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<document id="1AF2C2DBDAA7693E216FBD2E898EFAF9" ID-CLB-Dataset="21199" ID-DOI="10.11646/zootaxa.4615.1.10" ID-GBIF-Dataset="806f3a8f-219d-48ff-9b05-8ff10546940a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995294" ID-ZooBank="70175096-AA78-4B74-BF38-44804F8FAD59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598056528139" checkinUser="felipe" docAuthor="Benetti, Cesar J., Short, Andrew E. Z. &amp; Michat, Mariano C." docDate="2019" docId="7E421778FF8FFFB3FF10F915A68D58FA" docLanguage="en" docName="zootaxa.4615.1.10.pdf" docOrigin="Zootaxa 4615 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hamadiana chapadensis Benetti &amp; Short &amp; Michat 2019, sp. n." docType="treatment" docUuid="88532963-8F7F-412B-A7BB-33CE812276BC" docUuidSource="ZooBank" docVersion="7" lastPageNumber="179" masterDocId="827B6F00FF8EFFB0FF87FFF0A2645971" masterDocTitle="Hamadiana chapadensis, a new genus and species of diving beetle from Brazil (Coleoptera, Dytiscidae, Laccophilinae, Laccophilini)" masterLastPageNumber="184" masterPageNumber="176" pageNumber="177" updateTime="1698845169827" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="581001FF095DFCF4E20533D16BEF89AE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9A350B36C02020FA06D8E34310A9A857">
<mods:title id="65DA2513A95DBBB48001D30811483D75">Hamadiana chapadensis, a new genus and species of diving beetle from Brazil (Coleoptera, Dytiscidae, Laccophilinae, Laccophilini)</mods:title>
</mods:titleInfo>
<mods:name id="A52E3C6122CF1088530C99BF3CBFCA07" type="personal">
<mods:role id="7F9A59B851B019A0F6173B8B137514F7">
<mods:roleTerm id="3D94E40D27866ABCE8AF681215267A62">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3471C5EBE7361DC2749B87DF8FBA9677">Benetti, Cesar J.</mods:namePart>
<mods:affiliation id="DCE0C16C533709C7DBF163598BB087D5">Coordenação de Biodiversidade, Programa de Pós-Graduação em Entomologia (PPGEnt), Instituto Nacional de Pesquisas da Amazônia (INPA), Av. André Araújo 2936, CEP 69067 - 375, Manaus, AM, Brazil. E-mail: cjbenetti @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="062BFD272EED0DA7BD378FAAC7B6D198" type="personal">
<mods:role id="8F891A9826A5878E215D38387277F993">
<mods:roleTerm id="2FF6F4C582A92AEA7AD64D9876DF5B5F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C7A7C20436646B6D8239D13BC3B4D35">Short, Andrew E. Z.</mods:namePart>
<mods:affiliation id="CBCC02F065A4360DAF88D62CB1089FC5">Department of Ecology &amp; Evolutionary Biology, Division of Entomology, Biology Institute, University of Kansas, Lawrence, United States. E-mail: aezshort @ ku. edu</mods:affiliation>
</mods:name>
<mods:name id="2DB3B3B5173A257EF4D10DB6D455B674" type="personal">
<mods:role id="6B98D95295D98F8F9D53E074B700F1C8">
<mods:roleTerm id="AF2917034AB4D35A5188F1511B42E9D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B07AC52773F1049CF02219E6E440300">Michat, Mariano C.</mods:namePart>
<mods:affiliation id="D8D067A4703C024D68968BECD4BC3B67">Laboratory of Entomology, IBBEA, CONICET-UBA., DBBE-FCEN, University of Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FF11F803F5D0F21225DFFC7568C4ABBB">text</mods:typeOfResource>
<mods:relatedItem id="98DFE543B05FED133E975C28146C1A76" type="host">
<mods:titleInfo id="883593AF3B61619C116245BFA9BD2C95">
<mods:title id="53475ED27076EB0040F9C67C762B3354">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6C6C0509D65C92A655DD1E8094A8529E">
<mods:date id="2B5B53F730A40AB01E012EC1AEDEC016">2019</mods:date>
<mods:detail id="9DC944072E3B6F5F070F2521B33B2700" type="pubDate">
<mods:number id="4CB177901E21EC72DB38E12F42DDD160">2019-06-12</mods:number>
</mods:detail>
<mods:detail id="B2D4605662E83C219BB37E7D3F7E1325" type="volume">
<mods:number id="6834246116DE557EAF586BB9EEF6982C">4615</mods:number>
</mods:detail>
<mods:detail id="F597B97FF3E7C85A3F0477CD9EF5A269" type="issue">
<mods:number id="DA6694E11B7D34E864A648CF9100F99C">1</mods:number>
</mods:detail>
<mods:extent id="8E5BBAA2CC5605CEC52FC27D3BE04A47" unit="page">
<mods:start id="2FE1EF306C9B135C48BFFC849D04F5D0">176</mods:start>
<mods:end id="1F204AE290C31F0FE7BF8AE3698AE212">184</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FD027631D06E269FEEDC05136BF5AAB8">journal article</mods:classification>
<mods:identifier id="64C3636A4BD6663FAEF7EAD840B1B32C" type="CLB-Dataset">21199</mods:identifier>
<mods:identifier id="3DCA70C2BDB981CD498C3FD1F104FBF5" type="DOI">10.11646/zootaxa.4615.1.10</mods:identifier>
<mods:identifier id="B2DC7C20D6CBB5132D26793BA0FE4AC8" type="GBIF-Dataset">806f3a8f-219d-48ff-9b05-8ff10546940a</mods:identifier>
<mods:identifier id="B2F5981E69121B62E0DBA37AA28228B5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5F4BBFF59AC5171EF0ECF751C4937625" type="Zenodo-Dep">3995294</mods:identifier>
<mods:identifier id="F1471E64F17FA20829F870E9A359D897" type="ZooBank">70175096-AA78-4B74-BF38-44804F8FAD59</mods:identifier>
</mods:mods>
<treatment id="7E421778FF8FFFB3FF10F915A68D58FA" ID-DOI="http://doi.org/10.5281/zenodo.4324073" ID-GBIF-Taxon="167292782" ID-Zenodo-Dep="4324073" LSID="urn:lsid:zoobank.org:act:88532963-8F7F-412B-A7BB-33CE812276BC" httpUri="http://treatment.plazi.org/id/7E421778FF8FFFB3FF10F915A68D58FA" lastPageId="3" lastPageNumber="179" pageId="1" pageNumber="177">
<subSubSection id="BEF1F5E5FF8FFFB1FF10F915A0685F8E" box="[151,524,1765,1792]" pageId="1" pageNumber="177" type="nomenclature">
<paragraph id="F654A66EFF8FFFB1FF10F915A0685F8E" blockId="1.[151,957,1765,1863]" box="[151,524,1765,1792]" pageId="1" pageNumber="177">
<heading id="AD1C1102FF8FFFB1FF10F915A0685F8E" bold="true" box="[151,524,1765,1792]" fontSize="11" level="1" pageId="1" pageNumber="177" reason="1">
<emphasis id="C49F7A7CFF8FFFB1FF10F915A0685F8E" bold="true" box="[151,524,1765,1792]" pageId="1" pageNumber="177">
<taxonomicName id="31EBDDEDFF8FFFB1FF10F915A3A65E71" ID-CoL="8LZ6Q" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[151,450,1765,1792]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="177" phylum="Arthropoda" rank="species" species="chapadensis" status="sp. nov.">
<emphasis id="C49F7A7CFF8FFFB1FF10F915A3A65E71" bold="true" box="[151,450,1765,1792]" italics="true" pageId="1" pageNumber="177">Hamadiana chapadensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFACC707FF8FFFB1FE4DF915A0685F8E" box="[458,524,1765,1791]" pageId="1" pageNumber="177" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB1FF10F8F9A1D95E36" pageId="1" pageNumber="177" type="description">
<paragraph id="F654A66EFF8FFFB1FF10F8F9A37D5E52" blockId="1.[151,957,1765,1863]" box="[151,281,1800,1827]" pageId="1" pageNumber="177">
(
<figureCitation id="6ED0BAEBFF8FFFB1FF18F8F9A3745E52" box="[159,272,1800,1827]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–7" captionStart-2="FIGURE 8–11" captionStart-3="FIGURES 12–15" captionStartId-0="4.[151,264,1536,1561]" captionStartId-1="5.[151,264,1803,1828]" captionStartId-2="6.[151,250,1220,1245]" captionStartId-3="6.[151,264,1972,1997]" captionTargetBox-0="[151,1435,538,1448]" captionTargetBox-1="[151,1435,181,1732]" captionTargetBox-2="[184,1320,233,1174]" captionTargetBox-3="[163,1335,1312,1937]" captionTargetId-0="figure@4.[151,1435,538,1448]" captionTargetId-1="figure@5.[151,1435,181,1732]" captionTargetId-2="figure@6.[169,1418,174,1209]" captionTargetId-3="figure@6.[151,1435,1296,1960]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionText-0="FIGURES 1–2. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." captionText-1="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." captionText-2="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." captionText-3="FIGURES 12–15. Hamadiana chapadensis sp. n.: 12) median lobe, dorsal view; 13) median lobe, left lateral view; 14) left paramere, left lateral view; 15) right paramere, left lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3995296" figureDoi-1="http://doi.org/10.5281/zenodo.3995298" figureDoi-2="http://doi.org/10.5281/zenodo.3995300" figureDoi-3="http://doi.org/10.5281/zenodo.3995302" httpUri-0="https://zenodo.org/record/3995296/files/figure.png" httpUri-1="https://zenodo.org/record/3995298/files/figure.png" httpUri-2="https://zenodo.org/record/3995300/files/figure.png" httpUri-3="https://zenodo.org/record/3995302/files/figure.png" pageId="1" pageNumber="177">Figs 1–15</figureCitation>
)
</paragraph>
<paragraph id="F654A66EFF8FFFB1FF10F8DCA1D95E36" blockId="1.[151,957,1765,1863]" box="[151,957,1836,1863]" pageId="1" pageNumber="177">
<uri id="827AAA6CFF8FFFB1FF10F8DCA1D95E36" box="[151,957,1836,1863]" pageId="1" pageNumber="177">
urn:lsid:zoobank.org:act:
<uuid id="824D9CBBFF8FFFB1FE21F8DCA1D95E36" box="[422,957,1836,1863]" pageId="1" pageNumber="177">88532963-8F7F-412B-A7BB-33CE812276BC</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8FFFB2FF10F885A7E959A6" lastPageId="2" lastPageNumber="178" pageId="1" pageNumber="177" type="materials_examined">
<paragraph id="F654A66EFF8FFFB1FF10F885A3795EC2" blockId="1.[151,1436,1908,1971]" pageId="1" pageNumber="177">
<emphasis id="C49F7A7CFF8FFFB1FF10F885A35F5EFE" bold="true" box="[151,315,1908,1935]" pageId="1" pageNumber="177">Type locality.</emphasis>
<materialsCitation id="4683AC33FF8FFFB1FEC1F885A0885EFE" ID-GBIF-Occurrence="2846841301" box="[326,748,1909,1935]" country="Brazil" location="Goias State" pageId="1" pageNumber="177" specimenCount="1" stateProvince="Goias">
<collectingCountry id="8EFCE6FEFF8FFFB1FEC1F885A3EB5EFE" box="[326,399,1909,1935]" name="Brazil" pageId="1" pageNumber="177">Brazil</collectingCountry>
:
<collectingRegion id="342F688CFF8FFFB1FE18F885A0435EFE" box="[415,551,1909,1935]" country="Brazil" name="Goias" pageId="1" pageNumber="177">Goiás State</collectingRegion>
, Alto Paraíso de
</materialsCitation>
<materialsCitation id="4683AC33FF8FFFB1FD7FF885A37C5EC2" ID-GBIF-Occurrence="2846841302" collectorName="County" country="Brazil" latitude="-14.283334" location="Goias" longLatPrecision="1288" longitude="-47.75" pageId="1" pageNumber="177" specimenCount="1" stateProvince="Goias">
<collectingRegion id="342F688CFF8FFFB1FD7FF885A15D5EFE" box="[760,825,1909,1935]" country="Brazil" name="Goias" pageId="1" pageNumber="177">Goiás</collectingRegion>
<collectorName id="5B1EC3B8FF8FFFB1FCC1F885A1FE5EFE" box="[838,922,1909,1935]" pageId="1" pageNumber="177">County</collectorName>
, “Rio dos Couros, Cachoeira” (ca.
<geoCoordinate id="93DFC0A9FF8FFFB1FAC7F885A7F85EFE" box="[1344,1436,1909,1935]" degrees="14" direction="south" minutes="17" orientation="latitude" pageId="1" pageNumber="177" precision="925" value="-14.283334">14°17’S</geoCoordinate>
<geoCoordinate id="93DFC0A9FF8FFFB1FF10F868A3745EC2" box="[151,272,1944,1971]" degrees="047" direction="west" minutes="45" orientation="longitude" pageId="1" pageNumber="177" precision="925" value="-47.75">047°45’W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FF69A7E959A6" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<materialsCitation id="4683AC33FF8CFFB2FF40FF69A7E959A6" ID-GBIF-Occurrence="2846841303" collectingDate="2017-06-21" collectionCode="INPA" collectorName="N. Hamada" country="Brazil" determinerName="Michat" location="Alto Paraiso" pageId="2" pageNumber="178" specimenCount="2" stateProvince="Goias" typeStatus="holotype">
<emphasis id="C49F7A7CFF8CFFB2FF40FF69A35F59C2" bold="true" box="[199,315,153,179]" pageId="2" pageNumber="178">
<typeStatus id="295018CCFF8CFFB2FF40FF69A35259C2" box="[199,310,153,179]" pageId="2" pageNumber="178" type="holotype">Holotype</typeStatus>
.
</emphasis>
m*, “
<collectingCountry id="8EFCE6FEFF8CFFB2FEFBFF69A3A059C2" box="[380,452,153,179]" name="Brazil" pageId="2" pageNumber="178">Brasil</collectingCountry>
,
<collectingRegion id="342F688CFF8CFFB2FE48FF69A07759C2" box="[463,531,153,179]" country="Brazil" name="Goias" pageId="2" pageNumber="178">Goiás</collectingRegion>
,
<location id="F334F0B5FF8CFFB2FD9BFF69A0C259C2" LSID="urn:lsid:plazi:treatment:7E421778FF8FFFB3FF10F915A68D58FA:F334F0B5FF8CFFB2FD9BFF69A0C259C2" box="[540,678,153,179]" country="Brazil" name="Alto Paraiso" pageId="2" pageNumber="178" stateProvince="Goias">Alto Paraíso</location>
de
<collectingRegion id="342F688CFF8CFFB2FD4AFF69A17559C2" box="[717,785,153,179]" country="Brazil" name="Goias" pageId="2" pageNumber="178">Goiás</collectingRegion>
, Rio dos Couros, cachoeira,
<date id="825580AEFF8CFFB2FBC9FF69A6AD59C2" box="[1102,1225,153,179]" pageId="2" pageNumber="178" value="2017-06-21">
<collectingDate id="92117946FF8CFFB2FBC9FF69A6AD59C2" box="[1102,1225,153,179]" pageId="2" pageNumber="178" value="2017-06-21">21.vi.2017</collectingDate>
</date>
, leg.
<collectorName id="5B1EC3B8FF8CFFB2FA84FF69A7E559C2" box="[1283,1409,153,179]" pageId="2" pageNumber="178">N. Hamada</collectorName>
<emphasis id="C49F7A7CFF8CFFB2FA0FFF6AA2C959A6" italics="true" pageId="2" pageNumber="178">et al</emphasis>
.”, “
<typeStatus id="295018CCFF8CFFB2FF5FFF4DA32259A6" box="[216,326,189,215]" pageId="2" pageNumber="178" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="31EBDDEDFF8CFFB2FED6FF4DA3B559A6" authority="Benetti &amp; Short &amp; Michat, 2019" authorityName="Benetti &amp; Short &amp; Michat" authorityYear="2019" box="[337,465,189,215]" class="Insecta" family="Dytiscidae" genus="Hamadiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="178" phylum="Arthropoda" rank="genus" status="gen. nov.">Hamadiana</taxonomicName>
<emphasis id="C49F7A7CFF8CFFB2FE5FFF4DA04959A6" bold="true" box="[472,557,189,215]" pageId="2" pageNumber="178">
<taxonomicNameLabel id="DFACC707FF8CFFB2FE5FFF4DA04359A6" box="[472,551,189,215]" pageId="2" pageNumber="178" rank="genus">gen. n.</taxonomicNameLabel>
,
</emphasis>
chapadensis
<emphasis id="C49F7A7CFF8CFFB2FD45FF4DA16659A6" bold="true" box="[706,770,189,215]" pageId="2" pageNumber="178">
<taxonomicNameLabel id="DFACC707FF8CFFB2FD45FF4DA16659A6" box="[706,770,189,215]" pageId="2" pageNumber="178" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Benetti, Short &amp;
<determinerName id="03884AAAFF8CFFB2FC4FFF4DA67359A6" box="[968,1047,189,215]" pageId="2" pageNumber="178">Michat</determinerName>
det. 2019” [red, printed] (
<collectionCode id="90FA3EABFF8CFFB2FABBFF4DA7E659A6" box="[1340,1410,189,215]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="178">INPA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8CFFB3FF40FF10A3FB598A" lastPageId="3" lastPageNumber="179" pageId="2" pageNumber="178" type="description">
<paragraph id="F654A66EFF8CFFB2FF40FF10A0905833" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FF10A33D598A" bold="true" box="[199,345,224,251]" pageId="2" pageNumber="178">Description.</emphasis>
<emphasis id="C49F7A7CFF8CFFB2FEE7FF11A3DD598A" box="[352,441,225,251]" italics="true" pageId="2" pageNumber="178">Habitus</emphasis>
(
<figureCitation id="6ED0BAEBFF8CFFB2FE40FF11A04E598A" box="[455,554,225,251]" captionStart="FIGURES 1–2" captionStartId="4.[151,264,1536,1561]" captionTargetBox="[151,1435,538,1448]" captionTargetId="figure@4.[151,1435,538,1448]" captionTargetPageId="4" captionText="FIGURES 1–2. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3995296" httpUri="https://zenodo.org/record/3995296/files/figure.png" pageId="2" pageNumber="178">Figs 1–2</figureCitation>
). Body shape oval, short and broad, gradually narrowed posteriorly, with apex slightly acuminate; lateral outline continuously curved from head to elytra; dorsally strongly convex, ventrally slightly convex. Maximum width at about mid length.
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FEBDA0F158FB" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FEBDA30D5817" box="[199,361,333,358]" italics="true" pageId="2" pageNumber="178">Measurements</emphasis>
(n = 1). TL:
<quantity id="31130B8BFF8CFFB2FE7CFEBCA0305816" box="[507,596,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="178" unit="mm" value="3.1">3.1 mm</quantity>
; TL-H:
<quantity id="31130B8BFF8CFFB2FD2AFEBBA1705816" box="[685,788,331,359]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="2" pageNumber="178" unit="mm" value="2.85">2.85 mm</quantity>
; MW:
<quantity id="31130B8BFF8CFFB2FCE7FEBCA1DD5816" box="[864,953,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="178" unit="mm" value="1.9">1.9 mm</quantity>
; EL:
<quantity id="31130B8BFF8CFFB2FC72FEBBA6385816" box="[1013,1116,331,359]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="2" pageNumber="178" unit="mm" value="2.35">2.35 mm</quantity>
; HW:
<quantity id="31130B8BFF8CFFB2FB24FEBCA6985816" box="[1187,1276,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="178" unit="mm" value="1.2">1.2 mm</quantity>
; PW:
<quantity id="31130B8BFF8CFFB2FAB8FEBCA7FC5816" box="[1343,1432,332,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="178" unit="mm" value="1.7">1.7 mm</quantity>
; EW:
<quantity id="31130B8BFF8CFFB2FF57FE9FA35358FB" box="[208,311,367,394]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="178" unit="mm" value="0.75">0.75 mm</quantity>
; TL/MW: 1.63; HW/EW: 1.61.
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FE64A6735B11" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FE64A36C58DF" box="[199,264,404,430]" italics="true" pageId="2" pageNumber="178">Color</emphasis>
(
<figureCitation id="6ED0BAEBFF8CFFB2FE90FE64A31D58DF" box="[279,377,404,430]" captionStart="FIGURES 1–2" captionStartId="4.[151,264,1536,1561]" captionTargetBox="[151,1435,538,1448]" captionTargetId="figure@4.[151,1435,538,1448]" captionTargetPageId="4" captionText="FIGURES 1–2. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3995296" httpUri="https://zenodo.org/record/3995296/files/figure.png" pageId="2" pageNumber="178">Figs 1, 2</figureCitation>
). Head orangish; pronotum brown with two broad dark yellow to orangish spots on lateral margins and one vague light spot on middle region; elytra brown, each with four well marked yellow spots (one basal, one submedian reaching lateral margin, one submedian near suture and one subapical) and two vague light spots on apical third (one elongate near suture and one on apex); prosternum, procoxae and basal 1/3 of epipleura dark yellow; metaventrite, metacoxae, abdominal ventrites and apical 3/4 of epipleura brown to reddish-brown (ventrite 1 and posterior margins of ventrites 3–5 somewhat lighter); legs reddish-brown.
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FD9AA3D05BD9" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FD9AA36F5BF5" box="[199,267,618,644]" italics="true" pageId="2" pageNumber="178">Head.</emphasis>
Surface smooth, shiny, without microreticulation, with fine impressed shallow punctures near anterior and inner margin of eyes.
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FD42A7285A62" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FD42A35A5BBA" box="[199,318,690,715]" italics="true" pageId="2" pageNumber="178">Pronotum.</emphasis>
Surface smooth, shiny, with fine microreticulation composed of polygonal cells; with fine regular micropunctation and well impressed wide punctures mostly along lateral sides and fore margin, arranged in more or less regular series. Basal margin nearly straight, lateral margins bordered, posterior angles rounded (
<figureCitation id="6ED0BAEBFF8CFFB2FB7EFD09A75A5A62" box="[1273,1342,761,787]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FCEDA3735D7C" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FCEDA3715A46" box="[199,277,797,823]" italics="true" pageId="2" pageNumber="178">Elytra.</emphasis>
Surface smooth, shiny, with fine microreticulation composed of polygonal cells; with fine, regular, shallow micropunctation and faintly impressed, hardly visible wide punctures, sparse and arranged in irregular, longitudinal series, mainly on apical third. Epipleuron without microreticulation, with some sparse punctures at middle. Scutellum not visible when elytra closed (
<figureCitation id="6ED0BAEBFF8CFFB2FDF0FC78A0DF5AD3" box="[631,699,904,930]" captionStart="FIGURES 1–2" captionStartId="4.[151,264,1536,1561]" captionTargetBox="[151,1435,538,1448]" captionTargetId="figure@4.[151,1435,538,1448]" captionTargetPageId="4" captionText="FIGURES 1–2. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3995296" httpUri="https://zenodo.org/record/3995296/files/figure.png" pageId="2" pageNumber="178">Fig. 1</figureCitation>
). Elytra elongate, apically slightly acuminate, attenuate and narrowly bordered laterally; suture between elytron and epipleuron faintly visible dorsally on apical third; epipleuron broad up to level of hind margin of metacoxal plates, then strongly narrowed, distinctly attenuate just before elytral tip (
<figureCitation id="6ED0BAEBFF8CFFB2FF43FC03A36D5D7C" box="[196,265,1011,1037]" captionStart="FIGURES 1–2" captionStartId="4.[151,264,1536,1561]" captionTargetBox="[151,1435,538,1448]" captionTargetId="figure@4.[151,1435,538,1448]" captionTargetPageId="4" captionText="FIGURES 1–2. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3995296" httpUri="https://zenodo.org/record/3995296/files/figure.png" pageId="2" pageNumber="178">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FBE7A3925CCB" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FBE7A3275D40" box="[199,323,1047,1073]" italics="true" pageId="2" pageNumber="178">Underside.</emphasis>
Surface smooth, shiny; microreticulation present on metacoxal processes and abdominal ventrites, composed of elongate cells; metaventrite with several punctures in middle region; metacoxa with stridulatory file in form of a series of parallel, closely spaced ridges (
<figureCitation id="6ED0BAEBFF8CFFB2FD4BFBAEA1755D09" box="[716,785,1117,1144]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Fig. 5</figureCitation>
); last abdominal ventrite with shallow scratchlike microsculpture on posterior half (
<figureCitation id="6ED0BAEBFF8CFFB2FE3AFB72A06B5DED" box="[445,527,1154,1180]" captionStart="FIGURE 8–11" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="2" pageNumber="178">Fig. 10</figureCitation>
). Prosternum and head separated by a shallow step; prosternum with small median ridge. Prosternal process elongate, lanceolate and bordered, clearly carinate, reaching mesocoxae but not extending beyond their hind margins; laterally compressed at level of procoxae, apically simple, with apex elongate, thornlike, very acute (
<figureCitation id="6ED0BAEBFF8CFFB2FED6FB1DA3FD5C76" box="[337,409,1261,1287]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Figs 4</figureCitation>
,
<figureCitation id="6ED0BAEBFF8CFFB2FE22FB1DA3D15C76" box="[421,437,1261,1287]" captionStart="FIGURE 8–11" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="2" pageNumber="178">8</figureCitation>
). Metasternal wings thin and arched; metacoxal lines converging anteriad in posterior 3/4 of their length, slightly diverging anteriad in anterior 1/4 (
<figureCitation id="6ED0BAEBFF8CFFB2FCA5FAE1A10F5C5A" box="[802,875,1296,1323]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Figs 5</figureCitation>
,
<figureCitation id="6ED0BAEBFF8CFFB2FCF0FAE1A1E35C5A" box="[887,903,1297,1323]" captionStart="FIGURE 8–11" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="2" pageNumber="178">9</figureCitation>
). Metacoxal process distally bilobed, with hind margin deeply incised and medially slightly protruded backwards (
<figureCitation id="6ED0BAEBFF8CFFB2FCF9FAC5A1A25C3E" box="[894,966,1332,1359]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Figs 5</figureCitation>
,
<figureCitation id="6ED0BAEBFF8CFFB2FC56FAC5A1855C3E" box="[977,993,1333,1359]" captionStart="FIGURE 8–11" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="2" pageNumber="178">9</figureCitation>
). Ventrite 3 with a medial protrusion on posterior margin (= median process raised apically) (
<figureCitation id="6ED0BAEBFF8CFFB2FD63FAA8A1435C02" box="[740,807,1368,1395]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Fig. 6</figureCitation>
) being probably a male sexual character; last abdominal ventrite as long as previous two ventrites combined, almost straight at sides, with anterolateral angles rounded and hind margin indented (
<figureCitation id="6ED0BAEBFF8CFFB2FE11FA50A38D5CCB" box="[406,489,1440,1466]" captionStart="FIGURE 8–11" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="2" pageNumber="178">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="F654A66EFF8CFFB2FF40FA34A0645F8D" blockId="2.[151,1437,153,2038]" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40FA34A3605CAC" box="[199,260,1476,1501]" italics="true" pageId="2" pageNumber="178">Legs.</emphasis>
Fore and middle legs long and slender; tarsomeres longer than wide, tarsomere 5 almost as long as tarsomeres 3 and 4 combined; pro- and mesotarsal claws slightly shorter than tarsomere 5; protarsal claws subequal in length and shape, unmodified; mesotibial spurs short, larger one not longer than first two tarsomeres together; posterior surface of mesofemora with a series of 3–4 long, stiff setae; swimming setae on mesotibiae present only apically. Metatibiae short, broad, about 2/3 as long as metafemur, with two simple apical spurs being acuminate at tip (
<figureCitation id="6ED0BAEBFF8CFFB2FF41F986A3695FE0" box="[198,269,1654,1681]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Fig. 6</figureCitation>
); metatarsomeres 1–4 with well developed, elongate lobes on posterolateral angles, each lobe bearing small set of flat spines and a strong spine on inner margin (
<figureCitation id="6ED0BAEBFF8CFFB2FCBBF96AA1E55FC4" box="[828,897,1690,1717]" captionStart="FIGURES 3–7" captionStartId="5.[151,264,1803,1828]" captionTargetBox="[151,1435,181,1732]" captionTargetId="figure@5.[151,1435,181,1732]" captionTargetPageId="5" captionText="FIGURES 3–7. Hamadiana chapadensis sp. n.: 3) posterior pronotal angle; 4) prosternal process; 5) metacoxa, arrow indicating stridulatory file; 6) abdomen and right hind-leg, arrows indicating simple apical tibial spurs and medial marginal protrusion on abdominal ventrite 3; 7) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995298" httpUri="https://zenodo.org/record/3995298/files/figure.png" pageId="2" pageNumber="178">Fig. 7</figureCitation>
); metatarsomere 5 bilobed, inner lobe long and acute, outer lobe about half shorter, bearing single, short, curved, apically rounded claw; claw about as long as outer lobe of metatarsomere (
<figureCitation id="6ED0BAEBFF8CFFB2FE26F911A3965F8D" box="[417,498,1761,1788]" captionStart="FIGURE 8–11" captionStartId="6.[151,250,1220,1245]" captionTargetBox="[184,1320,233,1174]" captionTargetId="figure@6.[169,1418,174,1209]" captionTargetPageId="6" captionText="FIGURE 8–11. Hamadiana chapadensis sp. n., ventral view: 8) prosternal process; 9) metacoxal processes; 10) last abdominal ventrite; 11) left metatarsus." figureDoi="http://doi.org/10.5281/zenodo.3995300" httpUri="https://zenodo.org/record/3995300/files/figure.png" pageId="2" pageNumber="178">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="F654A66EFF8CFFB3FF40F8F5A1E059A6" blockId="2.[151,1437,153,2038]" lastBlockId="3.[151,1436,152,395]" lastPageId="3" lastPageNumber="179" pageId="2" pageNumber="178">
<emphasis id="C49F7A7CFF8CFFB2FF40F8F5A36C5E6E" box="[199,264,1797,1823]" italics="true" pageId="2" pageNumber="178">Male.</emphasis>
Pro- and mesotarsomeres 1–3 not dilated, each one bearing ventrally a pair of stalked suction palettes (
<figureCitation id="6ED0BAEBFF8CFFB2FF27F8D9A28C5E32" box="[160,232,1833,1859]" captionStart="FIGURES 1–2" captionStartId="4.[151,264,1536,1561]" captionTargetBox="[151,1435,538,1448]" captionTargetId="figure@4.[151,1435,538,1448]" captionTargetPageId="4" captionText="FIGURES 1–2. Hamadiana chapadensis sp. n.: 1) habitus, dorsal view; 2) habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3995296" httpUri="https://zenodo.org/record/3995296/files/figure.png" pageId="2" pageNumber="178">Fig. 2</figureCitation>
). Aedeagus asymmetrical as in other genera of
<taxonomicName id="31EBDDEDFF8CFFB2FC94F8D9A1C05E32" baseAuthorityName="Miller &amp; Bergsten" baseAuthorityYear="2016" box="[787,932,1833,1859]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="178" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
. Median lobe in dorsal view (
<figureCitation id="6ED0BAEBFF8CFFB2FA84F8D9A73C5E32" box="[1283,1368,1833,1859]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1972,1997]" captionTargetBox="[163,1335,1312,1937]" captionTargetId="figure@6.[151,1435,1296,1960]" captionTargetPageId="6" captionText="FIGURES 12–15. Hamadiana chapadensis sp. n.: 12) median lobe, dorsal view; 13) median lobe, left lateral view; 14) left paramere, left lateral view; 15) right paramere, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3995302" httpUri="https://zenodo.org/record/3995302/files/figure.png" pageId="2" pageNumber="178">Fig. 12</figureCitation>
) with lateral margins slightly curved, basal 1/2 broader, distal 1/2 narrower, tip slightly curved; in lateral view (
<figureCitation id="6ED0BAEBFF8CFFB2FAC5F8BDA7F75E16" box="[1346,1427,1869,1895]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1972,1997]" captionTargetBox="[163,1335,1312,1937]" captionTargetId="figure@6.[151,1435,1296,1960]" captionTargetPageId="6" captionText="FIGURES 12–15. Hamadiana chapadensis sp. n.: 12) median lobe, dorsal view; 13) median lobe, left lateral view; 14) left paramere, left lateral view; 15) right paramere, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3995302" httpUri="https://zenodo.org/record/3995302/files/figure.png" pageId="2" pageNumber="178">Fig. 13</figureCitation>
) elongate, with curved thicker base, distal portion subparallel at basal 2/3, narrowed towards apex at distal 1/3, ending in curved rounded apex; ventral margin sinuous, dorsal margin straight in middle region, with small protrusion at distal 1/4 near apex. Parameres short, right one larger, both reaching less than half of length of median lobe. Left paramere (
<figureCitation id="6ED0BAEBFF8CFFB2FE89F82CA33B5E87" box="[270,351,2012,2038]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1972,1997]" captionTargetBox="[163,1335,1312,1937]" captionTargetId="figure@6.[151,1435,1296,1960]" captionTargetPageId="6" captionText="FIGURES 12–15. Hamadiana chapadensis sp. n.: 12) median lobe, dorsal view; 13) median lobe, left lateral view; 14) left paramere, left lateral view; 15) right paramere, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3995302" httpUri="https://zenodo.org/record/3995302/files/figure.png" pageId="2" pageNumber="178">Fig. 14</figureCitation>
) in lateral view with ventral margin almost straight and dorsal margin convex; with small pointed apex and two long setae at tip. Right paramere (
<figureCitation id="6ED0BAEBFF8DFFB3FD28FF69A16459C2" box="[687,768,152,179]" captionStart="FIGURES 12–15" captionStartId="6.[151,264,1972,1997]" captionTargetBox="[163,1335,1312,1937]" captionTargetId="figure@6.[151,1435,1296,1960]" captionTargetPageId="6" captionText="FIGURES 12–15. Hamadiana chapadensis sp. n.: 12) median lobe, dorsal view; 13) median lobe, left lateral view; 14) left paramere, left lateral view; 15) right paramere, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3995302" httpUri="https://zenodo.org/record/3995302/files/figure.png" pageId="3" pageNumber="179">Fig. 15</figureCitation>
) in lateral view subtriangular, wider at base, ventral margin almost straight, dorsal margin slightly sinuous; lacking setae at tip.
</paragraph>
<paragraph id="F654A66EFF8DFFB3FF40FF11A3FB598A" blockId="3.[151,1436,152,395]" box="[199,415,225,251]" pageId="3" pageNumber="179">
<emphasis id="C49F7A7CFF8DFFB3FF40FF11A347598A" box="[199,291,225,251]" italics="true" pageId="3" pageNumber="179">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8DFFB3FF40FEF5A76E5832" pageId="3" pageNumber="179" type="etymology">
<paragraph id="F654A66EFF8DFFB3FF40FEF5A76E5832" blockId="3.[151,1436,152,395]" pageId="3" pageNumber="179">
<emphasis id="C49F7A7CFF8DFFB3FF40FEF5A32A586E" bold="true" box="[199,334,261,287]" pageId="3" pageNumber="179">Etymology.</emphasis>
This species is named after the “Chapada dos Veadeiros”, a plateau in northeast of the
<collectingRegion id="342F688CFF8DFFB3FA9DFEF5A7FC586E" box="[1306,1432,261,287]" country="Brazil" name="Goias" pageId="3" pageNumber="179">Goiás state</collectingRegion>
, Central
<collectingCountry id="8EFCE6FEFF8DFFB3FF76FED9A35C5832" box="[241,312,297,323]" name="Brazil" pageId="3" pageNumber="179">Brazil</collectingCountry>
, where the
<typeStatus id="295018CCFF8DFFB3FE3FFED9A38C5832" box="[440,488,297,323]" pageId="3" pageNumber="179">type</typeStatus>
locality is located. The name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="BEF1F5E5FF8DFFB3FF40FEBCA68D58FA" pageId="3" pageNumber="179" type="distribution">
<paragraph id="F654A66EFF8DFFB3FF40FEBCA68D58FA" blockId="3.[151,1436,152,395]" pageId="3" pageNumber="179">
<emphasis id="C49F7A7CFF8DFFB3FF40FEBCA3945816" bold="true" box="[199,496,332,359]" pageId="3" pageNumber="179">Distribution and ecology.</emphasis>
Central
<collectingCountry id="8EFCE6FEFF8DFFB3FDC9FEBDA0F15816" box="[590,661,333,359]" name="Brazil" pageId="3" pageNumber="179">Brazil</collectingCountry>
, currently only known from the state of
<collectingRegion id="342F688CFF8DFFB3FBCAFEBDA6EA5816" box="[1101,1166,333,359]" country="Brazil" name="Goias" pageId="3" pageNumber="179">Goiás</collectingRegion>
(
<figureCitation id="6ED0BAEBFF8DFFB3FB1AFEBDA68B5819" box="[1181,1263,333,360]" captionStart="FIGURE 16" captionStartId="7.[151,250,1345,1370]" captionTargetBox="[151,1435,181,1325]" captionTargetId="figure@7.[151,1435,181,1325]" captionTargetPageId="7" captionText="FIGURE 16. Type locality of Hamadiana chapadensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.3995304" httpUri="https://zenodo.org/record/3995304/files/figure.png" pageId="3" pageNumber="179">Fig. 16</figureCitation>
). The specimen was collected in a river, in rapids (
<figureCitation id="6ED0BAEBFF8DFFB3FD9FFE81A00E58FA" box="[536,618,369,395]" captionStart="FIGURE 17" captionStartId="8.[151,250,1164,1189]" captionTargetBox="[151,1435,181,1145]" captionTargetId="figure@8.[151,1435,181,1145]" captionTargetPageId="8" captionText="FIGURE 17. Habitat of Hamadiana chapadensis sp. n. Brazil: Goiás State, Alto Paraíso de Goiás County, “Rio dos Couros”." figureDoi="http://doi.org/10.5281/zenodo.3995306" httpUri="https://zenodo.org/record/3995306/files/figure.png" pageId="3" pageNumber="179">Fig. 17</figureCitation>
) at
<quantity id="31130B8BFF8DFFB3FD13FE80A0BF58FA" box="[660,731,368,395]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="179" unit="m" value="950.0">950 m</quantity>
a.s.l., in the tropical savanna biome “cerrado”.
</paragraph>
</subSubSection>
</treatment>
</document>
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