate; propleura with finer, sparser punctation but similar setation. Elytra (
<figureCitation id="6FC6A59EFFE1FF88D682DA0BFCAAF8C9" box="[736,808,1826,1852]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="12" pageNumber="381">Fig. 9</figureCitation>
) with complete rows of punctures without striae, equal in size to pronotal punctures, most in particular posteriorly bearing long, erect seta; intervals with a row of smaller punctures each bearing erect seta of same length, interval 9 with 6 distinct setiferous pores; intervals distinctly convex, alternate intervals 3, 5 and 7 much higher and nearly keel­like, surface shining. Femora with a pair of distinct identical teeth. Tibiae in males without distinct sexual character. Aedeagus see
<figureCitation id="6FC6A59EFFE0FF89D0CADCBDFB7CFE5B" box="[1192,1278,404,430]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="13" pageNumber="382">Fig. 22</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>