740 lines
97 KiB
XML
740 lines
97 KiB
XML
<document id="E053E5A8A93B6C0C6FF6215068E78E98" ID-CLB-Dataset="45529" ID-DOI="10.11646/zootaxa.3814.3.9" ID-GBIF-Dataset="e0c4a1bd-2e49-4b4a-86d9-b379ca472026" ID-ISSN="1175-5326" ID-Zenodo-Dep="225971" ID-ZooBank="128E7CA8-E8EB-446B-961A-C27838DFD357" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434953917" checkinUser="plazi" docAuthor="Hertwig, Stefan T., Min, Pui Yong, Haas, Alexander & Das, Indraneil" docDate="2014" docId="7E7FDE5BFF98FFC2FF5DFBB7FAEF6179" docLanguage="en" docName="zt03814p431.pdf" docOrigin="Zootaxa 3814 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ansonia vidua Hertwig, Min, Haas & Das, 2014, sp. nov." docType="treatment" docVersion="11" lastPageNumber="428" masterDocId="8246A623FF9BFFCBFFCAFFEAFF816505" masterDocTitle="Dressed in black. A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo)" masterLastPageNumber="431" masterPageNumber="419" pageNumber="422" updateTime="1698331971278" updateUser="ExternalLinkService">
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<mods:mods id="733711F8136572754394FD33F356B5C2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="872B59A21C9BACB7CA158DD27247DA8B">
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<mods:title id="ACB426DF6E83AB5E0663857B1CCCE0EC">Dressed in black. A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo)</mods:title>
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</mods:titleInfo>
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<mods:name id="28DA3472E55CA8CFB13419ADC0C70CE0" type="personal">
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<mods:role id="F1414A25585D474933DA0E3819C2CA40">
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<mods:roleTerm id="C2992F25ADDD714747AED3B3DBD72DDA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A5446222239D8D6BB192341E37B310EB">Hertwig, Stefan T.</mods:namePart>
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</mods:name>
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<mods:name id="154A8F2B96F77956464DB4FD73FECE91" type="personal">
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<mods:role id="CBEFEBB8A3BA090581AACA48B7A558F6">
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<mods:roleTerm id="DA8F9957A4021AA07FA131EF47EB922C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5F97EFDFF658A8D8140153DF4C2A9301">Min, Pui Yong</mods:namePart>
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</mods:name>
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<mods:roleTerm id="63404E8F249A918B37B720749A2FE1B2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B06488C496F59330C6FC312B0200B132">Haas, Alexander</mods:namePart>
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</mods:name>
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<mods:name id="52CFEF529CDD5635DB2CE389ACD0E7B0" type="personal">
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<mods:role id="ABB9B074BFD435523328D868743847D6">
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<mods:roleTerm id="4529F3947B354BAF67DE10E92D492040">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C6EEB094DDD068EBF1B2E8AD9C7715C">Das, Indraneil</mods:namePart>
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<mods:typeOfResource id="CE9A67FF752475E6796A1A90B7B916D9">text</mods:typeOfResource>
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<mods:title id="489D91801A48ABFD83FA9AD74C2363BE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="816AD0BE21B3C04E8D68F00598A8E968">
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<mods:date id="A0FACF1E7EF33F57DBF496788389B4E0">2014</mods:date>
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<mods:number id="DF1E5BAA6DE6BEDECCF1886C560DDA78">2014-06-10</mods:number>
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<mods:detail id="166E4B6B5581A28539338093EA505A1E" type="volume">
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<mods:number id="49C9418CB0E6EDE19BBB52A3AE04C20D">3814</mods:number>
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</mods:detail>
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<mods:detail id="32AF37BD1B9C40DB1F16B82B1288F98D" type="issue">
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<mods:number id="8977A6C7058B094302D42559F2240A0D">3</mods:number>
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</mods:mods>
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<subSubSection id="BECC3CC6FF98FFC8FF5DFBB7FE336172" box="[151,434,1117,1143]" pageId="3" pageNumber="422" type="nomenclature">
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<paragraph id="F6696F4DFF98FFC8FF5DFBB7FE336172" blockId="3.[151,434,1117,1143]" box="[151,434,1117,1143]" pageId="3" pageNumber="422">
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<heading id="AD21D821FF98FFC8FF5DFBB7FE336172" bold="true" box="[151,434,1117,1143]" fontSize="11" level="1" pageId="3" pageNumber="422" reason="1">
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<emphasis id="C4A2B35FFF98FFC8FF5DFBB7FE336172" bold="true" box="[151,434,1117,1143]" pageId="3" pageNumber="422">
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<emphasis id="C4A2B35FFF98FFC8FF5DFBB7FECC6172" bold="true" box="[151,333,1117,1143]" italics="true" pageId="3" pageNumber="422">
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<taxonomicName id="31D614CEFF98FFC8FF5DFBB7FEC66172" ID-CoL="66Y9Q" box="[151,327,1117,1143]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">Ansonia vidua</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="DF910E24FF98FFC8FE9EFBB7FE336172" box="[340,434,1117,1143]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="BECC3CC6FF98FFC8FF5DFB74FE6A608A" pageId="3" pageNumber="422" type="materials_examined">
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<paragraph id="F6696F4DFF98FFC8FF5DFB74FA046026" blockId="3.[151,1437,1182,2035]" pageId="3" pageNumber="422">
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<materialsCitation id="46BE6510FF98FFC8FF5DFB74FA046026" ID-GBIF-Occurrence="3320068301" accessionNumber="KJ488546" collectingDate="2012-02-28" collectionCode="NMBE" collectorName="Y. M. Pui & A. Jankowski & S. T. Hertwig" country="Malaysia" county="Sarawak" elevation="2152" latitude="3.9120166" location="Church Camp and the summit of Gunung Murud, near the intersection with Sungei Murud, Gunung Murud" longLatPrecision="1" longitude="115.49924" municipality="Pulong Tau National Park" pageId="3" pageNumber="422" specimenCode="NMBE 1061645, HJ12-125" specimenCount="1" typeStatus="holotype">
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<emphasis id="C4A2B35FFF98FFC8FF5DFB74FE8C61B2" bold="true" box="[151,269,1182,1207]" pageId="3" pageNumber="422">
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<typeStatus id="296DD1EFFF98FFC8FF5DFB74FE8961B2" box="[151,264,1182,1207]" pageId="3" pageNumber="422" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="A670C736FF98FFC8FEDEFB75FE5361B2" box="[276,466,1182,1207]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="3" pageNumber="422">NMBE 1061645</specimenCode>
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, field number
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<specimenCode id="A670C736FF98FFC8FDBFFB74FD6861B2" box="[629,745,1182,1207]" pageId="3" pageNumber="422">HJ12–125</specimenCode>
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(
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<figureCitation id="6EED73C8FF98FFC8FD32FB74FCE461B2" box="[760,869,1182,1207]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="6.[151,250,1071,1093]" captionStartId-1="7.[151,250,1827,1849]" captionStartId-2="8.[151,250,699,721]" captionStartId-3="8.[151,250,1629,1651]" captionTargetBox-0="[151,1436,193,1049]" captionTargetBox-1="[151,1436,193,1805]" captionTargetId-0="figure@6.[151,1436,193,1050]" captionTargetId-1="figure@7.[151,1436,193,1805]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." captionText-1="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." captionText-2="FIGURE 4. Head of holotype of Ansonia vidua sp. nov. in lateral view." captionText-3="FIGURE 5. Foot of holotype of Ansonia vidua sp. nov. in ventral view." httpUri-0="https://zenodo.org/record/225973/files/figure.png" httpUri-1="https://zenodo.org/record/225974/files/figure.png" pageId="3" pageNumber="422">Figs. 2–5</figureCitation>
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||
), GenBank accession number
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||
<accessionNumber id="E985F2AEFF98FFC8FB71FB74FAB061B2" box="[1211,1329,1182,1207]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ488546" pageId="3" pageNumber="422">KJ488546</accessionNumber>
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, from the summit trail between
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||
<location id="F3093996FF98FFC8FE44FB28FEC961FA" LSID="urn:lsid:plazi:treatment:7E7FDE5BFF98FFC2FF5DFBB7FAEF6179:F3093996FF98FFC8FE44FB28FEC961FA" country="Malaysia" county="Sarawak" latitude="3.9120166" longLatPrecision="1" longitude="115.49924" municipality="Pulong Tau National Park" name="Church Camp and the summit of Gunung Murud, near the intersection with Sungei Murud, Gunung Murud" pageId="3" pageNumber="422">Church Camp and the summit of Gunung Murud, near the intersection with Sungei Murud, Gunung Murud</location>
|
||
,
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<collectingMunicipality id="160DF537FF98FFC8FE9EFB0CFDFB61FA" box="[340,634,1254,1279]" pageId="3" pageNumber="422">Pulong Tau National Park</collectingMunicipality>
|
||
,
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||
<collectingCounty id="1F0817C1FF98FFC8FD4DFB0CFD6B61FA" box="[647,746,1254,1279]" pageId="3" pageNumber="422">Sarawak</collectingCounty>
|
||
, East
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||
<collectingCountry id="8EC12FDDFF98FFC8FCE4FB0CFC1461FA" box="[814,917,1254,1279]" name="Malaysia" pageId="3" pageNumber="422">Malaysia</collectingCountry>
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||
(Borneo), altitude
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||
<elevation id="7DFB887EFF98FFC8FBA1FB0CFB6E61FA" box="[1131,1263,1254,1279]" metricMagnitude="3" metricUnit="m" metricValue="2.152" pageId="3" pageNumber="422" unit="m" value="2152.0">
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<quantity id="312EC2A8FF98FFC8FBA1FB0CFB4961FB" box="[1131,1224,1254,1279]" metricMagnitude="3" metricUnit="m" metricValue="2.152" pageId="3" pageNumber="422" unit="m" value="2152.0">2,152 m</quantity>
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||
asl
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||
</elevation>
|
||
(
|
||
<geoCoordinate id="93E2098AFF98FFC8FB35FB0DFA1861FA" box="[1279,1433,1254,1279]" degrees="03" direction="north" minutes="54.721" orientation="latitude" pageId="3" pageNumber="422" precision="1" value="3.9120166">N 03°54.721'</geoCoordinate>
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||
,
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||
<geoCoordinate id="93E2098AFF98FFC8FF5DFAE1FEB36026" box="[151,306,1290,1315]" degrees="115" direction="east" minutes="29.954" orientation="longitude" pageId="3" pageNumber="422" precision="1" value="115.49924">E 115°29.954</geoCoordinate>
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, GPS WGS 84), coll.
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<collectorName id="5B230A9BFF98FFC8FDE3FAE1FD176026" box="[553,662,1290,1315]" pageId="3" pageNumber="422">Y. M. Pui</collectorName>
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||
,
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<collectorName id="5B230A9BFF98FFC8FD68FAE1FCBB6026" box="[674,826,1290,1315]" pageId="3" pageNumber="422">A. Jankowski</collectorName>
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and
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<collectorName id="5B230A9BFF98FFC8FCBBFAE1FB886026" box="[881,1033,1290,1315]" pageId="3" pageNumber="422">S. T. Hertwig</collectorName>
|
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,
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||
<date id="8268498DFF98FFC8FBDFFAE1FB616026" box="[1045,1248,1290,1315]" pageId="3" pageNumber="422" value="2012-02-28">
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||
<collectingDate id="922CB065FF98FFC8FBDFFAE1FB616026" box="[1045,1248,1290,1315]" pageId="3" pageNumber="422" value="2012-02-28">28 February 2012</collectingDate>
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||
</date>
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||
.
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<specimenCount id="E0D0A4C4FF98FFC8FB20FAE0FA006026" box="[1258,1409,1290,1315]" pageId="3" pageNumber="422" type="adult">Adult female</specimenCount>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="F6696F4DFF98FFC8FF0DFAC4FE6A608A" blockId="3.[151,1437,1182,2035]" pageId="3" pageNumber="422">
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||
<materialsCitation id="46BE6510FF98FFC8FF0DFAC4FE66608A" ID-GBIF-Occurrence="3320068302" accessionNumber="KJ488547" collectingDate="2013-03-10" collectionCode="NMBE" collectorName="Y. M. Pui & T. Einecke & T. Keilholz & S. T. Hertwig" location="25 m away from the locality of holotype at the summit trail" pageId="3" pageNumber="422" specimenCode="NMBE 1066153, SH13-291, Adult female" specimenCount="1" typeStatus="paratype">
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<emphasis id="C4A2B35FFF98FFC8FF0DFAC4FEBC6042" bold="true" box="[199,317,1326,1351]" pageId="3" pageNumber="422">
|
||
<typeStatus id="296DD1EFFF98FFC8FF0DFAC4FEB96042" box="[199,312,1326,1351]" pageId="3" pageNumber="422" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="A670C736FF98FFC8FE8EFAC5FD836042" box="[324,514,1326,1351]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="3" pageNumber="422">NMBE 1066153</specimenCode>
|
||
, field number
|
||
<specimenCode id="A670C736FF98FFC8FD6CFAC5FC9C6042" box="[678,797,1327,1351]" pageId="3" pageNumber="422">SH13–291</specimenCode>
|
||
(
|
||
<figureCitation id="6EED73C8FF98FFC8FCE7FAC4FCF16042" box="[813,880,1326,1351]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="3" pageNumber="422">Fig. 3</figureCitation>
|
||
C, D), GenBank accession number
|
||
<accessionNumber id="E985F2AEFF98FFC8FB35FAC4FAF56042" box="[1279,1396,1326,1351]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ488547" pageId="3" pageNumber="422">
|
||
KJ
|
||
<quantity id="312EC2A8FF98FFC8FAD4FAC4FAF56042" box="[1310,1396,1326,1351]" metricMagnitude="5" metricUnit="m" metricValue="4.8854725" pageId="3" pageNumber="422" unit="m" value="488547.25">488547</quantity>
|
||
</accessionNumber>
|
||
,
|
||
<location id="F3093996FF98FFC8FA4AFAC4FC8D606E" LSID="urn:lsid:plazi:treatment:7E7FDE5BFF98FFC2FF5DFBB7FAEF6179:F3093996FF98FFC8FA4AFAC4FC8D606E" name="25 m away from the locality of holotype at the summit trail" pageId="3" pageNumber="422">25 m away from the locality of holotype at the summit trail</location>
|
||
, coll.
|
||
<collectorName id="5B230A9BFF98FFC8FC85FAB9FC3D606E" box="[847,956,1362,1387]" pageId="3" pageNumber="422">Y. M. Pui</collectorName>
|
||
,
|
||
<collectorName id="5B230A9BFF98FFC8FC0DFAB9FBC0606E" box="[967,1089,1362,1387]" pageId="3" pageNumber="422">T. Einecke</collectorName>
|
||
,
|
||
<collectorName id="5B230A9BFF98FFC8FB86FAB9FB48606E" box="[1100,1225,1362,1387]" pageId="3" pageNumber="422">T. Keilholz</collectorName>
|
||
and
|
||
<collectorName id="5B230A9BFF98FFC8FB35FAB9FA19606E" box="[1279,1432,1362,1387]" pageId="3" pageNumber="422">S. T. Hertwig</collectorName>
|
||
,
|
||
<date id="8268498DFF98FFC8FF5DFA9DFEC7608A" box="[151,326,1398,1423]" pageId="3" pageNumber="422" value="2013-03-10">
|
||
<collectingDate id="922CB065FF98FFC8FF5DFA9DFEC7608A" box="[151,326,1398,1423]" pageId="3" pageNumber="422" value="2013-03-10">10 March 2013</collectingDate>
|
||
</date>
|
||
.
|
||
<specimenCode id="A670C736FF98FFC8FE9AFA9CFE66608A" box="[336,487,1398,1423]" pageId="3" pageNumber="422">Adult female</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BECC3CC6FF98FFC8FF0DFA70FE976266" pageId="3" pageNumber="422" type="diagnosis">
|
||
<paragraph id="F6696F4DFF98FFC8FF0DFA70FC7C60FE" blockId="3.[151,1437,1182,2035]" pageId="3" pageNumber="422">
|
||
<emphasis id="C4A2B35FFF98FFC8FF0DFA70FEC060B6" bold="true" box="[199,321,1434,1459]" pageId="3" pageNumber="422">Diagnosis.</emphasis>
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The new species is assigned to the genus
|
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<taxonomicName id="31D614CEFF98FFC8FCEDFA71FC0260B7" box="[807,899,1435,1458]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="3" pageNumber="422" phylum="Chordata" rank="genus">
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||
<emphasis id="C4A2B35FFF98FFC8FCEDFA71FC0260B7" box="[807,899,1435,1458]" italics="true" pageId="3" pageNumber="422">Ansonia</emphasis>
|
||
</taxonomicName>
|
||
based on its relatively slender body, long legs, indistinct subarticular tubercles, membraneous digitial webbing, the absence of parotoid glands, tympanum visible (
|
||
<bibRefCitation id="924712BCFF98FFC8FF54FA09FE9C60FE" author="Inger" box="[158,285,1506,1531]" firstAuthor="Inger" pageId="3" pageNumber="422" refString="Inger, R. F. (1960) A review of the Oriental toads of the genus Ansonia Stoliczka. Fieldiana: Zoology, 39, 473 - 503." type="journal article" year="1960">Inger 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="924712BCFF98FFC8FEE2FA08FEE060FE" author="Inger" box="[296,353,1506,1531]" firstAuthor="Inger" pageId="3" pageNumber="422" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">1966</bibRefCitation>
|
||
) and a robustly supported phylogenetic hypothesis (
|
||
<figureCitation id="6EED73C8FF98FFC8FC60FA08FC6D60FE" box="[938,1004,1506,1531]" captionStart="FIGURE 1" captionStartId="4.[151,250,1946,1968]" captionTargetBox="[150,1414,193,1895]" captionTargetId="figure@4.[150,1414,193,1895]" captionTargetPageId="4" captionText="FIGURE 1. Phylogenetic relationships of Ansonia vidua sp. nov. according to a RaxML-based Maximum Likelihood analysis of partial sequences of the mitochondrial 16 S rRNA gene. Node labels above branches show the bootstrapping support of ML analysis (only values> 50), below branches the posterior probability values of BI (only values> 70)." httpUri="https://zenodo.org/record/225972/files/figure.png" pageId="3" pageNumber="422">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F6696F4DFF98FFC8FF0DF9EDFE976266" blockId="3.[151,1437,1182,2035]" pageId="3" pageNumber="422">
|
||
<taxonomicName id="31D614CEFF98FFC8FF0DF9EDFEEF631B" box="[199,366,1542,1566]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="3" pageNumber="422" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF98FFC8FF0DF9EDFEEF631B" box="[199,366,1542,1566]" italics="true" pageId="3" pageNumber="422">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DF910E24FF98FFC8FEB1F9ECFE5A631B" box="[379,475,1542,1567]" pageId="3" pageNumber="422" rank="species">
|
||
<emphasis id="C4A2B35FFF98FFC8FEB1F9ECFE55631A" bold="true" box="[379,468,1542,1567]" pageId="3" pageNumber="422">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
can be differentiated from all other congeners by the following combination of characters: SVL 33.5 and
|
||
<quantity id="312EC2A8FF98FFC8FE72F9C1FD9A6347" box="[440,539,1579,1603]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" pageId="3" pageNumber="422" unit="mm" value="34.3">34.3 mm</quantity>
|
||
in the
|
||
<specimenCount id="E0D0A4C4FF98FFC8FDACF9C1FD6C6346" box="[614,749,1578,1603]" pageId="3" pageNumber="422" type="female">two females</specimenCount>
|
||
known; body not flattened; canthus rostralis slightly swollen; two low longitudinal interorbital ridges present; dorsum of body, flanks and upper surfaces of the limbs covered with numerous small, regular, low, rounded warts, resulting in a velvet-like appearance, only warts on scapular and temporal regions slightly enlarged; warts on dorsum, head and limbs terminating in a single fine, keratinous spine; first finger short, not reaching tip of second finger when adpressed; no sharp tarsal ridge; no skin flaps on posterior thigh near vent; third and fifth toe with 2½ phalanges free of web medially; colouration of the dorsal surface of head and body uniformly black-brown; limbs dark brown with lighter areas on joints of the limbs and phalanges; ventral surface uniformly dark grey to yellowish brown; dorsal and ventral parts of body and limbs completely patternless.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BECC3CC6FF98FFCEFF0DF884FE1A62F3" lastPageId="5" lastPageNumber="424" pageId="3" pageNumber="422" type="description">
|
||
<paragraph id="F6696F4DFF98FFCEFF0DF884FBD66461" blockId="3.[151,1437,1182,2035]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="424" pageId="3" pageNumber="422">
|
||
<emphasis id="C4A2B35FFF98FFC8FF0DF884FE6D6282" bold="true" box="[199,492,1902,1927]" pageId="3" pageNumber="422">
|
||
Description of
|
||
<typeStatus id="296DD1EFFF98FFC8FEB7F884FE676282" box="[381,486,1902,1927]" pageId="3" pageNumber="422" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Habitus slender; body as wide as head (
|
||
<figureCitation id="6EED73C8FF98FFC8FC03F884FBB66282" box="[969,1079,1902,1927]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,250,1071,1093]" captionStartId-1="7.[151,250,1827,1849]" captionTargetBox-0="[151,1436,193,1049]" captionTargetBox-1="[151,1436,193,1805]" captionTargetId-0="figure@6.[151,1436,193,1050]" captionTargetId-1="figure@7.[151,1436,193,1805]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." captionText-1="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri-0="https://zenodo.org/record/225973/files/figure.png" httpUri-1="https://zenodo.org/record/225974/files/figure.png" pageId="3" pageNumber="422">Figs. 2–3</figureCitation>
|
||
A, B), body not dorsoventrally flattened; head slightly longer than wide (
|
||
<tableCitation id="BB545AF6FF98FFC8FDA4F878FD4662AE" box="[622,711,1938,1963]" captionStart="TABLE 2" captionStartId="5.[151,239,398,421]" captionTargetBox="[159,1374,449,1122]" captionTargetPageId="5" captionText="TABLE 2. Measurements of holotype and paratype of Ansonia vidua sp. nov., for details and abbreviations see text." httpUri="http://table.plazi.org/id/A2A93FC5FF9EFFCEFF5DFE64FAF164A0" pageId="3" pageNumber="422" tableUuid="A2A93FC5FF9EFFCEFF5DFE64FAF164A0">Table 2</tableCitation>
|
||
); HWAJ/TBL 0.62; rostrum truncate in dorsal view, tip of snout rounded, longer than canthus, oblique in lateral view, projecting beyond tip of mandible, longitudinal, medial groove on the dorsal face of snout, rostral length greater than eye diameter; nostril lateral, closer to snout tip than eye (
|
||
<figureCitation id="6EED73C8FF9EFFCEFF07FF7DFE9065B5" box="[205,273,151,176]" captionStart="FIGURE 4" captionStartId="8.[151,250,699,721]" captionText="FIGURE 4. Head of holotype of Ansonia vidua sp. nov. in lateral view." pageId="5" pageNumber="424">Fig. 4</figureCitation>
|
||
); distinct vertical ridge between tip of snout and centre of maxilla (
|
||
<figureCitation id="6EED73C8FF9EFFCEFC37FF7DFBC965B5" box="[1021,1096,151,176]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Figs. 3</figureCitation>
|
||
A, B, 4); canthus swollen with rounded edge, slightly arcuate in lateral view; lores vertical and concave; eye diameter greater than eye-nostril distance; interorbital distance subequal to upper eyelid (
|
||
<tableCitation id="BB545AF6FF9EFFCEFCE4FF35FC0865FD" box="[814,905,223,248]" captionStart="TABLE 2" captionStartId="5.[151,239,398,421]" captionTargetBox="[159,1374,449,1122]" captionTargetPageId="5" captionText="TABLE 2. Measurements of holotype and paratype of Ansonia vidua sp. nov., for details and abbreviations see text." httpUri="http://table.plazi.org/id/A2A93FC5FF9EFFCEFF5DFE64FAF164A0" pageId="5" pageNumber="424" tableUuid="A2A93FC5FF9EFFCEFF5DFE64FAF164A0">Table 2</tableCitation>
|
||
); tympanum distinct, eye diameter 1.9 times greater than tympanic diameter (
|
||
<figureCitation id="6EED73C8FF9EFFCEFDC0FEEEFDD16419" box="[522,592,260,285]" captionStart="FIGURE 4" captionStartId="8.[151,250,699,721]" captionText="FIGURE 4. Head of holotype of Ansonia vidua sp. nov. in lateral view." pageId="5" pageNumber="424">Fig. 4</figureCitation>
|
||
,
|
||
<tableCitation id="BB545AF6FF9EFFCEFD94FEEEFD356419" box="[606,692,260,285]" captionStart="TABLE 2" captionStartId="5.[151,239,398,421]" captionTargetBox="[159,1374,449,1122]" captionTargetPageId="5" captionText="TABLE 2. Measurements of holotype and paratype of Ansonia vidua sp. nov., for details and abbreviations see text." httpUri="http://table.plazi.org/id/A2A93FC5FF9EFFCEFF5DFE64FAF164A0" pageId="5" pageNumber="424" tableUuid="A2A93FC5FF9EFFCEFF5DFE64FAF164A0">Table 2</tableCitation>
|
||
); two low, slightly curvestridges on head forming a parenthetical shape “()”extends from the anterior aspect of the interorbital region to a point approximately in line with the centre of the tympani (
|
||
<figureCitation id="6EED73C8FF9EFFCEFE83FEA6FE0A6461" box="[329,395,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
A), one supratympanic ridge in parotoid region (
|
||
<figureCitation id="6EED73C8FF9EFFCEFC61FEA6FC6F6461" box="[939,1006,332,357]" captionStart="FIGURE 2" captionStartId="6.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." httpUri="https://zenodo.org/record/225973/files/figure.png" pageId="5" pageNumber="424">Fig. 2</figureCitation>
|
||
and
|
||
<figureCitation id="6EED73C8FF9EFFCEFBEEFEA6FBB36461" box="[1060,1074,332,356]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">3</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<caption id="A2A93FC5FF9FFFCFFF5DF870FB0862E8" httpUri="https://zenodo.org/record/225972/files/figure.png" pageId="4" pageNumber="423" targetBox="[150,1414,193,1895]" targetPageId="4">
|
||
<paragraph id="F6696F4DFF9FFFCFFF5DF870FB0862E8" blockId="4.[151,1436,1946,2029]" pageId="4" pageNumber="423">
|
||
<emphasis id="C4A2B35FFF9FFFCFFF5DF870FE9262B5" bold="true" box="[151,275,1946,1968]" pageId="4" pageNumber="423">FIGURE 1.</emphasis>
|
||
Phylogenetic relationships of
|
||
<taxonomicName id="31D614CEFF9FFFCFFD8DF871FD5862B5" box="[583,729,1946,1968]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF9FFFCFFD8DF871FD5862B5" box="[583,729,1946,1968]" italics="true" pageId="4" pageNumber="423">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF9FFFCFFD2AF870FCB162B5" bold="true" box="[736,816,1946,1968]" pageId="4" pageNumber="423">
|
||
<taxonomicNameLabel id="DF910E24FF9FFFCFFD2AF870FCB162B5" box="[736,816,1946,1968]" pageId="4" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
according to a RaxML-based Maximum Likelihood analysis of partial sequences of the mitochondrial 16S rRNA gene. Node labels above branches show the bootstrapping support of ML analysis (only values>50), below branches the posterior probability values of BI (only values>70).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A2A93FC5FF9EFFCEFF5DFE64FAF164A0" ID-Table-UUID="A2A93FC5FF9EFFCEFF5DFE64FAF164A0" box="[151,1392,397,421]" httpUri="http://table.plazi.org/id/A2A93FC5FF9EFFCEFF5DFE64FAF164A0" pageId="5" pageNumber="424" targetBox="[159,1374,449,1122]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="F6696F4DFF9EFFCEFF5DFE64FAF164A0" blockId="5.[151,1392,397,421]" box="[151,1392,397,421]" pageId="5" pageNumber="424">
|
||
<emphasis id="C4A2B35FFF9EFFCEFF5DFE64FE8864A0" bold="true" box="[151,265,398,421]" pageId="5" pageNumber="424">TABLE 2.</emphasis>
|
||
Measurements of holotype and paratype of
|
||
<taxonomicName id="31D614CEFF9EFFCEFD2AFE64FCF964A0" box="[736,888,397,421]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF9EFFCEFD2AFE64FCF964A0" box="[736,888,397,421]" italics="true" pageId="5" pageNumber="424">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF9EFFCEFC4AFE64FC5464A0" bold="true" box="[896,981,398,421]" pageId="5" pageNumber="424">
|
||
<taxonomicNameLabel id="DF910E24FF9EFFCEFC4AFE64FC5464A0" box="[896,981,398,421]" pageId="5" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, for details and abbreviations see text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F6696F4DFF9EFFCEFF55FDE3FB636167" pageId="5" pageNumber="424">
|
||
<table id="84D69DEDFF9E0034FF55FE2BFADF6167" box="[159,1374,449,1122]" gridcols="6" gridrows="6" pageId="5" pageNumber="424">
|
||
<tr id="48E66D0FFF9E0034FF55FE2BFADF674D" box="[159,1374,449,584]" gridrow="0" pageId="5" pageNumber="424">
|
||
<th id="0B370473FF9E0034FF55FE2BFF69674D" box="[159,232,449,584]" gridcol="0" gridrow="0" pageId="5" pageNumber="424">SVL BW</th>
|
||
<th id="0B370473FF9E0034FE8FFE2BFE6F674D" box="[325,494,449,584]" gridcol="1" gridrow="0" pageId="5" pageNumber="424">holotype NMBE 1061645 33.51 12.07</th>
|
||
<th id="0B370473FF9E0034FDF8FE2BFD5B674D" box="[562,730,449,584]" gridcol="2" gridrow="0" pageId="5" pageNumber="424">paratype NMBE 1066153 34.28 10.93</th>
|
||
<th id="0B370473FF9E0034FCE3FE2BFCEE674D" box="[809,879,449,584]" gridcol="3" gridrow="0" pageId="5" pageNumber="424">HTYD VTYD</th>
|
||
<th id="0B370473FF9E0034FC04FE2BFBF7674D" box="[974,1142,449,584]" gridcol="4" gridrow="0" pageId="5" pageNumber="424">holotype NMBE 1061645 1.99 2.13</th>
|
||
<th id="0B370473FF9E0034FB7CFE2BFADF674D" box="[1206,1374,449,584]" gridcol="5" gridrow="0" pageId="5" pageNumber="424">paratype NMBE 1066153 2.01 2.26</th>
|
||
</tr>
|
||
<tr id="48E66D0FFF9E0034FF55FDB6FADF67C1" box="[159,1374,604,708]" gridrow="1" pageId="5" pageNumber="424">
|
||
<th id="0B370473FF9E0034FF55FDB6FF6967C1" box="[159,232,604,708]" gridcol="0" gridrow="1" pageId="5" pageNumber="424">AGD HL HWAE</th>
|
||
<td id="0B370473FF9E0034FE8FFDB6FE6F67C1" box="[325,494,604,708]" gridcol="1" gridrow="1" pageId="5" pageNumber="424">17.7 11.4 4.69</td>
|
||
<td id="0B370473FF9E0034FDF8FDB6FD5B67C1" box="[562,730,604,708]" gridcol="2" gridrow="1" pageId="5" pageNumber="424">16.4 11.36 4.9</td>
|
||
<td id="0B370473FF9E0034FCE3FDB6FCEE67C1" box="[809,879,604,708]" gridcol="3" gridrow="1" pageId="5" pageNumber="424">TBL THL HND</td>
|
||
<td id="0B370473FF9E0034FC04FDB6FBF767C1" box="[974,1142,604,708]" gridcol="4" gridrow="1" pageId="5" pageNumber="424">15.96 16.26 9.18</td>
|
||
<td id="0B370473FF9E0034FB7CFDB6FADF67C1" box="[1206,1374,604,708]" gridcol="5" gridrow="1" pageId="5" pageNumber="424">15.99 15.54 9.14</td>
|
||
</tr>
|
||
<tr id="48E66D0FFF9E0034FF55FD32FADF6645" box="[159,1374,728,832]" gridrow="2" pageId="5" pageNumber="424">
|
||
<th id="0B370473FF9E0034FF55FD32FF696645" box="[159,232,728,832]" gridcol="0" gridrow="2" pageId="5" pageNumber="424">HWAJ HD SL</th>
|
||
<td id="0B370473FF9E0034FE8FFD32FE6F6645" box="[325,494,728,832]" gridcol="1" gridrow="2" pageId="5" pageNumber="424">9.88 4.63 4.16</td>
|
||
<td id="0B370473FF9E0034FDF8FD32FD5B6645" box="[562,730,728,832]" gridcol="2" gridrow="2" pageId="5" pageNumber="424">10.1 4.46 4.23</td>
|
||
<td id="0B370473FF9E0034FCE3FD32FCEE6645" box="[809,879,728,832]" gridcol="3" gridrow="2" pageId="5" pageNumber="424">FOT TI TII</td>
|
||
<td id="0B370473FF9E0034FC04FD32FBF76645" box="[974,1142,728,832]" gridcol="4" gridrow="2" pageId="5" pageNumber="424">12.88 1.68 2.49</td>
|
||
<td id="0B370473FF9E0034FB7CFD32FADF6645" box="[1206,1374,728,832]" gridcol="5" gridrow="2" pageId="5" pageNumber="424">12.69 1.7 2.58</td>
|
||
</tr>
|
||
<tr id="48E66D0FFF9E0034FF55FCBEFADF66B9" box="[159,1374,852,956]" gridrow="3" pageId="5" pageNumber="424">
|
||
<th id="0B370473FF9E0034FF55FCBEFF6966B9" box="[159,232,852,956]" gridcol="0" gridrow="3" pageId="5" pageNumber="424">NS EN IN</th>
|
||
<td id="0B370473FF9E0034FE8FFCBEFE6F66B9" box="[325,494,852,956]" gridcol="1" gridrow="3" pageId="5" pageNumber="424">1.79 2.44 2.85</td>
|
||
<td id="0B370473FF9E0034FDF8FCBEFD5B66B9" box="[562,730,852,956]" gridcol="2" gridrow="3" pageId="5" pageNumber="424">1.48 2.49 2.51</td>
|
||
<td id="0B370473FF9E0034FCE3FCBEFCEE66B9" box="[809,879,852,956]" gridcol="3" gridrow="3" pageId="5" pageNumber="424">TIII TIV TV</td>
|
||
<td id="0B370473FF9E0034FC04FCBEFBF766B9" box="[974,1142,852,956]" gridcol="4" gridrow="3" pageId="5" pageNumber="424">3.79 7.3 4.76</td>
|
||
<td id="0B370473FF9E0034FB7CFCBEFADF66B9" box="[1206,1374,852,956]" gridcol="5" gridrow="3" pageId="5" pageNumber="424">4.0 7.53 4.81</td>
|
||
</tr>
|
||
<tr id="48E66D0FFF9E0034FF55FC3AFADF613D" box="[159,1374,976,1080]" gridrow="4" pageId="5" pageNumber="424">
|
||
<th id="0B370473FF9E0034FF55FC3AFF69613D" box="[159,232,976,1080]" gridcol="0" gridrow="4" pageId="5" pageNumber="424">ED UE IO</th>
|
||
<td id="0B370473FF9E0034FE8FFC3AFE6F613D" box="[325,494,976,1080]" gridcol="1" gridrow="4" pageId="5" pageNumber="424">3.73 4.27 4.1</td>
|
||
<td id="0B370473FF9E0034FDF8FC3AFD5B613D" box="[562,730,976,1080]" gridcol="2" gridrow="4" pageId="5" pageNumber="424">3.85 4.29 4.2</td>
|
||
<td id="0B370473FF9E0034FCE3FC3AFCEE613D" box="[809,879,976,1080]" gridcol="3" gridrow="4" pageId="5" pageNumber="424">FI FII FIII</td>
|
||
<td id="0B370473FF9E0034FC04FC3AFBF7613D" box="[974,1142,976,1080]" gridcol="4" gridrow="4" pageId="5" pageNumber="424">2.96 3.63 6.63</td>
|
||
<td id="0B370473FF9E0034FB7CFC3AFADF613D" box="[1206,1374,976,1080]" gridcol="5" gridrow="4" pageId="5" pageNumber="424">3.08 3.89 7.25</td>
|
||
</tr>
|
||
<tr id="48E66D0FFF9E0034FF55FBA6FADF6167" box="[159,1374,1100,1122]" gridrow="5" pageId="5" pageNumber="424">
|
||
<th id="0B370473FF9E0034FF55FBA6FF696167" box="[159,232,1100,1122]" gridcol="0" gridrow="5" pageId="5" pageNumber="424">TED</th>
|
||
<td id="0B370473FF9E0034FE8FFBA6FE6F6167" box="[325,494,1100,1122]" gridcol="1" gridrow="5" pageId="5" pageNumber="424">0.49</td>
|
||
<td id="0B370473FF9E0034FDF8FBA6FD5B6167" box="[562,730,1100,1122]" gridcol="2" gridrow="5" pageId="5" pageNumber="424">0.41</td>
|
||
<td id="0B370473FF9E0034FCE3FBA6FCEE6167" box="[809,879,1100,1122]" gridcol="3" gridrow="5" pageId="5" pageNumber="424">FIV</td>
|
||
<td id="0B370473FF9E0034FC04FBA6FBF76167" box="[974,1142,1100,1122]" gridcol="4" gridrow="5" pageId="5" pageNumber="424">4.31</td>
|
||
<td id="0B370473FF9E0034FB7CFBA6FADF6167" box="[1206,1374,1100,1122]" gridcol="5" gridrow="5" pageId="5" pageNumber="424">4.29</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="F6696F4DFF9EFFCEFF0DFB4BFD226097" blockId="5.[151,1436,1185,2038]" pageId="5" pageNumber="424">
|
||
Tips of fingers IV and V weakly expanded, spatulate, without distinct discs; first finger (II) much shorter than second (III), only reaching middle of second phalange of second finger; subarticular tubercles low; palm smooth with large, distinct rounded palmar tubercle. Tips of all toes rounded; fifth toe projecting farther than third; toes not fully webbed, first toe with one phalange free of web, second with 1½ phalanges free of web medially, third and fifth with 2½ phalanges free of web medially, fourth toe with 4 phalanges free of web medially; subarticular tubercles indistinct; two metatarsal tubercles, both oval and slightly raised (
|
||
<figureCitation id="6EED73C8FF9EFFCEFC29FABFFBA9606B" box="[995,1064,1365,1390]" captionStart="FIGURE 5" captionStartId="8.[151,250,1629,1651]" captionText="FIGURE 5. Foot of holotype of Ansonia vidua sp. nov. in ventral view." pageId="5" pageNumber="424">Fig. 5</figureCitation>
|
||
); no tarsal fold (
|
||
<figureCitation id="6EED73C8FF9EFFCEFB2AFABFFAA5606B" box="[1248,1316,1365,1390]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
). Left foot slightly deformed, toes shortened (
|
||
<figureCitation id="6EED73C8FF9EFFCEFDD1FA93FDDC6097" box="[539,605,1401,1426]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="F6696F4DFF9EFFCEFF0DFA77FF3B63B7" blockId="5.[151,1436,1185,2038]" pageId="5" pageNumber="424">
|
||
Skin of sides, back and dorsal surfaces of limbs uniformly covered with small, low, rounded tubercles; shagreened skin has a characteristic velvet-like appearance; tubercles not arranged in distinct rows, tubercles on top of head and eyelids smaller (
|
||
<figureCitation id="6EED73C8FF9EFFCEFE14FA0FFDCD60FB" box="[478,588,1509,1534]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="6.[151,250,1071,1093]" captionStartId-1="7.[151,250,1827,1849]" captionStartId-2="8.[151,250,699,721]" captionTargetBox-0="[151,1436,193,1049]" captionTargetBox-1="[151,1436,193,1805]" captionTargetId-0="figure@6.[151,1436,193,1050]" captionTargetId-1="figure@7.[151,1436,193,1805]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." captionText-1="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." captionText-2="FIGURE 4. Head of holotype of Ansonia vidua sp. nov. in lateral view." httpUri-0="https://zenodo.org/record/225973/files/figure.png" httpUri-1="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Figs. 2–4</figureCitation>
|
||
), slightly enlarged tubercles on scapular and temporal region and posterior of tympanum (
|
||
<figureCitation id="6EED73C8FF9EFFCEFE8AF9E3FE2E6327" box="[320,431,1545,1570]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="6.[151,250,1071,1093]" captionStartId-1="7.[151,250,1827,1849]" captionStartId-2="8.[151,250,699,721]" captionTargetBox-0="[151,1436,193,1049]" captionTargetBox-1="[151,1436,193,1805]" captionTargetId-0="figure@6.[151,1436,193,1050]" captionTargetId-1="figure@7.[151,1436,193,1805]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." captionText-1="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." captionText-2="FIGURE 4. Head of holotype of Ansonia vidua sp. nov. in lateral view." httpUri-0="https://zenodo.org/record/225973/files/figure.png" httpUri-1="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Figs. 2–4</figureCitation>
|
||
); one row of enlarged, spineless tubercles with darker pigmentation beneath mandibular symphysis (
|
||
<figureCitation id="6EED73C8FF9EFFCEFEEBF9C7FEE86343" box="[289,361,1581,1606]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
B), followed posteriorly by few smaller, spineless, irregularly arranged tubercles (
|
||
<figureCitation id="6EED73C8FF9EFFCEFAFBF9C7FAF96343" box="[1329,1400,1581,1606]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
B); tubercles on dorsal surfaces of head, body, and limbs tipped with one small, regularly shaped dark brown spine, tubercles on sides, venter and throat without spines (
|
||
<figureCitation id="6EED73C8FF9EFFCEFD20F99FFCAC638B" box="[746,813,1653,1678]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
A); no skin flaps on posterior thigh near vent (
|
||
<figureCitation id="6EED73C8FF9EFFCEFAF4F99FFA01638B" box="[1342,1408,1653,1678]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="F6696F4DFF9EFFCEFF0DF957FC866247" blockId="5.[151,1436,1185,2038]" pageId="5" pageNumber="424">
|
||
Colour in preservative uniformly dark brown above, slightly lighter areas on sides and on joints of limbs and phalanges; venter uniformly brownish grey, throat and chest uniformly yellowish brown (
|
||
<figureCitation id="6EED73C8FF9EFFCEFBB0F90BFB3A63FF" box="[1146,1211,1761,1786]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
A, B). Colour in life uniformly black-brown above, limbs dark brown with lighter areas on joints of the limbs and phalanges (
|
||
<figureCitation id="6EED73C8FF9EFFCEFA82F8EFFA0E621B" box="[1352,1423,1797,1822]" captionStart="FIGURE 2" captionStartId="6.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." httpUri="https://zenodo.org/record/225973/files/figure.png" pageId="5" pageNumber="424">Fig. 2</figureCitation>
|
||
). Iris black with small irregular intense red spots (
|
||
<figureCitation id="6EED73C8FF9EFFCEFD7FF8C3FD7B6247" box="[693,762,1833,1858]" captionStart="FIGURE 2" captionStartId="6.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Ansonia vidua sp. nov. (NMBE 1061645) in life." httpUri="https://zenodo.org/record/225973/files/figure.png" pageId="5" pageNumber="424">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F6696F4DFF9EFFCEFF0DF8A7FE1A62F3" blockId="5.[151,1436,1185,2038]" pageId="5" pageNumber="424">
|
||
<emphasis id="C4A2B35FFF9EFFCEFF0DF8A7FEBA6263" bold="true" box="[199,315,1869,1894]" pageId="5" pageNumber="424">Variation.</emphasis>
|
||
In the
|
||
<typeStatus id="296DD1EFFF9EFFCEFE44F8A4FE6F6263" box="[398,494,1870,1894]" pageId="5" pageNumber="424" type="paratype">paratype</typeStatus>
|
||
(
|
||
<tableCitation id="BB545AF6FF9EFFCEFE34F8A7FDD46263" box="[510,597,1869,1894]" captionStart="TABLE 2" captionStartId="5.[151,239,398,421]" captionTargetBox="[159,1374,449,1122]" captionTargetPageId="5" captionText="TABLE 2. Measurements of holotype and paratype of Ansonia vidua sp. nov., for details and abbreviations see text." httpUri="http://table.plazi.org/id/A2A93FC5FF9EFFCEFF5DFE64FAF164A0" pageId="5" pageNumber="424" tableUuid="A2A93FC5FF9EFFCEFF5DFE64FAF164A0">Table 2</tableCitation>
|
||
,
|
||
<figureCitation id="6EED73C8FF9EFFCEFDABF8A7FD226263" box="[609,675,1869,1894]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
C, D), skin on sides is slightly smoother, abdomen and throat finely granular, tubercles smaller and indistinct, only a single row of enlarged tubercles with darker pigmentation present posterior of the mandibular symphysis. Skin fold on rear thigh near vent is a preservation artifact (
|
||
<figureCitation id="6EED73C8FF9EFFCEFAC5F87FFAD562AB" box="[1295,1364,1941,1966]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
C, D). Lighter areas on sides larger in preservative, colour of throat changes from light yellowish grey to yellowish brown anteriorly (
|
||
<figureCitation id="6EED73C8FF9EFFCEFED9F837FED462F3" box="[275,341,2013,2038]" captionStart="FIGURE 3" captionStartId="7.[151,250,1827,1849]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@7.[151,1436,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Preserved type material of Ansonia vidua sp. nov.: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype." httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="5" pageNumber="424">Fig. 3</figureCitation>
|
||
C, D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A2A93FC5FF9DFFCDFF5DFBC5FC0B6140" box="[151,906,1071,1093]" httpUri="https://zenodo.org/record/225973/files/figure.png" pageId="6" pageNumber="425" targetBox="[151,1436,193,1049]" targetPageId="6">
|
||
<paragraph id="F6696F4DFF9DFFCDFF5DFBC5FC0B6140" blockId="6.[151,906,1071,1093]" box="[151,906,1071,1093]" pageId="6" pageNumber="425">
|
||
<emphasis id="C4A2B35FFF9DFFCDFF5DFBC5FE936140" bold="true" box="[151,274,1071,1093]" pageId="6" pageNumber="425">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="31D614CEFF9DFFCDFE53FBDAFDAB6140" box="[409,554,1071,1093]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF9DFFCDFE53FBDAFDAB6140" box="[409,554,1071,1093]" italics="true" pageId="6" pageNumber="425">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF9DFFCDFDFBFBC5FD006140" bold="true" box="[561,641,1071,1093]" pageId="6" pageNumber="425">
|
||
<taxonomicNameLabel id="DF910E24FF9DFFCDFDFBFBC5FD006140" box="[561,641,1071,1093]" pageId="6" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMBE 1061645) in life.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="BECC3CC6FF9DFFC2FF0DFB86FB65675A" lastPageId="9" lastPageNumber="428" pageId="6" pageNumber="425" type="reference_group">
|
||
<paragraph id="F6696F4DFF9DFFC2FF0DFB86FD6F64A9" blockId="6.[151,1437,1132,2021]" lastBlockId="9.[151,1437,151,1149]" lastPageId="9" lastPageNumber="428" pageId="6" pageNumber="425">
|
||
<emphasis id="C4A2B35FFF9DFFCDFF0DFB86FEE46180" bold="true" box="[199,357,1132,1157]" pageId="6" pageNumber="425">Comparisons</emphasis>
|
||
<taxonomicName id="31D614CEFF9DFFCDFEA7FB87FD8F6180" ID-CoL="66Y9Q" box="[365,526,1133,1157]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF9DFFCDFEA7FB87FD8F6180" box="[365,526,1133,1157]" italics="true" pageId="6" pageNumber="425">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF9DFFCDFDDFFB86FDEE6180" bold="true" box="[533,623,1132,1157]" pageId="6" pageNumber="425">
|
||
<taxonomicNameLabel id="DF910E24FF9DFFCDFDDFFB86FDEE6180" box="[533,623,1132,1157]" pageId="6" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished morphologically from the 12 described congeneric Bornean species on the basis of an apomorphic combination of diagnostic characters (see ‘Diagnosis’). Bornean congeners differ from
|
||
<taxonomicName id="31D614CEFF9DFFCDFE6BFB5FFD8261C8" ID-CoL="66Y9Q" box="[417,515,1205,1229]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="vidua">
|
||
<emphasis id="C4A2B35FFF9DFFCDFE6BFB5FFD8261C8" box="[417,515,1205,1229]" italics="true" pageId="6" pageNumber="425">A. vidua</emphasis>
|
||
</taxonomicName>
|
||
as follows:
|
||
<taxonomicName id="31D614CEFF9DFFCDFD51FB5FFC6E61C8" ID-CoL="5V9WG" authority="Inger, 1960" authorityName="Inger" authorityYear="1960" box="[667,1007,1204,1229]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="albomaculata">
|
||
<emphasis id="C4A2B35FFF9DFFCDFD51FB5FFD2D61C9" box="[667,684,1205,1228]" italics="true" pageId="6" pageNumber="425">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF9DFFCDFD75FB5FFCD861C8" box="[703,857,1205,1229]" italics="true" pageId="6" pageNumber="425">albomaculata</emphasis>
|
||
<bibRefCitation id="924712BCFF9DFFCDFCACFB5FFC6E61C8" author="Inger" box="[870,1007,1204,1229]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1960) A review of the Oriental toads of the genus Ansonia Stoliczka. Fieldiana: Zoology, 39, 473 - 503." type="journal article" year="1960">Inger, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
: interorbital ridges absent; tympanic annulum partially hidden by thickened skin; sharp tarsal ridge present; toes almost fully webbed; light band running from the posterior corner of the eye to the axilla, often with a subocular light spot (
|
||
<bibRefCitation id="924712BCFF9DFFCDFB7EFB17FAB46010" author="Inger" box="[1204,1333,1276,1301]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFA88FB17FEBF603C" author="Inger & Stuebing" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF9DFFCDFE99FACBFE53603D" ID-CoL="679WG" box="[339,466,1312,1336]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="echinata">
|
||
<emphasis id="C4A2B35FFF9DFFCDFE99FACBFE53603D" box="[339,466,1312,1336]" italics="true" pageId="6" pageNumber="425">A. echinata</emphasis>
|
||
</taxonomicName>
|
||
: interorbital ridges absent; dorsum with rounded tubercles both small and large; large ventrolateral tubercles (up to half diameter of tympanum) tipped with small black spines; spinose tubercles on top of snout; third and fifth toe with 1–2 phalanges free of web; limbs with dark crossbars (
|
||
<bibRefCitation id="924712BCFF9DFFCDFBA3FA83FAF06084" author="Inger & Stuebing" box="[1129,1393,1384,1409]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Stuebing, R. B. (2009) New species and new records of Bornean frogs (Amphibia: Anura). Raffles Bulletin of Zoology, 57, 527 - 535." type="journal article" year="2009">Inger & Stuebing 2009</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF9DFFCDFA4EFA83FF7A60A0" ID-CoL="66Y8H" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="fuliginea">
|
||
<emphasis id="C4A2B35FFF9DFFCDFA4EFA83FA146085" box="[1412,1429,1385,1408]" italics="true" pageId="6" pageNumber="425">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF9DFFCDFF5DFA67FF7A60A0" box="[151,251,1421,1445]" italics="true" pageId="6" pageNumber="425">fuliginea</emphasis>
|
||
</taxonomicName>
|
||
: larger body size (females
|
||
<quantity id="312EC2A8FF9DFFCDFD83FA67FD4D60A0" box="[585,716,1421,1445]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="6" pageNumber="425" unit="mm" value="41.0" valueMax="44.0" valueMin="38.0">38–44 mm</quantity>
|
||
); interorbital ridges absent; snout vertical in profile and not projecting; tympanum ca. 50% of eye diameter; submandibular spines arranged in a double row (
|
||
<bibRefCitation id="924712BCFF9DFFCDFB19FA5BFAD060CC" author="Inger" box="[1235,1361,1456,1481]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1960) A review of the Oriental toads of the genus Ansonia Stoliczka. Fieldiana: Zoology, 39, 473 - 503." type="journal article" year="1960">Inger 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="924712BCFF9DFFCDFA96FA5AFA1660CC" author="Inger" box="[1372,1431,1456,1481]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFF5DFA3FFE1E60E8" author="Inger & Stuebing" box="[151,415,1492,1517]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF9DFFCDFE72FA3FFD2F60E8" ID-CoL="66XWM" authority="Inger, 1966" authorityName="Inger" authorityYear="1966" box="[440,686,1492,1517]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="C4A2B35FFF9DFFCDFE72FA3FFE4860E9" box="[440,457,1493,1516]" italics="true" pageId="6" pageNumber="425">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF9DFFCDFE13FA3FFD9E60E8" box="[473,543,1493,1517]" italics="true" pageId="6" pageNumber="425">guibei</emphasis>
|
||
<bibRefCitation id="924712BCFF9DFFCDFDE2FA3FFD2F60E8" author="Inger" box="[552,686,1492,1517]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
: interorbital ridges absent; interorbital distance much greater than width of upper eyelid; tympanum diameter 25–33% of eye diameter; first finger reaches tip of second finger when adpressed; web reaches tips of first, second, third, and fifth toes; dorsal and dorsolateral warts topped with clusters of dark spinules; oblique flap of skin present on each side of vent; upper surfaces dark brown with irregular lighter spots on top of head and back, underside light grey with indistinct dark spots (
|
||
<bibRefCitation id="924712BCFF9DFFCDFC35F98FFBFE6378" author="Inger" box="[1023,1151,1636,1661]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFB43F98FFA0A6378" author="Inger & Stuebing" box="[1161,1419,1636,1661]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF9DFFCDFF5DF963FE9C63A5" ID-CoL="66Y8M" box="[151,285,1672,1696]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="hanitschi">
|
||
<emphasis id="C4A2B35FFF9DFFCDFF5DF963FF2963A5" box="[151,168,1673,1696]" italics="true" pageId="6" pageNumber="425">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF9DFFCDFF7DF962FE9C63A5" box="[183,285,1672,1696]" italics="true" pageId="6" pageNumber="425">hanitschi</emphasis>
|
||
</taxonomicName>
|
||
: body flattened; narrower head (HWAJ/TBL 0.50–0.56); interorbital ridges absent; third and fifth toe with 1.5–2 phalanges free of web; skin on back with heterogeneous small rounded warts and a few oval warts, a few warts form short, slender ridges on the back; dark greenish-grey to reddish-brown colouration with dorsal markings, limbs with black crossbars, venter with pattern of dark blotches(
|
||
<bibRefCitation id="924712BCFF9DFFCDFBDBF91EFB1A6208" author="Dring" box="[1041,1179,1780,1805]" firstAuthor="Dring" pageId="6" pageNumber="425" refString="Dring, J. C. M. (1983) Some new frogs from Sarawak. Amphibia-Reptilia, 4, 103 - 115." type="journal article" year="1983">Dring 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFB61F91FFAAE6208" author="Inger" box="[1195,1327,1780,1805]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFAF5F91FFEBD6234" author="Inger & Stuebing" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF9DFFCDFE9AF8F3FE4E6235" ID-CoL="5V9WK" box="[336,463,1816,1840]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="latidisca">
|
||
<emphasis id="C4A2B35FFF9DFFCDFE9AF8F3FEE16235" box="[336,352,1817,1840]" italics="true" pageId="6" pageNumber="425">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF9DFFCDFEA5F8F2FE4E6235" box="[367,463,1816,1840]" italics="true" pageId="6" pageNumber="425">latidisca</emphasis>
|
||
</taxonomicName>
|
||
: larger body size (females
|
||
<quantity id="312EC2A8FF9DFFCDFD37F8F2FC226234" box="[765,931,1816,1841]" metricMagnitude="-2" metricUnit="m" metricValue="5.595" metricValueMax="5.9799999999999995" metricValueMin="5.21" pageId="6" pageNumber="425" unit="mm" value="55.95" valueMax="59.8" valueMin="52.1">52.1–59.8 mm</quantity>
|
||
); canthus rostralis sharp, straight; interorbital ridges absent, but two rows of conical interorbital tubercles; tympanum diameter half of eye; tips of fingers dilated into truncate discs (that of third finger equal in width to tympanum); green colouration above with a distinct pattern of irregular brown spots, abdomen brown with scattered yellowish spots (
|
||
<bibRefCitation id="924712BCFF9DFFCDFC30F86FFBFC6298" author="Inger" box="[1018,1149,1924,1949]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFB47F86EFAE66298" author="Matsui" box="[1165,1383,1924,1949]" etAl="et al." firstAuthor="Matsui" pageId="6" pageNumber="425" refString="Matsui, M., Nishikawa, K., Yeo, S. T. & Eto, K. (2012) Notes on a rare Bornean bufonid Ansonia latidisca Inger, 1966, with special reference to its phylogenetic position. Current Herpetology, 31, 87 - 96." type="journal article" year="2012">
|
||
Matsui
|
||
<emphasis id="C4A2B35FFF9DFFCDFB2CF86FFAA26298" box="[1254,1315,1925,1949]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF9DFFCDFA4EF86FFF7762C5" ID-CoL="5V9WL" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="leptopus">
|
||
<emphasis id="C4A2B35FFF9DFFCDFA4EF86FFA146299" box="[1412,1429,1925,1948]" italics="true" pageId="6" pageNumber="425">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF9DFFCDFF5DF842FF7762C5" box="[151,246,1960,1984]" italics="true" pageId="6" pageNumber="425">leptopus</emphasis>
|
||
</taxonomicName>
|
||
: larger body size (females to
|
||
<quantity id="312EC2A8FF9DFFCDFD84F842FD2262C4" box="[590,675,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="425" unit="mm" value="60.0">60 mm</quantity>
|
||
); interorbital ridge absent; tip of first finger reaches swollen tip of second; chest and belly grey with darker mottling (
|
||
<bibRefCitation id="924712BCFF9DFFCDFD39F827FCF362E0" author="Inger" box="[755,882,1996,2021]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFCB4F827FBE162E0" author="Inger & Dring" box="[894,1120,1996,2021]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Dring, J. C. M. (1988) Taxonomic and ecological relations of Bornean stream toads allied to Ansonia leptopus (Gunther) (Anura: Bufonidae). Malayan Nature Journal, 41, 461 - 471." type="journal article" year="1988">Inger & Dring 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF9DFFCDFBA1F827FAF062E0" author="Inger & Stuebing" box="[1131,1393,1996,2021]" firstAuthor="Inger" pageId="6" pageNumber="425" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF93FFC3FF5DF94EFEB263B9" ID-CoL="66XWL" box="[151,307,1700,1724]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="longidigita">
|
||
<emphasis id="C4A2B35FFF93FFC3FF5DF94EFF2963BE" box="[151,168,1700,1723]" italics="true" pageId="8" pageNumber="427">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF93FFC3FF72F94EFEB263B9" box="[184,307,1700,1724]" italics="true" pageId="8" pageNumber="427">longidigita</emphasis>
|
||
</taxonomicName>
|
||
: much larger size (females to
|
||
<quantity id="312EC2A8FF93FFC3FD58F949FD6663B9" box="[658,743,1699,1724]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="427" unit="mm" value="65.0">65 mm</quantity>
|
||
); canthus rostralis sharply angular; tip of first finger reaches swollen tip of second; third and fifth toe with one to two phalanges free of web; abdomen with faint dark spots on a pale background or dark with obscure pale spots, limbs with obscure light crossbars (
|
||
<bibRefCitation id="924712BCFF93FFC3FBA3F906FB6B6201" author="Inger" box="[1129,1258,1771,1796]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">Inger 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF93FFC3FB32F906FF53622D" author="Inger & Dring" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. & Dring, J. C. M. (1988) Taxonomic and ecological relations of Bornean stream toads allied to Ansonia leptopus (Gunther) (Anura: Bufonidae). Malayan Nature Journal, 41, 461 - 471." type="journal article" year="1988">Inger & Dring 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF93FFC3FF14F8FAFE65622D" author="Inger & Stuebing" box="[222,484,1807,1832]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF93FFC3FE32F8FAFDE46222" ID-CoL="66XXR" box="[504,613,1808,1831]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="minuta">
|
||
<emphasis id="C4A2B35FFF93FFC3FE32F8FAFD886222" box="[504,521,1808,1831]" italics="true" pageId="8" pageNumber="427">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF93FFC3FDDDF8FAFDE46222" box="[535,613,1808,1831]" italics="true" pageId="8" pageNumber="427">minuta</emphasis>
|
||
</taxonomicName>
|
||
: smaller size (females
|
||
<quantity id="312EC2A8FF93FFC3FCA2F8FAFC67622D" box="[872,998,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="8" pageNumber="427" unit="mm" value="25.5" valueMax="28.0" valueMin="23.0">23–28 mm</quantity>
|
||
); interorbital ridges absent; interorbital distance 1.3 width of upper eyelid; distinct tarsal ridge; toes are three-fourth webbed in females; dorsum brown with irregular small orange or yellowish dots, side of head with dark and light bars with a whitish area below eye, abdomen pale with black spots (
|
||
<bibRefCitation id="924712BCFF93FFC3FDC1F896FD0A6291" author="Inger" box="[523,651,1915,1940]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. (1960) A review of the Oriental toads of the genus Ansonia Stoliczka. Fieldiana: Zoology, 39, 473 - 503." type="journal article" year="1960">Inger 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="924712BCFF93FFC3FD53F891FD556291" author="Inger" box="[665,724,1915,1940]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF93FFC3FD28F896FC6E6291" author="Inger & Stuebing" box="[738,1007,1915,1940]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF93FFC3FBCFF896FB186291" ID-CoL="66XXS" box="[1029,1177,1916,1940]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="platysoma">
|
||
<emphasis id="C4A2B35FFF93FFC3FBCFF896FB976296" box="[1029,1046,1916,1939]" italics="true" pageId="8" pageNumber="427">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF93FFC3FBECF896FB186291" box="[1062,1177,1916,1940]" italics="true" pageId="8" pageNumber="427">platysoma</emphasis>
|
||
</taxonomicName>
|
||
: smaller size (females
|
||
<quantity id="312EC2A8FF93FFC3FF5DF875FE9762BD" box="[151,278,1951,1976]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="8" pageNumber="427" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20–25 mm</quantity>
|
||
); head and body flattened; interorbital ridges absent; skin with heterogeneous warts; 1–1.5 phalanges free of web on third toe; abdomen with light and dark mottling; limbs with light crossbars (
|
||
<bibRefCitation id="924712BCFF93FFC3FB01F82EFACF62D9" author="Inger" box="[1227,1358,1987,2012]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. (1960) A review of the Oriental toads of the genus Ansonia Stoliczka. Fieldiana: Zoology, 39, 473 - 503." type="journal article" year="1960">Inger 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="924712BCFF93FFC3FA96F829FA1662D9" author="Inger" box="[1372,1431,1987,2012]" firstAuthor="Inger" pageId="8" pageNumber="427" refString="Inger, R. F. (1966) The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology, 52, 1 - 402." type="journal article" year="1966">1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF92FFC2FF5DFF7DFE0465B5" author="Malkmus" box="[151,389,151,176]" etAl="et al." firstAuthor="Malkmus" pageId="9" pageNumber="428" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. & Kosuch, J. (2002) Amphibians & Reptiles of Mount Kinabalu (North Borneo). A. R. G. Gantner Verlag, Ruggell, 424 pp." type="book" year="2002">
|
||
Malkmus
|
||
<emphasis id="C4A2B35FFF92FFC2FEC2FF73FEC065B5" box="[264,321,152,176]" italics="true" pageId="9" pageNumber="428">et al.</emphasis>
|
||
2002
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF92FFC2FE53FF72FDA565B5" ID-CoL="66XXP" box="[409,548,152,176]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="spinulifer">
|
||
<emphasis id="C4A2B35FFF92FFC2FE53FF72FE2B65AA" box="[409,426,152,175]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FE72FF72FDA565B5" box="[440,548,152,176]" italics="true" pageId="9" pageNumber="428">spinulifer</emphasis>
|
||
</taxonomicName>
|
||
: larger body size (females to
|
||
<quantity id="312EC2A8FF92FFC2FCB9FF7DFC7165B5" box="[883,1008,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="9" pageNumber="428" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40–45 mm</quantity>
|
||
); interorbital ridges absent; tip of first finger reaches swollen tip of second; dorsal and dorsolateral warts elevated, large and tipped with one to three substantial black spines; top of head and body black with a yellowish or whitish oval patch between shoulders and a light streak along flanks, venter black with small cream-coloured spots or black marbling (
|
||
<bibRefCitation id="924712BCFF92FFC2FB7AFEEEFA166419" author="Inger & Dring" box="[1200,1431,260,285]" firstAuthor="Inger" pageId="9" pageNumber="428" refString="Inger, R. F. & Dring, J. C. M. (1988) Taxonomic and ecological relations of Bornean stream toads allied to Ansonia leptopus (Gunther) (Anura: Bufonidae). Malayan Nature Journal, 41, 461 - 471." type="journal article" year="1988">Inger & Dring 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF92FFC2FF5DFEC2FE2D6445" author="Inger & Stuebing" box="[151,428,295,320]" firstAuthor="Inger" pageId="9" pageNumber="428" refString="Inger, R. F. & Stuebing, R. B. (2005) A Field Guide to the Frogs of Borneo. Second Edition. Natural History Publications (Borneo), Sdn. Bhd., Kota Kinabalu, viii + 201 pp." type="book" year="2005">Inger & Stuebing 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="31D614CEFF92FFC2FE0FFEC2FD646445" ID-CoL="66XY2" authority="Dring, 1983" authorityName="Dring" authorityYear="1983" box="[453,741,295,320]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="torrentis">
|
||
<emphasis id="C4A2B35FFF92FFC2FE0FFEC2FE57643A" box="[453,470,296,319]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FE23FEC2FDC9643A" box="[489,584,296,319]" italics="true" pageId="9" pageNumber="428">torrentis</emphasis>
|
||
<bibRefCitation id="924712BCFF92FFC2FD9FFECDFD646445" author="Dring" box="[597,741,295,320]" firstAuthor="Dring" pageId="9" pageNumber="428" refString="Dring, J. C. M. (1983) Some new frogs from Sarawak. Amphibia-Reptilia, 4, 103 - 115." type="journal article" year="1983">Dring, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
: interorbital ridges absent; dorsum covered with numerous heterogeneous rounded tubercles; tips of tubercles without spines, except on temporal and scapular region; blackish in colour with yellowish to pale brown markings on flanks, limbs with light crossbars, abdomen with large greybrown blotches separated by pale areas (
|
||
<bibRefCitation id="924712BCFF92FFC2FD91FE7EFD6064A9" author="Dring" box="[603,737,404,429]" firstAuthor="Dring" pageId="9" pageNumber="428" refString="Dring, J. C. M. (1983) Some new frogs from Sarawak. Amphibia-Reptilia, 4, 103 - 115." type="journal article" year="1983">Dring 1983</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A2A93FC5FF9CFFCCFF5DF8C9FB8C625D" httpUri="https://zenodo.org/record/225974/files/figure.png" pageId="7" pageNumber="426" targetBox="[151,1436,193,1805]" targetPageId="7">
|
||
<paragraph id="F6696F4DFF9CFFCCFF5DF8C9FB8C625D" blockId="7.[151,1436,1827,1880]" pageId="7" pageNumber="426">
|
||
<emphasis id="C4A2B35FFF9CFFCCFF5DF8C9FE92623C" bold="true" box="[151,275,1827,1849]" pageId="7" pageNumber="426">FIGURE 3.</emphasis>
|
||
Preserved type material of
|
||
<taxonomicName id="31D614CEFF9CFFCCFDE1F8CEFD3C623C" ID-CoL="66Y9Q" box="[555,701,1827,1849]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF9CFFCCFDE1F8CEFD3C623C" box="[555,701,1827,1849]" italics="true" pageId="7" pageNumber="426">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF9CFFCCFD0EF8C9FC95623C" bold="true" box="[708,788,1827,1849]" pageId="7" pageNumber="426">
|
||
<taxonomicNameLabel id="DF910E24FF9CFFCCFD0EF8C9FC95623C" box="[708,788,1827,1849]" pageId="7" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A. dorsal view of holotype (NMBE 1061645); B. ventral view of holotype; C. dorsal view of paratype (NMBE 1066153); D. ventral view of paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A2A93FC5FF93FFC3FF5DFD51FCF167D4" box="[151,880,699,721]" pageId="8" pageNumber="427" startId="8.[151,250,699,721]">
|
||
<paragraph id="F6696F4DFF93FFC3FF5DFD51FCF167D4" blockId="8.[151,880,699,721]" box="[151,880,699,721]" pageId="8" pageNumber="427">
|
||
<emphasis id="C4A2B35FFF93FFC3FF5DFD51FE9367D4" bold="true" box="[151,274,699,721]" pageId="8" pageNumber="427">FIGURE 4.</emphasis>
|
||
Head of holotype of
|
||
<taxonomicName id="31D614CEFF93FFC3FE20FD56FDFD67D4" ID-CoL="66Y9Q" box="[490,636,699,721]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF93FFC3FE20FD56FDFD67D4" box="[490,636,699,721]" italics="true" pageId="8" pageNumber="427">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF93FFC3FD48FD51FD5267D4" bold="true" box="[642,723,699,721]" pageId="8" pageNumber="427">
|
||
<taxonomicNameLabel id="DF910E24FF93FFC3FD48FD51FD5267D4" box="[642,723,699,721]" pageId="8" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A2A93FC5FF93FFC3FF5DF9B7FCF16376" box="[151,880,1629,1651]" pageId="8" pageNumber="427">
|
||
<paragraph id="F6696F4DFF93FFC3FF5DF9B7FCF16376" blockId="8.[151,880,1629,1651]" box="[151,880,1629,1651]" pageId="8" pageNumber="427">
|
||
<emphasis id="C4A2B35FFF93FFC3FF5DF9B7FE936376" bold="true" box="[151,274,1629,1651]" pageId="8" pageNumber="427">FIGURE 5.</emphasis>
|
||
Foot of holotype of
|
||
<taxonomicName id="31D614CEFF93FFC3FE29F9B4FDF46376" ID-CoL="66Y9Q" box="[483,629,1629,1651]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF93FFC3FE29F9B4FDF46376" box="[483,629,1629,1651]" italics="true" pageId="8" pageNumber="427">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF93FFC3FDB1F9B7FD4D6376" bold="true" box="[635,716,1629,1651]" pageId="8" pageNumber="427">
|
||
<taxonomicNameLabel id="DF910E24FF93FFC3FDB1F9B7FD4D6376" box="[635,716,1629,1651]" pageId="8" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F6696F4DFF92FFC2FF0DFE52FB65675A" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="428">
|
||
<taxonomicName id="31D614CEFF92FFC2FF0DFE52FEEB64D5" ID-CoL="66Y9Q" box="[199,362,440,464]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF92FFC2FF0DFE52FEEB64D5" box="[199,362,440,464]" italics="true" pageId="9" pageNumber="428">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF92FFC2FEBEFE5DFE5164D5" bold="true" box="[372,464,439,464]" pageId="9" pageNumber="428">
|
||
<taxonomicNameLabel id="DF910E24FF92FFC2FEBEFE5DFE5164D5" box="[372,464,439,464]" pageId="9" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unambiguously separated from
|
||
<taxonomicName id="31D614CEFF92FFC2FCAAFE52FC7F64D5" ID-CoL="66XWH" box="[864,1022,440,464]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="endauensis">
|
||
<emphasis id="C4A2B35FFF92FFC2FCAAFE52FCF064CA" box="[864,881,440,463]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FC48FE52FC7F64D5" box="[898,1022,440,464]" italics="true" pageId="9" pageNumber="428">endauensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FBC5FE52FB1564D5" ID-CoL="679WM" box="[1039,1172,440,464]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="inthanon">
|
||
<emphasis id="C4A2B35FFF92FFC2FBC5FE52FBA164CA" box="[1039,1056,440,463]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FBFAFE52FB1564D5" box="[1072,1172,440,464]" italics="true" pageId="9" pageNumber="428">inthanon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FB6FFE52FAF264D5" ID-CoL="5V9WN" box="[1189,1395,440,464]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="jeetsukumarani">
|
||
<emphasis id="C4A2B35FFF92FFC2FB6FFE52FB3764CA" box="[1189,1206,440,463]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FB0CFE52FAF264D5" box="[1222,1395,440,464]" italics="true" pageId="9" pageNumber="428">jeetsukumarani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FA4EFE52FF7564F1" ID-CoL="679WN" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="kraensis">
|
||
<emphasis id="C4A2B35FFF92FFC2FA4EFE52FA1464CA" box="[1412,1429,440,463]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FF5DFE36FF7564F1" box="[151,244,476,500]" italics="true" pageId="9" pageNumber="428">kraensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FECCFE36FEE764F1" ID-CoL="66XWK" box="[262,358,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="latiffi">
|
||
<emphasis id="C4A2B35FFF92FFC2FECCFE36FE9664F6" box="[262,279,476,499]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FEE3FE36FEE764F1" box="[297,358,476,500]" italics="true" pageId="9" pageNumber="428">latiffi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FEB2FE36FD8564F1" ID-CoL="679WK" box="[376,516,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="latirostra">
|
||
<emphasis id="C4A2B35FFF92FFC2FEB2FE36FE0864F6" box="[376,393,476,499]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FE51FE36FD8564F1" box="[411,516,476,500]" italics="true" pageId="9" pageNumber="428">latirostra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FDDCFE36FD2664F1" ID-CoL="679WL" box="[534,679,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="malayana">
|
||
<emphasis id="C4A2B35FFF92FFC2FDDCFE36FDA664F6" box="[534,551,476,499]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FDF3FE36FD2664F1" box="[569,679,476,500]" italics="true" pageId="9" pageNumber="428">malayana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FD70FE36FCD264F1" ID-CoL="5V9XR" box="[698,851,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="mcgregori">
|
||
<emphasis id="C4A2B35FFF92FFC2FD70FE36FD4A64F6" box="[698,715,476,499]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FD17FE37FCD264F1" box="[733,851,477,500]" italics="true" pageId="9" pageNumber="428">mcgregori</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FCA9FE36FC6264F1" ID-CoL="66XY3" box="[867,995,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="muelleri">
|
||
A.
|
||
<emphasis id="C4A2B35FFF92FFC2FC4CFE36FC6264F1" box="[902,995,476,500]" italics="true" pageId="9" pageNumber="428">muelleri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="C4A2B35FFF92FFC2FC3FFE36FB4764F6" box="[1013,1222,476,500]" italics="true" pageId="9" pageNumber="428">
|
||
<taxonomicName id="31D614CEFF92FFC2FC3FFE36FB2764F1" ID-CoL="679Y3" box="[1013,1190,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="penangensis">A. penangensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FB7FFE36FB4764F6" ID-CoL="679Y4" box="[1205,1222,476,499]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="siamensis">A</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FB12FE37FAC464F1" box="[1240,1349,477,500]" italics="true" pageId="9" pageNumber="428">siamensis</emphasis>
|
||
and
|
||
<taxonomicName id="31D614CEFF92FFC2FA4EFE36FE866712" ID-CoL="66XXQ" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="tiomanica">
|
||
<emphasis id="C4A2B35FFF92FFC2FA4EFE36FA1464F6" box="[1412,1429,476,499]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FF5DFDEAFE866712" box="[151,263,512,535]" italics="true" pageId="9" pageNumber="428">tiomanica</emphasis>
|
||
</taxonomicName>
|
||
by phylogenetic signal of sequence data (
|
||
<figureCitation id="6EED73C8FF92FFC2FD29FE15FCA7671D" box="[739,806,511,536]" captionStart="FIGURE 1" captionStartId="4.[151,250,1946,1968]" captionTargetBox="[150,1414,193,1895]" captionTargetId="figure@4.[150,1414,193,1895]" captionTargetPageId="4" captionText="FIGURE 1. Phylogenetic relationships of Ansonia vidua sp. nov. according to a RaxML-based Maximum Likelihood analysis of partial sequences of the mitochondrial 16 S rRNA gene. Node labels above branches show the bootstrapping support of ML analysis (only values> 50), below branches the posterior probability values of BI (only values> 70)." httpUri="https://zenodo.org/record/225972/files/figure.png" pageId="9" pageNumber="428">Fig. 1</figureCitation>
|
||
). The recently described species
|
||
<taxonomicName id="31D614CEFF92FFC2FB6BFDEAFE5A6738" ID-CoL="66Y8L" authority="Chan, Wood, Anuar, Muin, Quah, Sumarli" authorityName="Chan, Wood, Anuar, Muin, Quah, Sumarli" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="lumut">
|
||
<emphasis id="C4A2B35FFF92FFC2FB6BFDEAFB336712" box="[1185,1202,512,535]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FB0BFDEAFA80671D" box="[1217,1281,512,536]" italics="true" pageId="9" pageNumber="428">lumut</emphasis>
|
||
Chan, Wood, Anuar, Muin, Quah, Sumarli
|
||
</taxonomicName>
|
||
, and Grismer is the sister taxon to a clade consisting of
|
||
<taxonomicName id="31D614CEFF92FFC2FB81FDCEFA856739" box="[1099,1284,548,572]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="jeetskumaran">
|
||
<emphasis id="C4A2B35FFF92FFC2FB81FDCEFBDD673E" box="[1099,1116,548,571]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FBA0FDCEFA856739" box="[1130,1284,548,572]" italics="true" pageId="9" pageNumber="428">jeetskumaran</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FAC4FDCEFA1A6739" ID-CoL="679WL" box="[1294,1435,548,572]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="malayana">
|
||
A.
|
||
<emphasis id="C4A2B35FFF92FFC2FAE7FDCEFA1A6739" box="[1325,1435,548,572]" italics="true" pageId="9" pageNumber="428">malayana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="31D614CEFF92FFC2FF0CFDA2FDC26765" ID-CoL="679Y3" authority="Chan et al. 2014" authorityName="Chan et al." authorityYear="2014" box="[198,579,583,608]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="penangensis">
|
||
<emphasis id="C4A2B35FFF92FFC2FF0CFDA2FEEE675A" box="[198,367,584,607]" italics="true" pageId="9" pageNumber="428">A. penangensis</emphasis>
|
||
(
|
||
<bibRefCitation id="924712BCFF92FFC2FEB4FDADFDBA6765" author="Chan" box="[382,571,583,608]" etAl="et al." firstAuthor="Chan" pageId="9" pageNumber="428" refString="Chan, K. O., Wood, P. L., Anuar, S. M. S., Muin, M. A., Quah, E. S. H., Sumarli, A. X. Y. & Grismer, L. L. (2014) A new species of upland stream toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from northeastern peninsular Malaysia. Zootaxa, 3764 (4), 427 - 440. http: // dx. doi. org / 10.11646 / zootaxa. 3764.4.3" type="journal article" year="2014">
|
||
Chan
|
||
<emphasis id="C4A2B35FFF92FFC2FE08FDA3FE7A6765" box="[450,507,584,608]" italics="true" pageId="9" pageNumber="428">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and, therefore, also not directly related to
|
||
<taxonomicName id="31D614CEFF92FFC2FBD4FDA2FBFA6765" ID-CoL="66Y9Q" box="[1054,1147,584,608]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF92FFC2FBD4FDA2FBAE675A" box="[1054,1071,584,607]" italics="true" pageId="9" pageNumber="428">A</emphasis>
|
||
.
|
||
<emphasis id="C4A2B35FFF92FFC2FBF7FDA2FBFA6765" box="[1085,1147,584,608]" italics="true" pageId="9" pageNumber="428">vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF92FFC2FB49FDADFB5D6765" bold="true" box="[1155,1244,583,608]" pageId="9" pageNumber="428">
|
||
<taxonomicNameLabel id="DF910E24FF92FFC2FB49FDADFB5D6765" box="[1155,1244,583,608]" pageId="9" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BECC3CC6FF92FFC2FF0DFD86FAEF6179" pageId="9" pageNumber="428" type="etymology">
|
||
<paragraph id="F6696F4DFF92FFC2FF0DFD86FD0867C8" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="428">
|
||
<emphasis id="C4A2B35FFF92FFC2FF0DFD86FECF6780" bold="true" box="[199,334,620,645]" pageId="9" pageNumber="428">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="31D614CEFF92FFC2FDEDFD86FDE46781" box="[551,613,620,644]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="vidua">
|
||
<emphasis id="C4A2B35FFF92FFC2FDEDFD86FDE46781" box="[551,613,620,644]" italics="true" pageId="9" pageNumber="428">vidua</emphasis>
|
||
</taxonomicName>
|
||
means´widow´and refers to the uniform brownish-black colouration in life of this species, which is traditionally the colour of choice by widows. We suggest the English vernacular name ‘Murud Black Slender Toad’ for the species.
|
||
</paragraph>
|
||
<paragraph id="F6696F4DFF92FFC2FF0DFD3DFAEF6179" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="428">
|
||
<emphasis id="C4A2B35FFF92FFC2FF0DFD3DFE1567F5" bold="true" box="[199,404,727,752]" pageId="9" pageNumber="428">Ecological notes.</emphasis>
|
||
The
|
||
<typeStatus id="296DD1EFFF92FFC2FE13FD32FD8867F5" box="[473,521,728,752]" pageId="9" pageNumber="428">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="31D614CEFF92FFC2FD4DFD32FCAC67F5" box="[647,813,728,752]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="vidua" status="sp. nov.">
|
||
<emphasis id="C4A2B35FFF92FFC2FD4DFD32FCAC67F5" box="[647,813,728,752]" italics="true" pageId="9" pageNumber="428">Ansonia vidua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C4A2B35FFF92FFC2FCF0FD3DFC1B67F5" bold="true" box="[826,922,727,752]" pageId="9" pageNumber="428">
|
||
<taxonomicNameLabel id="DF910E24FF92FFC2FCF0FD3DFC1B67F5" box="[826,922,727,752]" pageId="9" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, consisting of
|
||
<specimenCount id="E0D0A4C4FF92FFC2FB9AFD32FB5267F5" box="[1104,1235,727,752]" pageId="9" pageNumber="428" type="female">two female</specimenCount>
|
||
individuals, was collected on two expeditions to the Pulong Tau National Park in
|
||
<date id="8268498DFF92FFC2FCADFD16FB906611" box="[871,1041,764,789]" pageId="9" pageNumber="428" value="2012" valueMax="2013">2012 and 2013</date>
|
||
. Males and larval stages of the new species are, as yet, unknown. Both specimens were found at night, resting on the leaves of lower vegetation at the edge of the summit trail to Gunung Murud, ca. two hours walk from Church Camp, a few meters from the edge of a small stream. The
|
||
<typeStatus id="296DD1EFFF92FFC2FEB2FC82FE296685" box="[376,424,872,896]" pageId="9" pageNumber="428">type</typeStatus>
|
||
locality is situated between this unnamed small side stream and the Sugei Murud that runs eastward from near the summit of Gunung Murud down to the plateau of the Kelabit Highlands. The
|
||
<typeStatus id="296DD1EFFF92FFC2FADAFC67FAC166A0" box="[1296,1344,909,933]" pageId="9" pageNumber="428">type</typeStatus>
|
||
locality is situated just below the summit ridge of the Gunung Murud massif. In this undisturbed montane mossy forest (
|
||
<bibRefCitation id="924712BCFF92FFC2FF54FC3EFEC166E9" author="Beaman" box="[158,320,980,1004]" firstAuthor="Beaman" pageId="9" pageNumber="428" refString="Beaman, J. H. (1999) Preliminary enumeration of the summit flora, Mount Murud, Kelabit Highlands, Sarawak. ASEAN Review of Biodiversity and Environmental Conservation, pp 23. Available from: http: // www. arbec. com. my / pdf / art 3 mayjun 99. pdf (accessed 6 May 2014)" type="journal article" year="1999">Beaman 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF92FFC2FE87FC3FFD0166E9" author="Beaman & Anderson" box="[333,640,980,1005]" firstAuthor="Beaman" pageId="9" pageNumber="428" refString="Beaman, J. H. & Anderson, C. (1997) The summit flora of Mt. Murud, Sarawak, Malaysia. Contributions from the University of Michigan Herbarium, 21, 85 - 141." type="journal article" year="1997">Beaman & Anderson 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="924712BCFF92FFC2FD47FC3EFCB166E8" author="Mjoberg" box="[653,816,980,1005]" firstAuthor="Mjoberg" pageId="9" pageNumber="428" refString="Mjoberg, E. (1925) An expedition to the Kelabit country and Mt. Murud, Sarawak. Geographical Review, 15, 411 - 427. http: // dx. doi. org / 10.2307 / 208563" type="journal article" year="1925">Mjöberg 1925</bibRefCitation>
|
||
), the following species of frogs have been recorded in sympatry:
|
||
<taxonomicName id="31D614CEFF92FFC2FEDAFC12FD8F6115" box="[272,526,1015,1040]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="C4A2B35FFF92FFC2FEDAFC12FE1C610A" box="[272,413,1016,1039]" italics="true" pageId="9" pageNumber="428">Limnonectes</emphasis>
|
||
cf.
|
||
<emphasis id="C4A2B35FFF92FFC2FE04FC12FD8F6115" box="[462,526,1016,1040]" italics="true" pageId="9" pageNumber="428">kuhlii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="C4A2B35FFF92FFC2FDD4FC12FC9C6115" box="[542,797,1016,1040]" italics="true" pageId="9" pageNumber="428">
|
||
<taxonomicName id="31D614CEFF92FFC2FDD4FC12FD7B6115" box="[542,762,1016,1040]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="mjobergi">Philautus mjobergi</taxonomicName>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FCC2FC13FC9C6115" box="[776,797,1017,1040]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="petersi">P.</taxonomicName>
|
||
</emphasis>
|
||
cf.
|
||
<emphasis id="C4A2B35FFF92FFC2FC86FC12FC19610A" box="[844,920,1016,1039]" italics="true" pageId="9" pageNumber="428">petersi</emphasis>
|
||
,
|
||
<taxonomicName id="31D614CEFF92FFC2FC63FC12FB346115" box="[937,1205,1016,1040]" class="Amphibia" family="Bufonidae" genus="Pelophryne" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="murudensis">
|
||
<emphasis id="C4A2B35FFF92FFC2FC63FC12FB346115" box="[937,1205,1016,1040]" italics="true" pageId="9" pageNumber="428">Pelophryne murudensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="31D614CEFF92FFC2FB3BFC12FE8C6131" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="montanum">
|
||
<emphasis id="C4A2B35FFF92FFC2FB3BFC12FE8C6131" italics="true" pageId="9" pageNumber="428">Leptobrachium montanum</emphasis>
|
||
</taxonomicName>
|
||
. The amphibian fauna of Gunung Murud has been described by
|
||
<bibRefCitation id="924712BCFF92FFC2FC3BFBF7FBE96130" author="Das" box="[1009,1128,1052,1077]" firstAuthor="Das" pageId="9" pageNumber="428" refString="Das, I. (2005) A new species of Polypedates (Anura: Rhacophoridae) from Gunung Murud, Sarawak (northwestern Borneo). Raffles Bulletin of Zoology, 53, 265 - 270." type="journal article" year="2005">Das (2005</bibRefCitation>
|
||
, 2008). The discovery of a new and possibly endemic species from the Gunung Murud massif underlines the importance of the Pulong Tau National Park in the protection of the highly diverse montane amphibian fauna of
|
||
<collectingCountry id="8EC12FDDFF92FFC2FBE7FB8EFB0C6178" box="[1069,1165,1124,1149]" name="Malaysia" pageId="9" pageNumber="428">Sarawak</collectingCountry>
|
||
and that of Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |