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<document id="60754B6D867A6ED93E5FE3B45D83EFD1" ID-CLB-Dataset="7571" ID-DOI="10.11646/zootaxa.4949.3.3" ID-GBIF-Dataset="39300142-6cea-4253-80bb-00856fdd5f95" ID-ISSN="1175-5326" ID-Zenodo-Dep="4640469" ID-ZooBank="C22ACA1A-4856-4979-BD7A-DFE37949C2D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616780319867" checkinUser="plazi" docAuthor="Fei, Xu-Dong &amp; Zhou, Hong-Zhang" docDate="2021" docId="7E5287DBE3542F7BFF2EFF48FBEF0777" docLanguage="en" docName="zootaxa.4949.3.3.pdf" docOrigin="Zootaxa 4949 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eccoptolonthus fuyuensis Fei &amp; Zhou 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="487" masterDocId="826BFFA3E3582F75FFB9FFDFFFE7017E" masterDocTitle="Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China" masterLastPageNumber="498" masterPageNumber="473" pageNumber="485" updateTime="1698935240280" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3606D2B768E15F82F042D90CB0C19083" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="41CF6A8E2B25094BC6E79FCFD307E2B2">
<mods:title id="F0DE5EEF02849984EE62F3E538A9B4B7">Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China</mods:title>
</mods:titleInfo>
<mods:name id="391CC434444AA399BF33D7CF60F9255E" type="personal">
<mods:role id="FBE90E3D02ADE6067B6B9EFD05CB87DD">
<mods:roleTerm id="4A60F3F07CCED54074BBA77CF81A5FE0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B080FB959122F1E18B3D259034BC3E1D">Fei, Xu-Dong</mods:namePart>
<mods:nameIdentifier id="4BCD5AD132E487AE483D5626616D221F" type="ORCID">0000-0003-0386-5918</mods:nameIdentifier>
<mods:affiliation id="B92CAB369C69D34353516745EF962A37">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China. &amp; feixudong @ ioz. ac. cn; https: // orcid. org / 0000 - 0003 - 0386 - 5918</mods:affiliation>
<mods:nameIdentifier id="D50F52C29979B479A38067CD97A4C6CF" type="email">feixudong@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="E02965B6CDF503E299529A39BCCE5A8B" type="personal">
<mods:role id="DA4C35914E7759D52203983C8E9C2CE4">
<mods:roleTerm id="CC20B73B39B895FB1F2DCF5E6A99D938">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47D7688F42DABE5391AE00FB15EFF083">Zhou, Hong-Zhang</mods:namePart>
<mods:affiliation id="A01563D245FCDD1071979A45E3EA5828">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<mods:title id="D7E2ABEB9680455394FFFB2F7D96EA39">Zootaxa</mods:title>
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<mods:part id="D5D21F2E350B6389A72C3CC82A463EC0">
<mods:date id="DFC8FA13028ACB413E386EFC0A7D114F">2021</mods:date>
<mods:detail id="D88C1E143355FB0C4EF3BFA4E41F7C89" type="pubDate">
<mods:number id="5E16B54408B35B8282A50B70990ABCDA">2021-03-26</mods:number>
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<mods:detail id="3CD08F1F64B6AD48BB37E1C323D19051" type="volume">
<mods:number id="D0AFB41CA2955CE0D264A2AC9BE035C5">4949</mods:number>
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<mods:number id="ACAE93250244E2F4EAF1175F216CC9AA">3</mods:number>
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<mods:classification id="A2A6589ACAFEC210BFC80714230AED7C">journal article</mods:classification>
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<mods:identifier id="1D12FF5520AACE1B1C4B2BC5B00A3AFB" type="DOI">10.11646/zootaxa.4949.3.3</mods:identifier>
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<treatment id="7E5287DBE3542F7BFF2EFF48FBEF0777" ID-DOI="http://doi.org/10.5281/zenodo.4640487" ID-GBIF-Taxon="180780157" ID-Zenodo-Dep="4640487" LSID="urn:lsid:plazi:treatment:7E5287DBE3542F7BFF2EFF48FBEF0777" httpUri="http://treatment.plazi.org/id/7E5287DBE3542F7BFF2EFF48FBEF0777" lastPageId="14" lastPageNumber="487" pageId="12" pageNumber="485">
<subSubSection id="BEE16546E3542F79FF2EFF48FD1601CF" box="[151,753,151,178]" pageId="12" pageNumber="485" type="nomenclature">
<paragraph id="F64436CDE3542F79FF2EFF48FD1601CF" blockId="12.[151,753,151,213]" box="[151,753,151,178]" pageId="12" pageNumber="485">
<heading id="AD0C81A1E3542F79FF2EFF48FD1601CF" bold="true" box="[151,753,151,178]" fontSize="11" level="1" pageId="12" pageNumber="485" reason="1">
<emphasis id="C48FEADFE3542F79FF2EFF48FD1601CF" bold="true" box="[151,753,151,178]" pageId="12" pageNumber="485">
5.
<taxonomicName id="31FB4D4EE3542F79FF0CFF48FD6101CF" ID-CoL="8XZXB" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[181,646,151,178]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="485" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE3542F79FF0CFF48FE0A01CF" bold="true" box="[181,493,151,178]" italics="true" pageId="12" pageNumber="485">Eccoptolonthus fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="DFBC57A4E3542F79FD2DFF48FD1601CF" box="[660,753,151,177]" pageId="12" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3542F79FF2EFF64FE9D01AB" box="[151,378,187,213]" pageId="12" pageNumber="485" type="description">
<paragraph id="F64436CDE3542F79FF2EFF64FE9D01AB" blockId="12.[151,753,151,213]" box="[151,378,187,213]" pageId="12" pageNumber="485">
(
<figureCitation id="6EC02A48E3542F79FF26FF64FEC101AB" box="[159,294,187,213]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="12" pageNumber="485">Figs. 7A–G</figureCitation>
,
<figureCitation id="6EC02A48E3542F79FE88FF64FE9501AB" box="[305,370,187,213]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="12" pageNumber="485">8A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3542F79FF2EFEDCFD9B003F" pageId="12" pageNumber="485" type="materials_examined">
<paragraph id="F64436CDE3542F79FF2EFEDCFD9B003F" blockId="12.[151,1437,259,682]" pageId="12" pageNumber="485">
<emphasis id="C48FEADFE3542F79FF2EFEDCFEB30063" bold="true" box="[151,340,259,285]" pageId="12" pageNumber="485">Type materials.</emphasis>
<materialsCitation id="46933C90E3542F79FEE7FEDCFD9B003F" ID-GBIF-Occurrence="3063281351" collectingDate="2009-06-28" collectionCode="IZ-CAS" collectorName="Wu Jie" country="China" elevation="400" location="Tianmu mountain" municipality="Linan" pageId="12" pageNumber="485" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="holotype">
<emphasis id="C48FEADFE3542F79FEE7FEDCFE350063" bold="true" box="[350,466,259,285]" pageId="12" pageNumber="485">
<typeStatus id="2940886FE3542F79FEE7FEDCFE2A0063" box="[350,461,259,285]" pageId="12" pageNumber="485" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="E0FDFD44E3542F79FE65FED8FE160063" box="[476,497,263,285]" pageId="12" pageNumber="485" type="male">♂</specimenCount>
,
<emphasis id="C48FEADFE3542F79FDBAFEDCFD320063" bold="true" box="[515,725,259,285]" pageId="12" pageNumber="485">
<collectingCountry id="8EEC765DE3542F79FDBAFEDCFDB70063" box="[515,592,259,285]" name="China" pageId="12" pageNumber="485">China</collectingCountry>
:
<collectingRegion id="343FF82FE3542F79FDD9FEDCFD370063" box="[608,720,259,285]" country="China" name="Zhejiang" pageId="12" pageNumber="485">Zhejiang</collectingRegion>
:
</emphasis>
<collectingMunicipality id="1620ACB7E3542F79FD66FEDCFCC20063" box="[735,805,259,285]" pageId="12" pageNumber="485">Linan</collectingMunicipality>
:
<location id="F3246016E3542F79FC8AFEDCFC1A0063" LSID="urn:lsid:plazi:treatment:7E5287DBE3542F7BFF2EFF48FBEF0777:F3246016E3542F79FC8AFEDCFC1A0063" box="[819,1021,259,285]" country="China" municipality="Linan" name="Tianmu mountain" pageId="12" pageNumber="485" stateProvince="Zhejiang">Tianmu mountain</location>
(30º2006′ N, 119º2654′ E),
<quantity id="31039B28E3542F79FAF2FEDCFA700060" box="[1355,1431,259,286]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="485" unit="m" value="400.0">
<elevation id="7DD6D1FEE3542F79FAF2FEDCFA700060" box="[1355,1431,259,286]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="485" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="8245100DE3542F79FF2EFEF8FEFD003F" box="[151,282,295,321]" pageId="12" pageNumber="485" value="2009-06-28">
<collectingDate id="9201E9E5E3542F79FF2EFEF8FEFD003F" box="[151,282,295,321]" pageId="12" pageNumber="485" value="2009-06-28">28.VI.2009</collectingDate>
</date>
, collected by
<collectorName id="5B0E531BE3542F79FE0AFEF7FDE6003F" box="[435,513,295,322]" pageId="12" pageNumber="485">Wu Jie</collectorName>
(
<collectionCode id="90EAAE08E3542F79FDA9FEF8FD96003F" box="[528,625,295,321]" pageId="12" pageNumber="485">IZ-CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3542F7BFF7EFE94FF2F0503" lastPageId="14" lastPageNumber="487" pageId="12" pageNumber="485" type="description">
<paragraph id="F64436CDE3542F79FF7EFE94FD8D00D0" blockId="12.[151,1437,259,682]" pageId="12" pageNumber="485">
<emphasis id="C48FEADFE3542F79FF7EFE94FEBE001B" bold="true" box="[199,345,331,357]" pageId="12" pageNumber="485">Description.</emphasis>
Length: BL=
<quantity id="31039B28E3542F79FE47FE94FD830018" box="[510,612,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="7.19" pageId="12" pageNumber="485" unit="mm" value="7.19">7.19 mm</quantity>
(HPL=
<quantity id="31039B28E3542F79FD7AFE94FCCA0018" box="[707,813,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="2.21" pageId="12" pageNumber="485" unit="mm" value="2.21">2.21 mm</quantity>
). Length of different body parts: HW=
<quantity id="31039B28E3542F79FB4BFE94FABD0018" box="[1266,1370,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="485" unit="mm" value="1.15">1.15 mm</quantity>
, HL=
<quantity id="31039B28E3542F79FF2EFEB0FF1B00F4" box="[151,252,367,394]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="485" unit="mm" value="0.95">0.95 mm</quantity>
(HW= HL= 1.25); ELL=
<quantity id="31039B28E3542F79FD9AFEB0FD6E00F4" box="[547,649,367,394]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="485" unit="mm" value="0.22">0.22 mm</quantity>
, TL=
<quantity id="31039B28E3542F79FD77FEB0FCD100F4" box="[718,822,367,394]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="485" unit="mm" value="0.17">0.17 mm</quantity>
; PL=
<quantity id="31039B28E3542F79FCC3FEB0FC0700F4" box="[890,992,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="12" pageNumber="485" unit="mm" value="1.26">1.26 mm</quantity>
, PW=
<quantity id="31039B28E3542F79FB97FEB0FB7500F4" box="[1070,1170,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="485" unit="mm" value="1.16">1.16 mm</quantity>
(PW: HW= 1.10); EL=
<quantity id="31039B28E3542F79FF2EFE4CFF1A00D0" box="[151,253,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="12" pageNumber="485" unit="mm" value="1.66">1.66 mm</quantity>
, EW=
<quantity id="31039B28E3542F79FEF2FE4CFE5500D0" box="[331,434,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="485" unit="mm" value="1.6">1.60 mm</quantity>
; AW=
<quantity id="31039B28E3542F79FE46FE4CFD8100D0" box="[511,614,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="485" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
<paragraph id="F64436CDE3542F79FF7EFE68FAB103D7" blockId="12.[151,1437,259,682]" pageId="12" pageNumber="485">Head black with slightly iridescent lustre. Pronotum dark reddish-brown. Antennae with antennomere I dark brown, apical-half of antennomere II–III dark brown, basal-half of antennomere II–III black, antennomere IV–V dark brown; due to damage, characters of apical segments unknown. Elytra black with suture and posterior margin markedly reddish-brown. Scutellum dark reddish-brown. Abdomen black with strongly iridescent lustre, posterior margin of each tergite narrowly reddish-brown. Mandibles and labrum reddish-brown, III, IV segments of maxillary and labial palpi yellowish-brown, base part of maxillary palpi reddish-brown. Profemur and mesofemur yellowishbrown, metafemur dark reddish-brown, tarsomere I dark brown, tarsi II–IV yellowish-brown, tarsi V yellow.</paragraph>
<caption id="A2846645E3542F79FF2EF84FFE070691" ID-DOI="http://doi.org/10.5281/zenodo.4640491" ID-Zenodo-Dep="4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="12" pageNumber="485" startId="12.[151,250,1936,1960]" targetBox="[205,1376,707,1907]" targetPageId="12">
<paragraph id="F64436CDE3542F79FF2EF84FFE070691" blockId="12.[151,1436,1935,2032]" pageId="12" pageNumber="485">
<emphasis id="C48FEADFE3542F79FF2EF84FFCE006D6" bold="true" box="[151,775,1935,1960]" pageId="12" pageNumber="485">
FIGURE 7.
<taxonomicName id="31FB4D4EE3542F79FEA0F850FD4C06D6" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[281,683,1935,1960]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="485" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE3542F79FEA0F850FDC006D9" bold="true" box="[281,551,1935,1959]" italics="true" pageId="12" pageNumber="485">Eccoptolonthus fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="DFBC57A4E3542F79FD0FF84FFCE006D6" box="[694,775,1936,1960]" pageId="12" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;
</paragraph>
</caption>
<caption id="A2846645E3552F78FF2EF8A3FDFF06A2" ID-DOI="http://doi.org/10.5281/zenodo.4640495" ID-Zenodo-Dep="4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="13" pageNumber="486" startId="13.[151,250,1916,1940]" targetBox="[270,1307,181,1878]" targetPageId="13">
<paragraph id="F64436CDE3552F78FF2EF8A3FDFF06A2" blockId="13.[151,1436,1916,2012]" pageId="13" pageNumber="486">
<emphasis id="C48FEADFE3552F78FF2EF8A3FCE806EA" bold="true" box="[151,783,1916,1940]" pageId="13" pageNumber="486">
FIGURE 8.
<taxonomicName id="31FB4D4EE3552F78FEA2F8A3FD5506EA" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[283,690,1916,1940]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="486" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE3552F78FEA2F8A3FDCD06EA" bold="true" box="[283,554,1916,1940]" italics="true" pageId="13" pageNumber="486">Eccoptolonthus fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
,
<taxonomicNameLabel id="DFBC57A4E3552F78FD07F8A3FCE806EA" box="[702,783,1916,1940]" pageId="13" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm.
</paragraph>
</caption>
<paragraph id="F64436CDE3562F7BFF7EFF48FADB00F7" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">Head of subquadrate shape, subparallel-sided, obtusely rounded at basal angles, slightly wider than long. Tempora almost evenly rounded, moderately and coarsely punctate; eyes large slightly convex, about 1.29 times as long as tempora. Dorsal surface of head with moderately numerous, large setiferous punctures, becoming sparser toward vertex; vertex largely impunctate. Entire head with distinct and dense microsculpture of transverse waves. Antennae long, antennomere I about two times as long as antennomere III, distinctly thickened towards apex, antennomere III about 1.25 times as long as antennomere II, following antennomeres gradually decreasing in length, but antennomere IV–V distinctly longer than wide; due to damage, characteristics of other antennomeres unknown.</paragraph>
<paragraph id="F64436CDE3562F7BFF7EFE4CFF330367" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">Pronotum approximate rectangular, slightly convex, posterior angles markedly rounded, pronotum almost as wide as head, parallel-sided; punctation moderate and fine, punctures separated by three to five times their diameter, narrowly impunctate along midline, the punctures approximately equal in size, microsculpture similar to that on head.</paragraph>
<paragraph id="F64436CDE3562F7BFF7EFDFCFD7203FB" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">Elytra about 1.3 times as along as pronotum, about 1.37 times as wide as pronotum; scutellum large, triangular; elytra and scutellum densely and finely punctate, punctures separated by one to two times their diameter, punctures much finer than those on head and pronotum.</paragraph>
<paragraph id="F64436CDE3562F7BFF7EFD50FCCA026B" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">Abdomen wide, narrowing towards apex, densely and finely punctate, punctures separated by two to three times their diameter; tergites III–V with two basal lines, elevated area between basal lines more sparsely punctate; punctation of basal portion of all tergites coarser than that on elytra, becoming sparser towards posterior margin of each tergite. Surface between punctures without microsculpture.</paragraph>
<paragraph id="F64436CDE3562F7BFF7EFCC0FDFE02B7" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">
Male: Sternite VIII moderately wide, approximate trapezoid, with obtusely triangular medioapical emargination (
<figureCitation id="6EC02A48E3562F7BFF6BFC9CFED70223" box="[210,304,835,861]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="14" pageNumber="487">Figs. 7F</figureCitation>
,
<figureCitation id="6EC02A48E3562F7BFE82FC9CFEBA0223" box="[315,349,835,861]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="14" pageNumber="487">8E</figureCitation>
). Genital segment with stylus of tergite IX simple, densely setae apically, and bearing two longer and stronger setae in the middle of tergite IX (
<figureCitation id="6EC02A48E3562F7BFD0AFCB8FCFE02FF" box="[691,793,871,897]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="14" pageNumber="487">Figs. 7G</figureCitation>
,
<figureCitation id="6EC02A48E3562F7BFC91FCB8FCAE02FF" box="[808,841,871,897]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="14" pageNumber="487">8F</figureCitation>
). Sternite IX with asymmetrical basal portion, apex deeply emarginate, with rod-like extensions (
<figureCitation id="6EC02A48E3562F7BFD2CFC54FD1E02DB" box="[661,761,907,933]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="14" pageNumber="487">Figs. 7D</figureCitation>
,
<figureCitation id="6EC02A48E3562F7BFCBFFC54FCCD02DB" box="[774,810,907,933]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="14" pageNumber="487">8C</figureCitation>
). Tergite X approximately trapezoid with slightly triangular emargination (
<figureCitation id="6EC02A48E3562F7BFEC3FC70FE3C02B7" box="[378,475,943,969]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="14" pageNumber="487">Figs. 7E</figureCitation>
,
<figureCitation id="6EC02A48E3562F7BFE5EFC70FDEB02B7" box="[487,524,943,969]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="14" pageNumber="487">8D</figureCitation>
).
</paragraph>
<paragraph id="F64436CDE3562F7BFF7EFC0CFF2F0503" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">
Aedeagus in ventral view with median lobe almost as wide as paramere, slightly expanded in the middle and narrowed into truncate apex, middle part of paramere narrowed, apex of paramere approximately circular, with dense peg setae irregularly arranged along apical portion of paramere (
<figureCitation id="6EC02A48E3562F7BFC17FBC4FBF7054B" box="[942,1040,1051,1077]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="14" pageNumber="487">Figs. 7B</figureCitation>
,
<figureCitation id="6EC02A48E3562F7BFBA2FBC4FBA6054B" box="[1051,1089,1051,1077]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="14" pageNumber="487">8A</figureCitation>
); in lateral view, apical portion moderately bent ventrally, paramere elongate, apical-half of paramere ellipsoidal, some fine setae on apex. (
<figureCitation id="6EC02A48E3562F7BFA85FBE0FA7F0527" box="[1340,1432,1087,1113]" captionStart="FIGURE 7" captionStartId="12.[151,250,1936,1960]" captionTargetBox="[205,1376,707,1907]" captionTargetId="figure-204@12.[199,1388,705,1912]" captionTargetPageId="12" captionText="FIGURE 7. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. sternite IX of male; E. tergite X of male; F. sternite VIII of male; g. stylus of tergite IX of male. Scale bars, A, B, C, D, E, F, G, H= 0.2 mm;" figureDoi="http://doi.org/10.5281/zenodo.4640491" httpUri="https://zenodo.org/record/4640491/files/figure.png" pageId="14" pageNumber="487">Figs.7C</figureCitation>
,
<figureCitation id="6EC02A48E3562F7BFF2EFBBCFF5C0503" box="[151,187,1123,1149]" captionStart="FIGURE 8" captionStartId="13.[151,250,1916,1940]" captionTargetBox="[270,1307,181,1878]" captionTargetId="figure-19@13.[265,1323,181,1892]" captionTargetPageId="13" captionText="FIGURE 8. Eccoptolonthus fuyuensis Fei &amp; Zhou, sp. nov. A. aedeagus, dorsal (parameral side) view; B. aedeagus, lateral view; C. sternite IX of male; D. tergite X of male; E. sternite VIII of male; F. stylus of tergite IX of male. Scale bars, A, B= 0.1 mm; C, D= 0.13 mm; E, F= 0.18 mm." figureDoi="http://doi.org/10.5281/zenodo.4640495" httpUri="https://zenodo.org/record/4640495/files/figure.png" pageId="14" pageNumber="487">8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3562F7BFF7EFB58FC3504E3" pageId="14" pageNumber="487" type="discussion">
<paragraph id="F64436CDE3562F7BFF7EFB58FC3504E3" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">
<emphasis id="C48FEADFE3562F7BFF7EFB58FEDC05DF" bold="true" box="[199,315,1159,1185]" pageId="14" pageNumber="487">Remarks.</emphasis>
<taxonomicName id="31FB4D4EE3562F7BFEF8FB58FD0305DF" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[321,740,1159,1185]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE3562F7BFEF8FB58FDBA05DF" box="[321,605,1159,1185]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
<emphasis id="C48FEADFE3562F7BFD53FB58FCA505DF" bold="true" box="[746,834,1159,1185]" pageId="14" pageNumber="487">
<taxonomicNameLabel id="DFBC57A4E3562F7BFD53FB58FCA505DF" box="[746,834,1159,1185]" pageId="14" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="31FB4D4EE3562F7BFBB5FB58FF3E05BB" authority="(Sharp, 1874)" authorityName="Schillhammer" authorityYear="2015" baseAuthorityName="Sharp" baseAuthorityYear="1874" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="rutiliventris">
<emphasis id="C48FEADFE3562F7BFBB5FB58FAA305DF" box="[1036,1348,1159,1185]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus rutiliventris</emphasis>
(
<bibRefCitation id="926A4B3CE3562F7BFAEBFB58FF3705BB" author="Sharp, D." pageId="14" pageNumber="487" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
but can be easily distinguished by the sensory peg setae of paramere uniformly distributed at the apex of paramere; in lateral view, the length of sensory peg setae about 0.25 times as long as width of the paramere (obviously shorter in
<taxonomicName id="31FB4D4EE3562F7BFEBEFB2CFD030473" authority="(Sharp, 1874))" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[263,740,1267,1293]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="rutilivenris">
<emphasis id="C48FEADFE3562F7BFEBEFB2CFDDF0473" box="[263,568,1267,1293]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus rutilivenris</emphasis>
(
<bibRefCitation id="926A4B3CE3562F7BFDFFFB2CFD360473" author="Sharp, D." box="[582,721,1267,1293]" pageId="14" pageNumber="487" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
))
</taxonomicName>
; in ventral view, paramere distinctly constricted at middle, this point about 0.36 times as broad as median lobe of aedeagus.
<taxonomicName id="31FB4D4EE3562F7BFCFAFAC8FB0C044F" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[835,1259,1303,1329]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE3562F7BFCFAFAC8FB87044F" box="[835,1120,1303,1329]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
<emphasis id="C48FEADFE3562F7BFB4AFAC8FAAB044F" bold="true" box="[1267,1356,1303,1329]" pageId="14" pageNumber="487">
<taxonomicNameLabel id="DFBC57A4E3562F7BFB4AFAC8FAAB044F" box="[1267,1356,1303,1329]" pageId="14" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from
<taxonomicName id="31FB4D4EE3562F7BFE00FAE4FC96042B" authority="(Sharp, 1874)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[441,881,1339,1365]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="gastralis">
<emphasis id="C48FEADFE3562F7BFE00FAE4FD37042B" box="[441,720,1339,1365]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus gastralis</emphasis>
(
<bibRefCitation id="926A4B3CE3562F7BFD66FAE4FC8E042B" author="Sharp, D." box="[735,873,1339,1365]" pageId="14" pageNumber="487" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
by the following characters: male sternite VIII of
<taxonomicName id="31FB4D4EE3562F7BFF2EFA80FDA30407" authority="Fei &amp; Zhou" authorityName="Fei &amp; Zhou" authorityYear="2021" box="[151,580,1375,1401]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="fuyuensis" status="sp. nov.">
<emphasis id="C48FEADFE3562F7BFF2EFA80FE510407" box="[151,438,1375,1401]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus fuyuensis</emphasis>
Fei &amp; Zhou
</taxonomicName>
<emphasis id="C48FEADFE3562F7BFDF4FA80FD4F0407" bold="true" box="[589,680,1375,1401]" pageId="14" pageNumber="487">
<taxonomicNameLabel id="DFBC57A4E3562F7BFDF4FA80FD4F0407" box="[589,680,1375,1401]" pageId="14" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with a pair of long appendages, (
<taxonomicName id="31FB4D4EE3562F7BFB90FA80FF3E04E3" authority="(Sharp, 1874)" baseAuthorityName="Sharp" baseAuthorityYear="1874" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="gastralis">
<emphasis id="C48FEADFE3562F7BFB90FA80FAA50407" box="[1065,1346,1375,1401]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus gastralis</emphasis>
(
<bibRefCitation id="926A4B3CE3562F7BFAEBFA80FF3704E3" author="Sharp, D." pageId="14" pageNumber="487" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
)
</taxonomicName>
without); in ventral view, paramere distinctly constricted at middle.
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3562F7BFF7EFA78FD990498" pageId="14" pageNumber="487" type="etymology">
<paragraph id="F64436CDE3562F7BFF7EFA78FD990498" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="487">
<emphasis id="C48FEADFE3562F7BFF7EFA78FEA904BF" bold="true" box="[199,334,1447,1473]" pageId="14" pageNumber="487">Etymology.</emphasis>
The specific epithet refers to the name of the
<typeStatus id="2940886FE3562F7BFCF4FA78FC9A04BF" box="[845,893,1447,1473]" pageId="14" pageNumber="487">type</typeStatus>
locality, Fuyu Mountain (Tianmu Mountain was named as Fuyu Mountain in ancient times).
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3562F7BFF7EFA31FBEF0777" box="[199,1032,1518,1545]" pageId="14" pageNumber="487" type="distribution">
<paragraph id="F64436CDE3562F7BFF7EFA31FBEF0777" blockId="14.[151,1437,151,1545]" box="[199,1032,1518,1545]" pageId="14" pageNumber="487">
<emphasis id="C48FEADFE3562F7BFF7EFA31FE860776" bold="true" box="[199,353,1518,1544]" pageId="14" pageNumber="487">Distribution.</emphasis>
This species is at present known only from the
<typeStatus id="2940886FE3562F7BFCCEFA30FC400777" box="[887,935,1519,1545]" pageId="14" pageNumber="487">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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