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<document id="67DB0EE782B45A946BC457C4573105AE" ID-CLB-Dataset="299337" ID-DOI="10.5281/zenodo.11067222" ID-GBIF-Dataset="1cb742f7-205f-4cc5-9136-b613d5ab059d" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067222" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052073787" checkinUser="felipe" docAuthor="LaBonte, James R. &amp; Maddison, David R." docDate="2023" docId="51533818FFFC4766FF85BAFAFDCCFED7" docLanguage="en" docName="ProcCalifAcadSci.67.18.401-432.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (18)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Medusapyga chehalis LaBonte 2023, new species" docType="treatment" docVersion="7" lastPageNumber="419" masterDocId="AD6A4060FFED4775FFFEBA21FFB5FFA3" masterDocTitle="Medusapyga LaBonte and Maddison, a New Genus of Anillini (Coleoptera: Carabidae: Trechinae) from the Pacific Northwest of the United States" masterLastPageNumber="432" masterPageNumber="401" pageNumber="418" updateTime="1719875109990" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A7EAAF1711CFBDC8C3B02A3C2FCBF0EC">Medusapyga LaBonte and Maddison, a New Genus of Anillini (Coleoptera: Carabidae: Trechinae) from the Pacific Northwest of the United States</mods:title>
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<mods:namePart id="55583E513C0F32FF4B62189A5A326B4F">LaBonte, James R.</mods:namePart>
<mods:affiliation id="F092BCB597238E54C7F94CD6A1C0C450">1570 Kenard Street NW, Salem, OR 97304, USA.</mods:affiliation>
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<mods:namePart id="F4CF624A28EF049E0187CCDA11986CB8">Maddison, David R.</mods:namePart>
<mods:affiliation id="0BE0861F14148B9C4EFD7D9A9F355E55">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA.</mods:affiliation>
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<mods:title id="478D91D60C5EB56E08CCC2F6F1085375">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="5860367C736C3AF53896EB1A4F6AFB4B">
<mods:date id="362DDB83E1C14C8B004E2E2B3DA886B0">2023</mods:date>
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<mods:number id="C2DEB36CAF7D86505C2026D22B4D229A">2023-02-15</mods:number>
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<emphasis id="EB8E551CFFFC4764FF85BAFAFEDCFF50" bold="true" box="[123,361,219,243]" italics="true" pageId="17" pageNumber="418">Medusapyga chehalis</emphasis>
LaBonte
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,
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.
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, deposited in the California Academy of Sciences, labeled: “
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NW Littlerock
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, elev. ~
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and
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”, “HOLOTYPE,
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<emphasis id="EB8E551CFFFC4764FDA7BB60FCF2FEFA" box="[601,839,321,345]" italics="true" pageId="17" pageNumber="418">Medusapyga chehalis</emphasis>
LaBonte
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59 (
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,
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), residing in the following collections and institutions:
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(4),
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(2),
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(2),
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(39),
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<collectionCode id="BFEB11CBFFFC4764FDAABB80FD14FE1A" box="[596,673,417,441]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="17" pageNumber="418" type="Museum">LACM</collectionCode>
(2),
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(2),
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(2),
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(4),
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(2).
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<collectingCounty id="3024F182FFFC4764FE72BBE0FD96FE7A" box="[396,547,449,473]" pageId="17" pageNumber="418">Thurston Co.</collectingCounty>
,
<locationDeviation id="C8F7902CFFFC4764FDC8BBE0FCD4FE7A" box="[566,865,449,473]" location="Littlerock" pageId="17" pageNumber="418">
<quantity id="1E0224EBFFFC4764FDC8BBE0FD32FE7A" box="[566,647,449,473]" metricMagnitude="3" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="418" unit="km" value="9.5">9.5 km</quantity>
NW of Littlerock
</locationDeviation>
,
<collectingMunicipality id="39211374FFFC4764FC8DBBE0FB24FE7A" box="[883,1169,449,473]" pageId="17" pageNumber="418">Falls Creek Campground</collectingMunicipality>
,
<geoCoordinate id="BCCEEFC9FFFC4764FF85BBC0FF5AFE5A" box="[123,239,481,505]" degrees="46.9333" direction="north" orientation="latitude" pageId="17" pageNumber="418" precision="5" value="46.9333">46.9333°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFC4764FF02BBC0FE31FE5A" box="[252,388,472,505]" degrees="123.1333" direction="west" orientation="longitude" pageId="17" pageNumber="418" precision="5" value="-123.1333">123.1333°W</geoCoordinate>
,
<quantity id="1E0224EBFFFC4764FE6ABBC0FE6BFE5A" box="[404,478,481,505]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="418" unit="m" value="200.0">
<elevation id="52D76E3DFFFC4764FE6ABBC0FE6BFE5A" box="[404,478,481,505]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="418" unit="m" value="200.0">200 m</elevation>
</quantity>
elevation,
<date id="AD44AFCEFFFC4764FDA1BBC0FD53FE5A" box="[607,742,481,505]" pageId="17" pageNumber="418" value="2000-04-30">
<collectingDate id="BD005626FFFC4764FDA1BBC0FD53FE5A" box="[607,742,481,505]" pageId="17" pageNumber="418" value="2000-04-30">30 IV 2000</collectingDate>
</date>
,
<collectorName id="740FECD8FFFC4764FD09BBC3FCDBFE5A" box="[759,878,481,505]" pageId="17" pageNumber="418">J. Longino</collectorName>
(
<specimenCount id="CFFC4287FFFC4764FC7ABBC0FC27FE5A" box="[900,914,481,505]" pageId="17" pageNumber="418">2</specimenCount>
) (#4212, ENT 145103
<collectionCode id="BFEB11CBFFFC4764FF85B820FF78FDBA" box="[123,205,513,537]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="17" pageNumber="418" type="Museum">LACM</collectionCode>
).
</materialsCitation>
<materialsCitation id="69928353FFFC4764FF21B820FBAEFD9A" ID-GBIF-Occurrence="4900808347" collectingDate="2007-05-06" collectingDateMax="2022-04-02" collectingDateMin="2000-04-30" collectionCode="CAS, CMNH, CNC, JRLC, LACM, MCZ, NHMUK, OSAC, WSU, R" collectorName="D. J. Cox &amp; J. R. LaBonte" country="United States of America" county="Thurston Co." elevation="389" latitude="46.962" location="Thurston Co." longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" pageId="17" pageNumber="418" specimenCount="4" specimenCount-female="32" specimenCount-male="27" stateProvince="Washington" typeStatus="paratype">
<collectingCountry id="A1EDC99EFFFC4764FF21B820FEA2FDBA" box="[223,279,513,537]" name="United States of America" pageId="17" pageNumber="418">USA</collectingCountry>
,
<collectingRegion id="1B3E47ECFFFC4764FEDCB820FE11FDBA" box="[290,420,513,537]" country="United States of America" name="Washington" pageId="17" pageNumber="418">Washington</collectingRegion>
,
<location id="DC25DFD5FFFC4764FE51B820FD8AFDBA" LSID="urn:lsid:plazi:treatment:51533818FFFC4766FF85BAFAFDCCFED7:DC25DFD5FFFC4764FE51B820FD8AFDBA" box="[431,575,513,537]" country="United States of America" county="Thurston Co." latitude="46.962" longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" name="Thurston Co." pageId="17" pageNumber="418" stateProvince="Washington">Thurston Co.</location>
,
<locationDeviation id="C8F7902CFFFC4764FDB5B820FCD2FDBA" box="[587,871,513,537]" location="Littlerock" pageId="17" pageNumber="418">
<quantity id="1E0224EBFFFC4764FDB5B820FD17FDBA" box="[587,674,513,537]" metricMagnitude="4" metricUnit="m" metricValue="1.13" pageId="17" pageNumber="418" unit="km" value="11.3">11.3 km</quantity>
NW of Littlerock
</locationDeviation>
,
<geoCoordinate id="BCCEEFC9FFFC4764FC8DB820FC41FDBA" box="[883,1012,513,537]" degrees="46.96000" direction="north" orientation="latitude" pageId="17" pageNumber="418" precision="1" value="46.96">46.96000°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFC4764FC02B820FB24FDBA" box="[1020,1169,513,537]" degrees="123.13495" direction="west" orientation="longitude" pageId="17" pageNumber="418" precision="1" value="-123.13495">123.13495°W</geoCoordinate>
, between
<quantity id="1E0224EBFFFC4764FF27B800FEE7FD9A" box="[217,338,545,569]" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" metricValueMax="4.12" metricValueMin="3.66" pageId="17" pageNumber="418" unit="m" value="389.0" valueMax="412.0" valueMin="366.0">
<elevation id="52D76E3DFFFC4764FF27B800FEE7FD9A" box="[217,338,545,569]" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" metricValueMax="4.12" metricValueMin="3.66" pageId="17" pageNumber="418" unit="m" value="389.0" valueMax="412.0" valueMin="366.0">366412 m</elevation>
</quantity>
elevation [Type locality].
<date id="AD44AFCEFFFC4764FD96B800FD7EFD9A" box="[616,715,545,569]" pageId="17" pageNumber="418" value="2007-05-06">
<collectingDate id="BD005626FFFC4764FD96B800FD7EFD9A" box="[616,715,545,569]" pageId="17" pageNumber="418" value="2007-05-06">6 V 2007</collectingDate>
</date>
,
<collectorName id="740FECD8FFFC4764FD2BB803FC87FD9A" box="[725,818,545,569]" pageId="17" pageNumber="418">D.J. Cox</collectorName>
and
<collectorName id="740FECD8FFFC4764FC9AB803FC45FD9A" box="[868,1008,546,569]" pageId="17" pageNumber="418">J.R. LaBonte</collectorName>
(
<specimenCount id="CFFC4287FFFC4764FC00B800FBBBFD9A" box="[1022,1038,545,569]" pageId="17" pageNumber="418">4</specimenCount>
).
</materialsCitation>
<materialsCitation id="69928353FFFC4764FBDFB800FE87FDFA" ID-GBIF-Occurrence="4900808326" collectingDate="2007-05-12" collectingDateMax="2022-04-02" collectingDateMin="2000-04-30" collectionCode="CAS, CMNH, CNC, JRLC, LACM, MCZ, NHMUK, OSAC, WSU, R" collectorName="J. R. LaBonte" country="United States of America" county="Thurston Co." elevation="200" latitude="46.962" location="Thurston Co." longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" pageId="17" pageNumber="418" specimenCount="7" specimenCount-female="32" specimenCount-male="27" stateProvince="Washington" typeStatus="paratype">
<date id="AD44AFCEFFFC4764FBDFB800FB27FD9A" box="[1057,1170,545,569]" pageId="17" pageNumber="418" value="2007-05-12">
<collectingDate id="BD005626FFFC4764FBDFB800FB27FD9A" box="[1057,1170,545,569]" pageId="17" pageNumber="418" value="2007-05-12">12 V 2007</collectingDate>
</date>
,
<collectorName id="740FECD8FFFC4764FF85B863FEB3FDFA" box="[123,262,578,601]" pageId="17" pageNumber="418">J.R. LaBonte</collectorName>
(
<specimenCount id="CFFC4287FFFC4764FEEBB860FE91FDFA" box="[277,292,577,601]" pageId="17" pageNumber="418">7</specimenCount>
).
</materialsCitation>
<materialsCitation id="69928353FFFC4764FEC6B860FDDEFDFA" ID-GBIF-Occurrence="4900808315" box="[312,619,577,601]" collectingDate="2007-05-23" collectingDateMax="2022-04-02" collectingDateMin="2000-04-30" collectionCode="CAS, CMNH, CNC, JRLC, LACM, MCZ, NHMUK, OSAC, WSU, R" collectorName="J. R. LaBonte" country="United States of America" county="Thurston Co." elevation="200" latitude="46.962" location="Thurston Co." longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" pageId="17" pageNumber="418" specimenCount="6" specimenCount-female="32" specimenCount-male="27" stateProvince="Washington" typeStatus="paratype">
<date id="AD44AFCEFFFC4764FEC6B860FE1CFDFA" box="[312,425,577,601]" pageId="17" pageNumber="418" value="2007-05-23">
<collectingDate id="BD005626FFFC4764FEC6B860FE1CFDFA" box="[312,425,577,601]" pageId="17" pageNumber="418" value="2007-05-23">23 V 2007</collectingDate>
</date>
,
<collectorName id="740FECD8FFFC4764FE4AB863FD8AFDFA" box="[436,575,578,601]" pageId="17" pageNumber="418">J.R. LaBonte</collectorName>
(
<specimenCount id="CFFC4287FFFC4764FDB0B860FDEBFDFA" box="[590,606,577,601]" pageId="17" pageNumber="418">6</specimenCount>
).
</materialsCitation>
<materialsCitation id="69928353FFFC4764FD8FB860FC7DFDD9" ID-GBIF-Occurrence="4900808351" collectingDate="2022-04-02" collectingDateMax="2022-04-02" collectingDateMin="2000-04-30" collectionCode="CAS, CMNH, CNC, JRLC, LACM, MCZ, NHMUK, OSAC, WSU, R" collectorName="J. R. LaBonte" country="United States of America" county="Thurston Co." elevation="315" latitude="46.962" location="Thurston Co." longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" pageId="17" pageNumber="418" specimenCount="4" specimenCount-female="32" specimenCount-male="27" stateProvince="Washington" typeStatus="paratype">
<collectingCountry id="A1EDC99EFFFC4764FD8FB860FD1CFDFA" box="[625,681,577,601]" name="United States of America" pageId="17" pageNumber="418">USA</collectingCountry>
,
<collectingRegion id="1B3E47ECFFFC4764FD4DB860FC80FDFA" box="[691,821,577,601]" country="United States of America" name="Washington" pageId="17" pageNumber="418">Washington</collectingRegion>
,
<location id="DC25DFD5FFFC4764FCC0B860FC79FDFA" LSID="urn:lsid:plazi:treatment:51533818FFFC4766FF85BAFAFDCCFED7:DC25DFD5FFFC4764FCC0B860FC79FDFA" box="[830,972,577,601]" country="United States of America" county="Thurston Co." latitude="46.962" longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" name="Thurston Co." pageId="17" pageNumber="418" stateProvince="Washington">Thurston Co.</location>
,
<locationDeviation id="C8F7902CFFFC4764FC29B860FF7DFDD9" location="Littlerock" pageId="17" pageNumber="418">
<quantity id="1E0224EBFFFC4764FC29B860FBACFDFA" box="[983,1049,577,601]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="418" unit="km" value="11.0">11 km</quantity>
NW of Littlerock
</locationDeviation>
,
<geoCoordinate id="BCCEEFC9FFFC4764FF2CB843FEF3FDD9" box="[210,326,610,634]" degrees="46.9564" direction="north" orientation="latitude" pageId="17" pageNumber="418" precision="5" value="46.9564">46.9564°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFC4764FEB3B843FE60FDD9" box="[333,469,610,634]" degrees="123.1366" direction="west" orientation="longitude" pageId="17" pageNumber="418" precision="5" value="-123.1366">123.1366°W</geoCoordinate>
,
<quantity id="1E0224EBFFFC4764FE21B843FD96FDDA" box="[479,547,610,634]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="17" pageNumber="418" unit="m" value="315.0">
<elevation id="52D76E3DFFFC4764FE21B843FD96FDDA" box="[479,547,610,634]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="17" pageNumber="418" unit="m" value="315.0">315 m</elevation>
</quantity>
elevation,
<date id="AD44AFCEFFFC4764FD66B843FCB0FDD9" box="[664,773,610,634]" pageId="17" pageNumber="418" value="2022-04-02">
<collectingDate id="BD005626FFFC4764FD66B843FCB0FDD9" box="[664,773,610,634]" pageId="17" pageNumber="418" value="2022-04-02">2 IV 2022</collectingDate>
</date>
,
<collectorName id="740FECD8FFFC4764FCEEB843FC29FDDA" box="[784,924,610,633]" pageId="17" pageNumber="418">J.R. LaBonte</collectorName>
(
<specimenCount id="CFFC4287FFFC4764FC55B843FC0EFDD9" box="[939,955,610,634]" pageId="17" pageNumber="418">4</specimenCount>
).
</materialsCitation>
<materialsCitation id="69928353FFFC4764FC30B843FEB1FD79" ID-GBIF-Occurrence="4900808327" collectingDate="2014-04-28" collectingDateMax="2022-04-02" collectingDateMin="2000-04-30" collectionCode="CAS, CMNH, CNC, JRLC, LACM, MCZ, NHMUK, OSAC, WSU, R" collectorName="D. H. Kavanaugh &amp; J. R. LaBonte" country="United States of America" county="Thurston Co." elevation="301" latitude="46.962" location="Thurston Co." longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" pageId="17" pageNumber="418" specimenCount="27" specimenCount-female="32" specimenCount-male="27" stateProvince="Washington" typeStatus="paratype">
<collectingCountry id="A1EDC99EFFFC4764FC30B843FBB3FDD9" box="[974,1030,610,634]" name="United States of America" pageId="17" pageNumber="418">USA</collectingCountry>
,
<collectingRegion id="1B3E47ECFFFC4764FBEEB843FB27FDD9" box="[1040,1170,610,634]" country="United States of America" name="Washington" pageId="17" pageNumber="418">Washington</collectingRegion>
,
<location id="DC25DFD5FFFC4764FF85B8A3FEBEFD39" LSID="urn:lsid:plazi:treatment:51533818FFFC4766FF85BAFAFDCCFED7:DC25DFD5FFFC4764FF85B8A3FEBEFD39" box="[123,267,642,666]" country="United States of America" county="Thurston Co." latitude="46.962" longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" name="Thurston Co." pageId="17" pageNumber="418" stateProvince="Washington">Thurston Co.</location>
,
<locationDeviation id="C8F7902CFFFC4764FEE9B8A3FD8EFD39" box="[279,571,642,666]" location="Littlerock" pageId="17" pageNumber="418">
<quantity id="1E0224EBFFFC4764FEE9B8A3FED7FD39" box="[279,354,642,666]" metricMagnitude="3" metricUnit="m" metricValue="9.3" pageId="17" pageNumber="418" unit="km" value="9.3">9.3 km</quantity>
NNW of Littlerock
</locationDeviation>
,
<geoCoordinate id="BCCEEFC9FFFC4764FDB9B8A3FD0EFD39" box="[583,699,642,666]" degrees="46.9620" direction="north" orientation="latitude" pageId="17" pageNumber="418" precision="5" value="46.962">46.9620°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFC4764FD3DB8A3FCFEFD39" box="[707,843,642,666]" degrees="123.1084" direction="west" orientation="longitude" pageId="17" pageNumber="418" precision="5" value="-123.1084">123.1084°W</geoCoordinate>
,
<quantity id="1E0224EBFFFC4764FCA9B8A3FC29FD3A" box="[855,924,642,666]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="17" pageNumber="418" unit="m" value="301.0">
<elevation id="52D76E3DFFFC4764FCA9B8A3FC29FD3A" box="[855,924,642,666]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="17" pageNumber="418" unit="m" value="301.0">301 m</elevation>
</quantity>
elevation,
<date id="AD44AFCEFFFC4764FBEDB8A3FB24FD39" box="[1043,1169,642,666]" pageId="17" pageNumber="418" value="2014-04-28">
<collectingDate id="BD005626FFFC4764FBEDB8A3FB24FD39" box="[1043,1169,642,666]" pageId="17" pageNumber="418" value="2014-04-28">28 IV 2014</collectingDate>
</date>
, J.R. LaBonte (
<specimenCount id="CFFC4287FFFC4764FEE0B883FE89FD19" box="[286,316,674,698]" pageId="17" pageNumber="418">27</specimenCount>
, 8 of which were extracted from soil and litter)
<collectorName id="740FECD8FFFC4764FC86B882FB85FD19" box="[888,1072,674,698]" pageId="17" pageNumber="418">D.H. Kavanaugh</collectorName>
and
<collectorName id="740FECD8FFFC4764FB92B882FF62FD79" pageId="17" pageNumber="418">J.R. LaBonte</collectorName>
(4).
</materialsCitation>
<materialsCitation id="69928353FFFC4764FEF5B8E3FBFBFD79" ID-GBIF-Occurrence="4900808342" box="[267,1102,706,730]" collectingDate="2021-04-22" collectingDateMax="2022-04-02" collectingDateMin="2000-04-30" collectionCode="CAS, CMNH, CNC, JRLC, LACM, MCZ, NHMUK, OSAC, WSU, R" collectorName="J. R. LaBonte" country="United States of America" county="Thurston Co." elevation="200" latitude="46.962" location="Thurston Co." longLatPrecision="6" longitude="-123.1084" municipality="Falls Creek Campground" pageId="17" pageNumber="418" specimenCount="3" specimenCount-female="32" specimenCount-male="27" stateProvince="Washington" typeStatus="paratype">
<date id="AD44AFCEFFFC4764FEF5B8E3FE3DFD79" box="[267,392,706,730]" pageId="17" pageNumber="418" value="2021-04-22">
<collectingDate id="BD005626FFFC4764FEF5B8E3FE3DFD79" box="[267,392,706,730]" pageId="17" pageNumber="418" value="2021-04-22">22 IV 2021</collectingDate>
</date>
,
<collectorName id="740FECD8FFFC4764FE6DB8E2FD95FD79" box="[403,544,707,730]" pageId="17" pageNumber="418">J.R. LaBonte</collectorName>
(
<specimenCount id="CFFC4287FFFC4764FDCFB8E3FD8AFD79" box="[561,575,706,730]" pageId="17" pageNumber="418">3</specimenCount>
, all of which were extracted from soil and litter)
</materialsCitation>
.
</paragraph>
<paragraph id="D945890EFFFC4764FF55B8C3FC20FCB9" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">Depending upon the GPS system used, the exact geographic coordinates and elevations may slightly differ among the labels of the different collectors and institutions.</paragraph>
<paragraph id="D945890EFFFC4764FF55B902FE64FCF9" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55B902FEE7FC99" bold="true" box="[171,338,803,826]" pageId="17" pageNumber="418">Type Locality.</emphasis>
<materialsCitation id="69928353FFFC4764FEA1B903FE64FCF9" ID-GBIF-Occurrence="4900808348" country="United States of America" county="Thurston Co." elevation="400" latitude="46.96" location="Thurston Co." longLatPrecision="6" longitude="-123.135" pageId="17" pageNumber="418" specimenCount="400" stateProvince="Washington">
<collectingCountry id="A1EDC99EFFFC4764FEA1B903FE22FC99" box="[351,407,802,826]" name="United States of America" pageId="17" pageNumber="418">USA</collectingCountry>
,
<collectingRegion id="1B3E47ECFFFC4764FE59B903FD9CFC99" box="[423,553,802,826]" country="United States of America" name="Washington" pageId="17" pageNumber="418">Washington</collectingRegion>
,
<collectingCounty id="3024F182FFFC4764FDC7B903FD7BFC99" box="[569,718,802,826]" pageId="17" pageNumber="418">Thurston Co.</collectingCounty>
,
<locationDeviation id="C8F7902CFFFC4764FD21B903FBA5FC99" box="[735,1040,802,826]" location="Littlerock" pageId="17" pageNumber="418">
<quantity id="1E0224EBFFFC4764FD21B903FC89FC99" box="[735,828,802,826]" metricMagnitude="4" metricUnit="m" metricValue="1.13" pageId="17" pageNumber="418" unit="km" value="11.3">11.3 km</quantity>
NW of Littlerock
</locationDeviation>
,
<geoCoordinate id="BCCEEFC9FFFC4764FBDFB903FB20FC99" box="[1057,1173,802,826]" degrees="46.9600" direction="north" orientation="latitude" pageId="17" pageNumber="418" precision="5" value="46.96">46.9600°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFC4764FF85B963FEB6FCF9" box="[123,259,834,858]" degrees="123.1350" direction="west" orientation="longitude" pageId="17" pageNumber="418" precision="5" value="-123.135">123.1350°W</geoCoordinate>
, elevation ~
<specimenCount id="CFFC4287FFFC4764FE7AB963FE64FCF9" box="[388,465,834,858]" count="400" pageId="17" pageNumber="418" type="generic">
<quantity id="1E0224EBFFFC4764FE7AB963FE78FCF9" box="[388,461,834,858]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="418" unit="m" value="400.0">400 m</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="91E0DA85FFFC4764FF55B942FB36FC38" pageId="17" pageNumber="418" type="etymology">
<paragraph id="D945890EFFFC4764FF55B942FB36FC38" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55B942FE9EFCD9" bold="true" box="[171,299,867,890]" pageId="17" pageNumber="418">Etymology.</emphasis>
The specific epithet refers to the southwestern
<collectingRegion id="1B3E47ECFFFC4764FCD2B943FC1EFCD9" box="[812,939,866,890]" country="United States of America" name="Washington" pageId="17" pageNumber="418">Washington</collectingRegion>
river drainage of this name and a community near the
<typeStatus id="064137ACFFFC4764FE25B9A2FDBCFC38" box="[475,521,899,923]" pageId="17" pageNumber="418">type</typeStatus>
locality. The name is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="91E0DA85FFFC4766FF55B982FDCCFED7" lastPageId="19" lastPageNumber="420" pageId="17" pageNumber="418" type="description">
<paragraph id="D945890EFFFC4764FF55B982FE62FC78" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55B982FE83FC19" bold="true" box="[171,310,931,954]" pageId="17" pageNumber="418">Description.</emphasis>
<emphasis id="EB8E551CFFFC4764FEBEB982FE21FC18" box="[320,404,931,955]" italics="true" pageId="17" pageNumber="418">Habitus</emphasis>
(
<figureCitation id="41C1958BFFFC4764FE56B982FDB4FC18" box="[424,513,931,955]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="17" pageNumber="418">Fig. 2B</figureCitation>
). SBL
<quantity id="1E0224EBFFFC4764FDAAB982FD42FC19" box="[596,759,931,955]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" metricValueMax="2.35" metricValueMin="2.03" pageId="17" pageNumber="418" unit="mm" value="2.19" valueMax="2.35" valueMin="2.03">2.032.35 mm</quantity>
. Color translucent testaceous to dark brown. Flattened in lateral view.
</paragraph>
<paragraph id="D945890EFFFC4764FF55B9C2FDA2FB98" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55B9C2FF50FC58" box="[171,229,995,1019]" italics="true" pageId="17" pageNumber="418">Head</emphasis>
(
<figureCitation id="41C1958BFFFC4764FF0BB9C2FEFEFC58" box="[245,331,995,1019]" captionStart="FIGURE" captionStartId="11.[155,167,793,812]" captionTargetBox="[182,1123,229,767]" captionTargetId="figure-340@11.[172,1132,216,777]" captionTargetPageId="11" captionText="FIGURE 7. Dorsal views of heads. (A) Medusapyga alsea. (B) M. chehalis. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067232" httpUri="https://zenodo.org/record/11067232/files/figure.png" pageId="17" pageNumber="418">Fig. 7B</figureCitation>
). The medial convexity formed by the frontolateral furrows does not end in a frontoclypeal prominence as in
<taxonomicName id="1EFAF28DFFFC4764FE2BBE25FD87FBB8" box="[469,562,1027,1051]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFFC4764FE2BBE25FD87FBB8" box="[469,562,1027,1051]" italics="true" pageId="17" pageNumber="418">M. alsea</emphasis>
</taxonomicName>
but instead gradually and evenly decreases in elevation anteriorly. Tempora obtusely angulate.
</paragraph>
<paragraph id="D945890EFFFC4764FF55BE65FBF2FA1F" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55BE65FEA0FBF8" box="[171,277,1092,1115]" italics="true" pageId="17" pageNumber="418">Pronotum</emphasis>
(
<figureCitation id="41C1958BFFFC4764FEDABE62FECCFBF8" box="[292,377,1091,1115]" captionStart="FIGURE" captionStartId="12.[155,167,666,685]" captionTargetBox="[195,1128,229,656]" captionTargetId="figure-339@12.[181,1141,216,662]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal views of pronota. (A) Medusapyga alsea. (B) M. chehalis. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067234" httpUri="https://zenodo.org/record/11067234/files/figure.png" pageId="17" pageNumber="418">Fig. 8B</figureCitation>
). PWm/
<collectionCode id="BFEB11CBFFFC4764FE2FBE62FE45FBF8" box="[465,496,1091,1115]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="17" pageNumber="418" type="Museum">PL</collectionCode>
~1.30, PWa/PWp ~1.10. Widest at about anterior quarter, posterior of insertion of large anteriolateral setae. Anterior angles rounded and, as essentially a continuation of the slightly arcuate and emarginate anterior margin, protrude only slightly beyond the anteromedial margin. Lateral margins slightly sinuate, approximately parallel only just anterior of posterior angles, oblique to greatest pronotal width, then convergently arcuate to anterior angles. Lateral explanations distinctly narrowly reflexed, width more or less uniform throughout except broadened posterior of posteromedial impression, widest at hind angles. Hind angles right to slightly obtuse, rounded, anterior of posterior margin. Width at posterior margin less than at hind angles. Lateral margins with fine serrations at and just anterior of hind angles, extending to posterior margins of hind angles. Disc not as strongly convex as in
<taxonomicName id="1EFAF28DFFFC4764FD3ABF44FC95FADF" box="[708,800,1380,1404]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFFC4764FD3ABF44FC95FADF" box="[708,800,1380,1404]" italics="true" pageId="17" pageNumber="418">M. alsea</emphasis>
</taxonomicName>
. One long, stout seta well anterior of each posterior angle (at approximately the anterior delimitation of the posteromedian impression) and a longer seta located laterally at approximately the anterior seventh of each side.
</paragraph>
<paragraph id="D945890EFFFC4764FF55BFE5FE01F93E" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55BFE5FF5BFA7F" box="[171,238,1476,1500]" italics="true" pageId="17" pageNumber="418">Elytra</emphasis>
(
<figureCitation id="41C1958BFFFC4764FEFFBFE5FEECFA7F" box="[257,345,1476,1500]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="17" pageNumber="418">Fig. 2B</figureCitation>
). Greatest width at mid-length. Slightly convex in lateral view, shallowly ascending from anterior margin to about anterior third, thereafter very shallowly and evenly descending to apex, more or less evenly and slightly convex in cross section. Each elytron with lateral margin finely denticulate from humerus to plical crossing. Humeri arcuate and obtusely rounded. Lateral margins slightly arcuately posteriorly divergent in anterior half, thereafter lateral margins evenly convergently arcuate to the separately rounded apices. Only very faintly indented at the plical crossing. Intervals flat.
</paragraph>
<paragraph id="D945890EFFFC4767FF55BC87FDAAFE50" blockId="17.[123,1174,219,1725]" lastBlockId="18.[123,622,219,851]" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418">
<emphasis id="EB8E551CFFFC4764FF55BC87FF51F91E" box="[171,228,1702,1725]" italics="true" pageId="17" pageNumber="418">Legs.</emphasis>
Profemoral spine of males near basal one-sixth of ventral face, with its length aboutwice its greatest diameter (
<figureCitation id="41C1958BFFFF4767FE57BAFAFDB5FF50" box="[425,512,219,243]" captionStart="FIGURE" captionStartId="9.[157,169,1658,1677]" captionTargetBox="[129,1168,1135,1647]" captionTargetId="figure-299@9.[120,1176,1127,1655]" captionTargetPageId="9" captionText="FIGURE 4. Left profemora, anterior view. (A) Medusapyga alsea, male. (B) M. alsea, female. (C) M. chehalis, male. (D) M. chehalis, female. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067228" httpUri="https://zenodo.org/record/11067228/files/figure.png" pageId="18" pageNumber="419">Fig. 4C</figureCitation>
). Females lack the elongate profemoral spine of males; however, in the same position they have a distinct obtuse angulation that can vary in expression to a short denticle (
<figureCitation id="41C1958BFFFF4767FE84BB7AFE64FED0" box="[378,465,347,371]" captionStart="FIGURE" captionStartId="9.[157,169,1658,1677]" captionTargetBox="[129,1168,1135,1647]" captionTargetId="figure-299@9.[120,1176,1127,1655]" captionTargetPageId="9" captionText="FIGURE 4. Left profemora, anterior view. (A) Medusapyga alsea, male. (B) M. alsea, female. (C) M. chehalis, male. (D) M. chehalis, female. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067228" httpUri="https://zenodo.org/record/11067228/files/figure.png" pageId="18" pageNumber="419">Fig. 4D</figureCitation>
). Females also have a carina extending from the profemoral base to this angulation, similar to that found in males. Mesotrochanter with posterior margin more or less evenly arcuate (
<figureCitation id="41C1958BFFFF4767FE4EBBFAFDA6FE50" box="[432,531,475,499]" captionStart="FIGURE" captionStartId="16.[539,551,527,546]" captionTargetBox="[507,1171,219,511]" captionTargetId="figure-370@16.[504,1176,216,514]" captionText="FIGURE 13. Male mesotrochanter, ventral view. (A) Medusapyga alsea, left trochanter. (B) M. chehalis, right trochanter, digitally reversed to make the image more comparable to (A). Scale bars 25 µm." figureDoi="http://doi.org/10.5281/zenodo.12612696" httpUri="https://zenodo.org/record/12612696/files/figure.png" pageId="18" pageNumber="419">Fig. 13B</figureCitation>
).
</paragraph>
<paragraph id="D945890EFFFF4767FF55BBDAFDA7FD51" blockId="18.[123,622,219,851]" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FF55BBDAFE3AFDB0" box="[171,399,507,531]" italics="true" pageId="18" pageNumber="419">Abdominal ventrites.</emphasis>
Apex of intercoxal process of second visible abdominal ventrite obtuse. There is a very small, shallow fovea on this ventrite just posterior of the intercoxal process between the metacoxae. It is in the shape of a broad trapezoid narrowest at the anterior and broadest at its posterior. The surface of the fovea is coarsely punctate.
</paragraph>
<paragraph id="D945890EFFFF4767FF55B8DAFBADFC50" blockId="18.[123,622,219,851]" lastBlockId="18.[123,1174,923,1715]" pageId="18" pageNumber="419">
In males there is a small medial laterally compressed carina (the “keel”) located at about slightly more than 1/5 the total length of visible abdominal ventrite 2 from the posterior margin thereof (
<figureCitation id="41C1958BFFFF4767FED5B95AFE24FC30" box="[299,401,891,915]" captionStart="FIGURE" captionStartId="16.[539,551,1050,1069]" captionTargetBox="[505,1171,631,1031]" captionTargetId="figure-404@16.[504,1176,630,1035]" captionText="FIGURE 14. Abdominal keel of a Medusapyga chehalis male. (A) Ventral view second visible abdominal ventrite; arrow points toward abdominal keel. (B) Lateral view. Scale bars 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12612698" httpUri="https://zenodo.org/record/12612698/files/figure.png" pageId="18" pageNumber="419">Fig. 14A</figureCitation>
). This keel is tallest at its posterior extent and is an elongate triangle in lateral perspective (
<figureCitation id="41C1958BFFFF4767FE64B99AFE4BFC70" box="[410,510,955,979]" captionStart="FIGURE" captionStartId="16.[539,551,1050,1069]" captionTargetBox="[505,1171,631,1031]" captionTargetId="figure-404@16.[504,1176,630,1035]" captionText="FIGURE 14. Abdominal keel of a Medusapyga chehalis male. (A) Ventral view second visible abdominal ventrite; arrow points toward abdominal keel. (B) Lateral view. Scale bars 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12612698" httpUri="https://zenodo.org/record/12612698/files/figure.png" pageId="18" pageNumber="419">Fig. 14B</figureCitation>
). It is most often sharply pointed (slightly rounded in a few males) at the tallest point and is perpendicularly declivous to the plane of the ventrite.
</paragraph>
<caption id="8D85D986FFFF4767FD55B97DFCB8FC01" ID-DOI="http://doi.org/10.5281/zenodo.12612700" ID-Zenodo-Dep="12612700" httpUri="https://zenodo.org/record/12612700/files/figure.png" pageId="18" pageNumber="419" startId="18.[683,695,860,879]" targetBox="[706,1176,216,848]" targetPageId="18">
<paragraph id="D945890EFFFF4767FD55B97DFCB8FC01" pageId="18" pageNumber="419">
<smallCapsWord id="DFA31FD2FFFF4767FD55B97DFD5DFCCD" baselines="874,874" box="[683,744,860,879]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="18" pageNumber="419">FIGURE</smallCapsWord>
15. Known geographic distributions of
<emphasis id="EB8E551CFFFF4767FD75B957FD53FC2B" box="[651,742,885,904]" italics="true" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FBB5B97DFB23FCCC" box="[1099,1174,860,879]" italics="true" pageId="18" pageNumber="419">Medusapyga</emphasis>
<taxonomicName id="1EFAF28DFFFF4767FD47B954FD53FC2B" box="[697,742,885,904]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="alsea">alsea</taxonomicName>
</emphasis>
(star) in
<collectingRegion id="1B3E47ECFFFF4767FCCCB954FCC7FC2B" box="[818,882,885,904]" country="United States of America" name="Oregon" pageId="18" pageNumber="419">Oregon</collectingRegion>
and
<taxonomicName id="1EFAF28DFFFF4767FC64B954FBE6FC2B" box="[922,1107,885,904]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFFF4767FC64B954FBE6FC2B" box="[922,1107,885,904]" italics="true" pageId="18" pageNumber="419">Medusapyga chehalis</emphasis>
</taxonomicName>
(circlein
<collectingRegion id="1B3E47ECFFFF4767FD5FB9AEFCBCFC01" box="[673,777,911,930]" country="United States of America" name="Washington" pageId="18" pageNumber="419">Washington</collectingRegion>
.
</paragraph>
</caption>
<paragraph id="D945890EFFFF4767FF55B9DAFB23FB90" blockId="18.[123,1174,923,1715]" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FF55B9DAFEFCFBB0" box="[171,329,1019,1043]" italics="true" pageId="18" pageNumber="419">Male genitalia</emphasis>
(
<figureCitation id="41C1958BFFFF4767FEA7B9DAFE68FBB0" box="[345,477,1019,1043]" captionStart="FIGURE" captionStartId="14.[155,167,910,929]" captionTargetBox="[120,624,216,894]" captionTargetId="figure-377@14.[120,645,216,902]" captionTargetPageId="14" captionText="FIGURE 11. Male aedeagus, left lateral view. (A) Medusapyga alsea, voucher JRL002. (B) M. alsea, voucher JRL013. (C) M. chehalis, voucher JRL012. (D) M. chehalis, voucher JRL010. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12613593" httpUri="https://zenodo.org/record/12613593/files/figure.png" pageId="18" pageNumber="419">Fig. 11C, D</figureCitation>
). Aedeagus with ventral margin somewhat sinuate, less arcuate than
<taxonomicName id="1EFAF28DFFFF4767FF51BE3DFEBFFB90" box="[175,266,1051,1075]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFFF4767FF51BE3DFEBFFB90" box="[175,266,1051,1075]" italics="true" pageId="18" pageNumber="419">M. alsea</emphasis>
</taxonomicName>
, with subapical downward bulge. Internal sac with small dark structure near dorsum.
</paragraph>
<paragraph id="D945890EFFFF4767FF55BE1DFCFAFA90" blockId="18.[123,1174,923,1715]" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FF55BE1DFE3DFBF0" bold="true" box="[171,392,1084,1107]" pageId="18" pageNumber="419">Sexual dimorphism</emphasis>
. In addition to the differences in protarsomeres and profemora, and lack of a medial preapical keel on visible abdominal ventrite 2, females are larger than males. Female SBL ranged from
<quantity id="1E0224EBFFFF4767FEC6BE5AFE67FB31" box="[312,466,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" metricValueMax="2.35" metricValueMin="2.13" pageId="18" pageNumber="419" unit="mm" value="2.24" valueMax="2.35" valueMin="2.13">2.132.35 mm</quantity>
and averaged
<quantity id="1E0224EBFFFF4767FD94BE5AFD79FB31" box="[618,716,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="18" pageNumber="419" unit="mm" value="2.25">2.25 mm</quantity>
. Male SBL ranged from
<quantity id="1E0224EBFFFF4767FC31BE5AFBDCFB31" box="[975,1129,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="2.105" metricValueMax="2.18" metricValueMin="2.03" pageId="18" pageNumber="419" unit="mm" value="2.105" valueMax="2.18" valueMin="2.03">2.032.18 mm</quantity>
and averaged
<quantity id="1E0224EBFFFF4767FF1FBEBAFEF1FB11" box="[225,324,1179,1203]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="419" unit="mm" value="2.1">2.10 mm</quantity>
. A two-tailed Students T-test indicated these differences in average length were highly significant (p = 0.0000003). The abdominal fovea on visible abdominal ventrite 2 is much longer and broader in females, about 1/3 the length of ventrite 2 from the metacoxae to the ventrites posterior margin. It is 2.4 times as long as the male fovea. The female fovea is also much more deeply impressed. It is almost circular, about as wide as long.
</paragraph>
<paragraph id="D945890EFFFF4767FF55BF1DFB23FA10" blockId="18.[123,1174,923,1715]" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FF55BF1DFEA8FAF0" bold="true" box="[171,285,1340,1363]" pageId="18" pageNumber="419">Variation.</emphasis>
The posterolateral pronotal margins immediately anterior of the posterior margin of some specimens were slightly obliquely divergent, rather than parallel. Although most specimens were pale testaceous, a few specimens were very dark brown, almost black. A few of the paler individuals may have been teneral; this appears certain for a male with a clearly teneral aedeagus.
</paragraph>
<paragraph id="D945890EFFFF4767FF55BF9DFEF4F9D1" blockId="18.[123,1174,923,1715]" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FF55BF9DFEF9FA70" bold="true" box="[171,332,1468,1491]" pageId="18" pageNumber="419">Identification.</emphasis>
Characters of
<taxonomicName id="1EFAF28DFFFF4767FE18BF9DFDD7FA70" box="[486,610,1467,1491]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFFF4767FE18BF9DFDD7FA70" box="[486,610,1467,1491]" italics="true" pageId="18" pageNumber="419">M. chehalis</emphasis>
</taxonomicName>
distinguishing it from
<taxonomicName id="1EFAF28DFFFF4767FCAABF9DFC1AFA70" box="[852,943,1467,1491]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFFF4767FCAABF9DFC1AFA70" box="[852,943,1467,1491]" italics="true" pageId="18" pageNumber="419">M. alsea</emphasis>
</taxonomicName>
include smaller average length, depressed and less robust habitus, absence of a frontoclypeal prominence, pronotum without protruding anterior angles and with obtuse and rounded posterior angles, females with angulate to faintly denticulate profemoral bases, mesotrochanters with evenly rounded posterior margins, punctate abdominal foveae, and with a less arcuate aedeagus lacking a C-shaped structure in the internal sac.
</paragraph>
<paragraph id="D945890EFFFF4766FF55BC5DFDCCFED7" blockId="18.[123,1174,923,1715]" lastBlockId="19.[123,1174,219,372]" lastPageId="19" lastPageNumber="420" pageId="18" pageNumber="419">
<emphasis id="EB8E551CFFFF4767FF55BC5DFE75F930" bold="true" box="[171,448,1660,1683]" pageId="18" pageNumber="419">Geographic distribution</emphasis>
(
<figureCitation id="41C1958BFFFF4767FE2EBC5AFD94F930" box="[464,545,1659,1683]" captionStart="FIGURE" captionStartId="18.[683,695,860,879]" captionTargetBox="[706,1176,216,848]" captionTargetId="figure-541@18.[648,1176,216,848]" captionText="FIGURE 15. Known geographic distributions of Medusa- pyga alsea (star) in Oregon and Medusapyga chehalis (circle) in Washington." figureDoi="http://doi.org/10.5281/zenodo.12612700" httpUri="https://zenodo.org/record/12612700/files/figure.png" pageId="18" pageNumber="419">Fig. 15</figureCitation>
).
<taxonomicName id="1EFAF28DFFFF4767FDC9BC5DFD03F930" box="[567,694,1659,1683]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFFF4767FDC9BC5DFD03F930" box="[567,694,1659,1683]" italics="true" pageId="18" pageNumber="419">M. chehalis</emphasis>
</taxonomicName>
is known only from four closely proximate sites:
<location id="DC25DFD5FFFF4767FF44BCBAFE83F910" LSID="urn:lsid:plazi:treatment:51533818FFFC4766FF85BAFAFDCCFED7:DC25DFD5FFFF4767FF44BCBAFE83F910" box="[186,310,1691,1715]" latitude="46.962" longLatPrecision="6" longitude="-123.1084" name="Falls Creek" pageId="18" pageNumber="419" stateProvince="Washington">Falls Creek</location>
Campground at
<geoCoordinate id="BCCEEFC9FFFF4767FE13BCBAFDD4F910" box="[493,609,1691,1715]" degrees="46.9333" direction="north" orientation="latitude" pageId="18" pageNumber="419" precision="5" value="46.9333">46.9333°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFF4767FD94BCBAFD44F910" box="[618,753,1691,1715]" degrees="123.1333" direction="west" orientation="longitude" pageId="18" pageNumber="419" precision="5" value="-123.1333">123.1333°W</geoCoordinate>
and
<quantity id="1E0224EBFFFF4767FCD6BCBAFCDBF911" box="[808,878,1691,1715]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="419" unit="m" value="200.0">
<elevation id="52D76E3DFFFF4767FCD6BCBAFCDBF911" box="[808,878,1691,1715]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="419" unit="m" value="200.0">200 m</elevation>
</quantity>
elevation, the type locality approximately 2.0 km to the north at
<geoCoordinate id="BCCEEFC9FFFE4766FDEEBAFAFD30FF50" box="[528,645,219,243]" degrees="46.9600" direction="north" orientation="latitude" pageId="19" pageNumber="420" precision="5" value="46.96">46.9600°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFE4766FD73BAFAFCA1FF50" box="[653,788,219,243]" degrees="123.1350" direction="west" orientation="longitude" pageId="19" pageNumber="420" precision="5" value="-123.135">123.1350°W</geoCoordinate>
and between
<quantity id="1E0224EBFFFE4766FC55BAFAFB90FF51" box="[939,1061,219,243]" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" metricValueMax="4.12" metricValueMin="3.66" pageId="19" pageNumber="420" unit="m" value="389.0" valueMax="412.0" valueMin="366.0">
<elevation id="52D76E3DFFFE4766FC55BAFAFB90FF51" box="[939,1061,219,243]" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" metricValueMax="4.12" metricValueMin="3.66" pageId="19" pageNumber="420" unit="m" value="389.0" valueMax="412.0" valueMin="366.0">366412 m</elevation>
</quantity>
elevation, a site approximately
<quantity id="1E0224EBFFFE4766FEA5BADAFE10FEB0" box="[347,421,251,275]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="420" unit="km" value="0.4">0.4 km</quantity>
SSW of the
<typeStatus id="064137ACFFFE4766FDD1BADDFDE8FEB7" box="[559,605,252,276]" pageId="19" pageNumber="420">type</typeStatus>
site at
<geoCoordinate id="BCCEEFC9FFFE4766FD54BADAFCAAFEB0" box="[682,799,251,275]" degrees="46.9564" direction="north" orientation="latitude" pageId="19" pageNumber="420" precision="5" value="46.9564">46.9564°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFE4766FCD8BADAFC18FEB0" box="[806,941,251,275]" degrees="123.1366" direction="west" orientation="longitude" pageId="19" pageNumber="420" precision="5" value="-123.1366">123.1366°W</geoCoordinate>
and
<quantity id="1E0224EBFFFE4766FC1CBADAFB93FEB0" box="[994,1062,251,275]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="19" pageNumber="420" unit="m" value="315.0">
<elevation id="52D76E3DFFFE4766FC1CBADAFB93FEB0" box="[994,1062,251,275]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="19" pageNumber="420" unit="m" value="315.0">315 m</elevation>
</quantity>
elevation, and a site approximately
<quantity id="1E0224EBFFFE4766FE77BB3AFE61FE90" box="[393,468,283,307]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="420" unit="km" value="2.6">2.6 km</quantity>
east of the
<typeStatus id="064137ACFFFE4766FDAFBB3DFDCAFE97" box="[593,639,284,308]" pageId="19" pageNumber="420">type</typeStatus>
locality at
<geoCoordinate id="BCCEEFC9FFFE4766FD06BB3AFCD8FE90" box="[760,877,283,307]" degrees="46.9620" direction="north" orientation="latitude" pageId="19" pageNumber="420" precision="5" value="46.962">46.9620°N</geoCoordinate>
<geoCoordinate id="BCCEEFC9FFFE4766FC8ABB3AFC49FE90" box="[884,1020,283,307]" degrees="123.1084" direction="west" orientation="longitude" pageId="19" pageNumber="420" precision="5" value="-123.1084">123.1084°W</geoCoordinate>
, at
<quantity id="1E0224EBFFFE4766FBDCBB3AFBD3FE90" box="[1058,1126,283,307]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="19" pageNumber="420" unit="m" value="301.0">
<elevation id="52D76E3DFFFE4766FBDCBB3AFBD3FE90" box="[1058,1126,283,307]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="19" pageNumber="420" unit="m" value="301.0">301 m</elevation>
</quantity>
elevation, all in southwestern
<collectingRegion id="1B3E47ECFFFE4766FE66BB1AFDAFFEF0" box="[408,538,315,339]" country="United States of America" name="Washington" pageId="19" pageNumber="420">Washington</collectingRegion>
. None of these sites are farther than
<quantity id="1E0224EBFFFE4766FC65BB1AFC51FEF0" box="[923,996,315,339]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="420" unit="km" value="2.6">2.6 km</quantity>
apart and bound a roughly triangular area of about 270 hectares.
</paragraph>
</subSubSection>
</treatment>
</document>