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<mods:title id="9A687A81B38851EF72D766B0CBFD7ADB">Revisiting Pagurus pilosipes (Stimpson, 1858) (Crustacea: Decapoda: Anomura: Paguridae)</mods:title>
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<mods:namePart id="81614E177B44EC8226924E3EBA261E5A">Yang, Chien-Hui</mods:namePart>
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<mods:namePart id="C85A7EE56399A55036617C52E3EF1E2B">Chan, Tin-Yam</mods:namePart>
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Genus
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<emphasis id="C4ACEA978579FFCBFF2FFF21FE44FF6B" bold="true" box="[238,412,221,247]" italics="true" pageId="2" pageNumber="43">Boninpagurus</emphasis>
<bibRefCitation id="92494B748579FFCBFE65FF21FCDFFF6B" author="Asakura" box="[420,775,221,247]" pageId="2" pageNumber="43" refString="Asakura, A. &amp; Tachikawa, H. (2004) Boninpagurus acanthocheles, a new genus and new species of hermit crab (Decapoda: Anomura: Paguridae) from shallow waters of the Ogasawara (Bonin) Islands, Japan. Journal of Crustacean Biology, 24, 157 - 167." type="journal article" year="2004">Asakura &amp; Tachikawa, 2004</bibRefCitation>
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<paragraph id="F66736858579FFCBFF56FEFCFC8AFE8B" blockId="2.[151,850,221,279]" box="[151,850,255,279]" pageId="2" pageNumber="43">[New Japanese name: Okinawa-akashima-hon-yadokari-zoku]</paragraph>
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<taxonomicName id="31D84D068579FFCBFF56FEB5FD5CFEC2" ID-CoL="72040a76-b803-4317-93e5-c4ba84281ff8" authority="Asakura &amp; Tachikawa, 2004: 157" authorityName="Asakura &amp; Tachikawa" authorityPageNumber="157" authorityYear="2004" box="[151,644,328,350]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="C4ACEA978579FFCBFF56FEB5FEFEFEC2" box="[151,294,329,350]" italics="true" pageId="2" pageNumber="43">Boninpagurus</emphasis>
<treatmentCitation id="777910948579FFCBFEEDFEB4FD5CFEC2" author="Asakura" box="[300,644,328,350]" page="157" pageId="2" pageNumber="43" year="2004">
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: 157
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.
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</paragraph>
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species.
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<taxonomicName id="31D84D068579FFCBFE81FE70FC05FE3E" authority="Asakura &amp; Tachikawa, 2004" authorityName="Asakura &amp; Tachikawa" authorityYear="2004" box="[320,989,394,419]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="acanthocheles">
<emphasis id="C4ACEA978579FFCBFE81FE70FD51FE3F" box="[320,649,395,419]" italics="true" pageId="2" pageNumber="43">Boninpagurus acanthocheles</emphasis>
<bibRefCitation id="92494B748579FFCBFD52FE76FC05FE3E" author="Asakura" box="[659,989,394,418]" pageId="2" pageNumber="43" refString="Asakura, A. &amp; Tachikawa, H. (2004) Boninpagurus acanthocheles, a new genus and new species of hermit crab (Decapoda: Anomura: Paguridae) from shallow waters of the Ogasawara (Bonin) Islands, Japan. Journal of Crustacean Biology, 24, 157 - 167." type="journal article" year="2004">Asakura &amp; Tachikawa, 2004</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="31D84D068579FFCBFBCAFE70FF02FE5A" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="pilosipes" status="comb. nov.">
<emphasis id="C4ACEA978579FFCBFBCAFE70FAC0FE3F" box="[1035,1304,395,419]" italics="true" pageId="2" pageNumber="43">Boninpagurus pilosipes</emphasis>
(Stimpson, 1858)
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,
<taxonomicNameLabel id="DF9F57EC8579FFCBFF27FE53FE9EFE5A" box="[230,326,430,454]" pageId="2" pageNumber="43">n. comb</taxonomicNameLabel>
]. Original designation.
</paragraph>
<paragraph id="F66736858579FFCBFF04FE2FFBFFFC06" blockId="2.[151,1436,394,1319]" pageId="2" pageNumber="43">
<emphasis id="C4ACEA978579FFCBFF04FE2FFE6BFE77" bold="true" box="[197,435,466,491]" pageId="2" pageNumber="43">Emended diagnosis.</emphasis>
Eleven pairs of biserial phyllobranchiate gills, including 2 pairs of arthrobranchiae above bases of third maxilliped to fourth pereopod and 1 pleurobranch on seventh thoracic somite (above base of fourth pereopod). Rostrum (
<figureCitation id="6EE32A008579FFCBFE4AFDE6FE08FDAE" box="[395,464,538,562]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
A) broadly triangular, not particularly developed. Ocular acicles (
<figureCitation id="6EE32A008579FFCBFB00FDE6FADEFDAE" box="[1217,1286,538,562]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
A) each with submarginal spine. Antennal peduncle (
<figureCitation id="6EE32A008579FFCBFD93FDC2FD4DFDCA" box="[594,661,574,598]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
A) with supernumerary segmentation; acicles well developed. Maxillule with produced external lobe on endopod (
<figureCitation id="6EE32A008579FFCBFD67FD9EFD34FDE6" box="[678,748,610,634]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
B). Third maxilliped with well developed crista dentata consisting of fine row of corneous teeth and accessory tooth on ischium (
<figureCitation id="6EE32A008579FFCBFC71FD7AFC2DFD02" box="[944,1013,646,670]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
D). Chelipeds distinctly unequal with right longer and stouter than left (
<figureCitation id="6EE32A008579FFCBFDCEFD56FD88FD5E" box="[527,592,682,706]" captionStart="FIGURE 1" captionStartId="5.[151,250,1825,1848]" captionTargetBox="[151,1436,195,1776]" captionTargetId="figure@5.[151,1436,195,1776]" captionTargetPageId="5" captionText="FIGURE 1. Boninpagurus pilopsipes (Stimpson, 1858), n. comb. A, D, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands; B, male (SL 1.7 mm; CBM-ZC 10207), Maehama, Chichi-jima Island, Ogasawara Islands; C, E, male with rhizocephalan parasite attached to pleon (SL 3.0 mm; CBM-ZC 10000), Hayama, Sagami Bay. A, B, C, entire animal, dorsal view; D, E, close up of cephalic appendages, showing differences in color pattern of antennular peduncle." httpUri="https://zenodo.org/record/202570/files/figure.png" pageId="2" pageNumber="43">Figs 1</figureCitation>
AC, 3, 4A, C). Second and third pereopods generally similar (
<figureCitation id="6EE32A008579FFCBFAD8FD56FA83FD5E" box="[1305,1371,682,706]" captionStart="FIGURE 1" captionStartId="5.[151,250,1825,1848]" captionTargetBox="[151,1436,195,1776]" captionTargetId="figure@5.[151,1436,195,1776]" captionTargetPageId="5" captionText="FIGURE 1. Boninpagurus pilopsipes (Stimpson, 1858), n. comb. A, D, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands; B, male (SL 1.7 mm; CBM-ZC 10207), Maehama, Chichi-jima Island, Ogasawara Islands; C, E, male with rhizocephalan parasite attached to pleon (SL 3.0 mm; CBM-ZC 10000), Hayama, Sagami Bay. A, B, C, entire animal, dorsal view; D, E, close up of cephalic appendages, showing differences in color pattern of antennular peduncle." httpUri="https://zenodo.org/record/202570/files/figure.png" pageId="2" pageNumber="43">Figs 1</figureCitation>
AC, 4D, E). Fourth pereopods semichelate, lacking preungual process on dactylus; propodal rasp of more than 2 rows of corneous scales (
<figureCitation id="6EE32A008579FFCBFE9AFD0EFE7AFC96" box="[347,418,754,778]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
E). Fifth pereopods chelate. Coxa of right fifth pereopod in male bearing sexual tube of medium length, directed laterally or posterolaterally (
<figureCitation id="6EE32A008579FFCBFD32FCEAFCEDFCB2" box="[755,821,790,814]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
G, H, L); coxa of left fifth pereopod with short protrusion of vas deferens (
<figureCitation id="6EE32A008579FFCBFE44FCC6FE11FCCE" box="[389,457,826,850]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
J). Female with paired gonopores. Male pleon with 3 unequally biramous left pleopods; female pleon with 4 unequally biramous left pleopods. Telson (
<figureCitation id="6EE32A008579FFCBFCA8FCA2FC75FCEA" box="[873,941,862,886]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
K) with lateral indentations; posterior lobes separated by median cleft, left larger; terminal margins oblique, each with spines.
</paragraph>
</subSubSection>
<subSubSection id="BEC2650E8579FFCBFF04FC5BFC60FABB" pageId="2" pageNumber="43" type="discussion">
<paragraph id="F66736858579FFCBFF04FC5BFDF2FBEE" blockId="2.[151,1436,394,1319]" pageId="2" pageNumber="43">
<emphasis id="C4ACEA978579FFCBFF04FC5BFEE1FC23" bold="true" box="[197,313,935,959]" pageId="2" pageNumber="43">Remarks.</emphasis>
The original generic diagnosis of
<taxonomicName id="31D84D068579FFCBFD01FC5BFC87FC22" box="[704,863,935,958]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="C4ACEA978579FFCBFD01FC5BFC87FC22" box="[704,863,935,958]" italics="true" pageId="2" pageNumber="43">Boninpagurus</emphasis>
</taxonomicName>
given by
<bibRefCitation id="92494B748579FFCBFC13FC5AFAFAFC22" author="Asakura" box="[978,1314,934,958]" pageId="2" pageNumber="43" refString="Asakura, A. &amp; Tachikawa, H. (2004) Boninpagurus acanthocheles, a new genus and new species of hermit crab (Decapoda: Anomura: Paguridae) from shallow waters of the Ogasawara (Bonin) Islands, Japan. Journal of Crustacean Biology, 24, 157 - 167." type="journal article" year="2004">Asakura &amp; Tachikawa (2004)</bibRefCitation>
contains a few minor discrepancies or inconsistencies, and therefore is slightly emended herewith. The rostrum was described as “well developed”, but examination of the present series of specimens has shown that the rostrum is usually broadly triangular, not showing particular development. The original authors stated that the propodal-carpal articulation of the left cheliped was rotated counter-clockwise 30° from the horizontal plane, but we found no particular degree of rotation in our specimens.
</paragraph>
<paragraph id="F66736858579FFCBFF04FB83FC60FABB" blockId="2.[151,1436,394,1319]" pageId="2" pageNumber="43">
As noted above, males of the newly collected specimens from Okinawa Islands have the right sexual tube of a medium length (about 5.0 times of the coxal length; see
<figureCitation id="6EE32A008579FFCBFCE4FB5EFCB2FB26" box="[805,874,1186,1210]" captionStart="FIGURE 2" captionStartId="6.[151,250,1824,1847]" captionTargetBox="[248,1353,208,1793]" captionTargetId="figure@6.[233,1376,193,1806]" captionTargetPageId="6" captionText="FIGURE 2. Boninpagurus pilosipes (Stimpson, 1858), n. comb. A I, male (SL 3.0 mm; CBM-ZC 10000), Sagami Bay; J, female (SL 2.7 mm; CMNH-ZC 629), Boso Peninsula; L, male (SL 2.8 mm; CBM-ZC 10206), Kouri Island, Okinawa Islands. A, shield and cephalic appendages, dorsal view; B, endopod of left maxillule, lateral view; C, left third maxilliped, lateral view; D, same, ischium and basis, ventral view (setae omitted); E, dactylus and propodus of left fourth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, L, coxae of fifth pereopods and eighth thoracic sternite, ventral view; H, right sexual tube, lateral view; I, left coxa of fifth pereopod, lateral view; J, coxae of third pereopods and sixth thoracic sternite. Scale bars: Scale bars: 1 mm for A, C, G J, L; 0.5 mm for B, D F, K." httpUri="https://zenodo.org/record/202571/files/figure.png" pageId="2" pageNumber="43">Fig. 2</figureCitation>
C). In every diagnostic aspect,
<taxonomicName id="31D84D068579FFCBFB11FB58FA44FB27" box="[1232,1436,1187,1211]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA978579FFCBFB11FB58FA44FB27" box="[1232,1436,1187,1211]" italics="true" pageId="2" pageNumber="43">Pagurus pilosipes</emphasis>
</taxonomicName>
agrees with the diagnosis of
<taxonomicName id="31D84D068579FFCBFE1CFB34FDA4FB43" box="[477,636,1224,1247]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="C4ACEA978579FFCBFE1CFB34FDA4FB43" box="[477,636,1224,1247]" italics="true" pageId="2" pageNumber="43">Boninpagurus</emphasis>
</taxonomicName>
given by
<bibRefCitation id="92494B748579FFCBFD31FB3AFB9AFB42" author="Asakura" box="[752,1090,1222,1246]" pageId="2" pageNumber="43" refString="Asakura, A. &amp; Tachikawa, H. (2004) Boninpagurus acanthocheles, a new genus and new species of hermit crab (Decapoda: Anomura: Paguridae) from shallow waters of the Ogasawara (Bonin) Islands, Japan. Journal of Crustacean Biology, 24, 157 - 167." type="journal article" year="2004">Asakura &amp; Tachikawa (2004)</bibRefCitation>
, and thus the species is herein transferred to
<taxonomicName id="31D84D068579FFCBFEFFFB10FE05FA9F" box="[318,477,1260,1283]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="C4ACEA978579FFCBFEFFFB10FE05FA9F" box="[318,477,1260,1283]" italics="true" pageId="2" pageNumber="43">Boninpagurus</emphasis>
</taxonomicName>
. As dicussed below, conspecificity between
<taxonomicName id="31D84D068579FFCBFC31FB10FBA0FA9F" box="[1008,1144,1259,1283]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="C4ACEA978579FFCBFC31FB10FBA0FA9F" box="[1008,1144,1259,1283]" italics="true" pageId="2" pageNumber="43">B. pilosipes</emphasis>
</taxonomicName>
and
<taxonomicName id="31D84D068579FFCBFB78FB10FAA6FA9F" box="[1209,1406,1259,1283]" class="Malacostraca" family="Paguridae" genus="Boninpagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="acanthocheles">
<emphasis id="C4ACEA978579FFCBFB78FB10FAA6FA9F" box="[1209,1406,1259,1283]" italics="true" pageId="2" pageNumber="43">B. acanthocheles</emphasis>
</taxonomicName>
is strongly supported by comparisons of morphology and barcoding gene.
</paragraph>
</subSubSection>
</treatment>
</document>