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<document id="E5BE67EE5837F7488E2F1F198AA5CB03" ID-CLB-Dataset="299337" ID-DOI="10.5281/zenodo.11067222" ID-GBIF-Dataset="1cb742f7-205f-4cc5-9136-b613d5ab059d" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067222" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052073787" checkinUser="felipe" docAuthor="LaBonte, James R. &amp; Maddison, David R." docDate="2023" docId="51533818FFE54778FEA4BF2EFF02FAF1" docLanguage="en" docName="ProcCalifAcadSci.67.18.401-432.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (18)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Medusapyga LaBonte and Maddison 2023" docType="treatment" docVersion="7" lastPageNumber="414" masterDocId="AD6A4060FFED4775FFFEBA21FFB5FFA3" masterDocTitle="Medusapyga LaBonte and Maddison, a New Genus of Anillini (Coleoptera: Carabidae: Trechinae) from the Pacific Northwest of the United States" masterLastPageNumber="432" masterPageNumber="401" pageNumber="409" updateTime="1719875109990" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="646AAA0B555F5B23BFBC6EC142D8D5F5">Medusapyga LaBonte and Maddison, a New Genus of Anillini (Coleoptera: Carabidae: Trechinae) from the Pacific Northwest of the United States</mods:title>
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<mods:namePart id="955ED856E83C489564CD5F344D67A391">LaBonte, James R.</mods:namePart>
<mods:affiliation id="E6C08AA965B8D6614840F205F3FBF008">1570 Kenard Street NW, Salem, OR 97304, USA.</mods:affiliation>
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<mods:namePart id="03A7FCAED16516D52CAFD922D60438F0">Maddison, David R.</mods:namePart>
<mods:affiliation id="5F929000E91E412DA8888EE2669C9587">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA.</mods:affiliation>
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<mods:title id="C00B105256A99087010904403D3C1A0D">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="C7786E6CD6B521279888F1A56436E8C3">2023</mods:date>
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<treatment id="51533818FFE54778FEA4BF2EFF02FAF1" ID-DOI="http://doi.org/10.5281/zenodo.12613595" ID-GBIF-Taxon="234396642" ID-Zenodo-Dep="12613595" LSID="urn:lsid:plazi:treatment:51533818FFE54778FEA4BF2EFF02FAF1" httpUri="http://treatment.plazi.org/id/51533818FFE54778FEA4BF2EFF02FAF1" lastPageId="13" lastPageNumber="414" pageId="8" pageNumber="409">
<subSubSection id="91E0DA85FFE5477DFEA4BF2EFC03FA8A" box="[346,950,1295,1321]" pageId="8" pageNumber="409" type="nomenclature">
<paragraph id="D945890EFFE5477DFEA4BF2EFC03FA8A" blockId="8.[346,950,1295,1321]" box="[346,950,1295,1321]" pageId="8" pageNumber="409">
<heading id="820D3E62FFE5477DFEA4BF2EFC03FA8A" box="[346,950,1295,1321]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="409" reason="4">
<taxonomicName id="1EFAF28DFFE5477DFEA4BF2EFC03FA8A" authority="LaBonte and Maddison" authorityName="LaBonte and Maddison" authorityYear="2023" box="[346,950,1295,1321]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EB8E551CFFE5477DFEA4BF2EFC03FA8A" bold="true" box="[346,950,1295,1321]" pageId="8" pageNumber="409">
<emphasis id="EB8E551CFFE5477DFEA4BF2EFE46FA8A" bold="true" box="[346,499,1295,1321]" italics="true" pageId="8" pageNumber="409">Medusapyga</emphasis>
LaBonte and Maddison, new genus
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91E0DA85FFE5477DFF55BF65FC08FAF8" box="[171,957,1347,1371]" pageId="8" pageNumber="409" type="type_taxon">
<paragraph id="D945890EFFE5477DFF55BF65FC08FAF8" blockId="8.[123,1174,1347,1725]" box="[171,957,1347,1371]" pageId="8" pageNumber="409">
<emphasis id="EB8E551CFFE5477DFF55BF65FE8AFAF8" bold="true" box="[171,319,1348,1371]" pageId="8" pageNumber="409">
<typeStatus id="064137ACFFE5477DFF55BF65FF56FAF8" box="[171,227,1348,1371]" pageId="8" pageNumber="409">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="1EFAF28DFFE5477DFEB8BF62FCB6FAF8" authority="LaBonte" authorityName="LaBonte" authorityYear="2023" box="[326,771,1347,1371]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="alsea" status="sp. nov.">
<emphasis id="EB8E551CFFE5477DFEB8BF62FDBBFAF8" box="[326,526,1347,1371]" italics="true" pageId="8" pageNumber="409">Medusapyga alsea</emphasis>
LaBonte, new species
</taxonomicName>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="91E0DA85FFE5477DFF55BF45FC33F9BF" pageId="8" pageNumber="409" type="etymology">
<paragraph id="D945890EFFE5477DFF55BF45FBD0FA5F" blockId="8.[123,1174,1347,1725]" pageId="8" pageNumber="409">
<emphasis id="EB8E551CFFE5477DFF55BF45FE9EFAD8" bold="true" box="[171,299,1380,1403]" pageId="8" pageNumber="409">Etymology.</emphasis>
The generic name refers to the bifurcate setae on laterotergite IX of females. The Latin
<taxonomicName id="1EFAF28DFFE5477DFF44BFA5FEBBFA3F" authorityName="Linnaeus" authorityYear="1758" box="[186,270,1412,1436]" class="Magnoliopsida" family="Violaceae" genus="Medusa" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="409" phylum="Tracheophyta" rank="genus">
<emphasis id="EB8E551CFFE5477DFF44BFA5FEBBFA3F" box="[186,270,1412,1436]" italics="true" pageId="8" pageNumber="409">Medusa</emphasis>
</taxonomicName>
alludes to the mythical monster (a gorgon) with snakes rather than hair on her head and the similarity of the beetles’ setae to the forked tongues of snakes, while
<emphasis id="EB8E551CFFE5477DFC48BF84FC5FFA1F" box="[950,1002,1445,1468]" italics="true" pageId="8" pageNumber="409">pyga</emphasis>
is derived from the Greek noun
<emphasis id="EB8E551CFFE5477DFED7BFE4FEEEFA7F" box="[297,347,1477,1500]" italics="true" pageId="8" pageNumber="409">pyge</emphasis>
for rump, referring to the placement of the setae and that these can often be observed protruding from the posterior of the abdomen. The gender of this name is feminine.
</paragraph>
<paragraph id="D945890EFFE5477DFF55BC24FC33F9BF" blockId="8.[123,1174,1347,1725]" box="[171,902,1540,1565]" pageId="8" pageNumber="409">
<emphasis id="EB8E551CFFE5477DFF55BC24FEF6F9BF" bold="true" box="[171,323,1541,1564]" pageId="8" pageNumber="409">Composition.</emphasis>
This genus includes the two species described below.
</paragraph>
</subSubSection>
<subSubSection id="91E0DA85FFE54778FF55BC04FF02FAF1" lastPageId="13" lastPageNumber="414" pageId="8" pageNumber="409" type="description">
<paragraph id="D945890EFFE5477CFF55BC04FCA2FDB0" blockId="8.[123,1174,1347,1725]" lastBlockId="9.[123,813,219,1011]" lastPageId="9" lastPageNumber="410" pageId="8" pageNumber="409">
<emphasis id="EB8E551CFFE5477DFF55BC04FE8FF99F" bold="true" box="[171,314,1573,1596]" pageId="8" pageNumber="409">Recognition.</emphasis>
Diagnostic characters of species of
<taxonomicName id="1EFAF28DFFE5477DFD41BC05FCF2F99F" box="[703,839,1572,1596]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="EB8E551CFFE5477DFD41BC05FCF2F99F" box="[703,839,1572,1596]" italics="true" pageId="8" pageNumber="409">Medusapyga</emphasis>
</taxonomicName>
include the field of long bifurcate setae on the ventral surface of the apical half of laterotergite IX of the female genitalia, the ventrobasal spine of the male profemur, and the two asymmetrically strongly dilated basal tarsomeres of the male protarsus, the first tarsomere with two rows and the second with one row of adhesive articulo-setae (
<bibRefCitation id="BD6BF4FFFFE5477DFE82BC84FE40F91E" author="STORK, N. E." box="[380,501,1701,1725]" pageId="8" pageNumber="409" pagination="173 - 306" refId="ref19071" refString="STORK, N. E. 1980. A scanning electron microscopy study of tarsal adhesive setae in the Coleoptera. Zoological Journal of the Linnean Society 68: 173 - 306." type="journal article" year="1980">Stork 1980</bibRefCitation>
) on the venter. Each species has one or more additional unique characters serving to distinguish them from members of previously described genera of North American
<taxonomicName id="1EFAF28DFFE4477CFD6DBADAFD5BFEB0" box="[659,750,251,275]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
. The mesotrochanters of both genders of
<taxonomicName id="1EFAF28DFFE4477CFE06BB3DFDE1FE90" box="[504,596,283,307]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE4477CFE06BB3DFDE1FE90" box="[504,596,283,307]" italics="true" pageId="9" pageNumber="410">M. alsea</emphasis>
</taxonomicName>
are quadrate, with a large, blunt, triangular tooth along the posterior margin. The profemora of female
<taxonomicName id="1EFAF28DFFE4477CFEBEBB7DFE0AFED0" box="[320,447,347,371]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE4477CFEBEBB7DFE0AFED0" box="[320,447,347,371]" italics="true" pageId="9" pageNumber="410">M. chehalis</emphasis>
</taxonomicName>
are distinctly angulate to dentate on the ventrobasal face and males of
<taxonomicName id="1EFAF28DFFE4477CFDEEBB5DFD38FE30" box="[528,653,379,403]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE4477CFDEEBB5DFD38FE30" box="[528,653,379,403]" italics="true" pageId="9" pageNumber="410">M. chehalis</emphasis>
</taxonomicName>
have a medial laterally compressed and posteriorly declivous keel on ventrite 2 (these species-specific characters will be addressed in more detail under the species descriptions below). Both species are relatively large for North American
<taxonomicName id="1EFAF28DFFE4477CFE1EBBDAFD83FDB0" box="[480,566,507,531]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="403" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
, especially
<taxonomicName id="1EFAF28DFFE4477CFD4DBBDDFCBAFDB0" box="[691,783,507,531]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE4477CFD4DBBDDFCBAFDB0" box="[691,783,507,531]" italics="true" pageId="9" pageNumber="410">M. alsea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D945890EFFE4477CFF55B83DFE63FD50" blockId="9.[123,813,219,1011]" pageId="9" pageNumber="410">
<emphasis id="EB8E551CFFE4477CFF55B83DFE83FD90" bold="true" box="[171,310,540,563]" pageId="9" pageNumber="410">Description.</emphasis>
<emphasis id="EB8E551CFFE4477CFEC2B83AFE25FD90" box="[316,400,539,563]" italics="true" pageId="9" pageNumber="410">Habitus</emphasis>
(
<figureCitation id="41C1958BFFE4477CFE61B83AFE57FD90" box="[415,482,539,563]" captionStart="FIGURE" captionStartId="1.[155,167,1465,1484]" captionTargetBox="[197,1115,216,1451]" captionTargetId="figure-74@1.[197,1115,216,1451]" captionTargetPageId="1" captionText="FIGURE 1. (A) Female of Medusapyga alsea from the type locality. (B) Female of M. chehalis from USA, Washington, Thurston Co., 11 km NW of Littlerock, approximately 0.4 km SSW of the type locality, 46.9564°N 123.1366°W, 315 m elevation. Both specimens were freshly killed, and then posed and photographed on soil from their habitats. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067224" httpUri="https://zenodo.org/record/11067224/files/figure.png" pageId="9" pageNumber="410">Figs 1</figureCitation>
,
<figureCitation id="41C1958BFFE4477CFE0EB83AFDB5FD90" box="[496,512,539,563]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="9" pageNumber="410">2</figureCitation>
). SBL
<quantity id="1E0224EBFFE4477CFDB5B83AFD5CFD90" box="[587,745,539,563]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" metricValueMax="2.65" metricValueMin="2.03" pageId="9" pageNumber="410" unit="mm" value="2.34" valueMax="2.65" valueMin="2.03">2.03–2.65 mm</quantity>
. Head large and broad, HW/HL ~0.88, HW/PWm ~0.75–0.81. Pronotum large, broad, PWm/EW ~0.69, with lateral margins sinuate anterior to posterior angles. Elytra convex to distinctly flattened in lateral perspective and ovoid in dorsal perspective, EW/EL ~0.65, EW/SBL ~0.36–0.39. Translucent pale yellowish brown to reddish brown to dark brown.
</paragraph>
<paragraph id="D945890EFFE4477CFF55B8DAFD80FC51" blockId="9.[123,813,219,1011]" pageId="9" pageNumber="410">Dorsal microsculpture imbricate, with the posterior margin of each sculpticell slightly raised. Sculpticell definition distinct but varies between species, among parts of the body, and among portions of those body parts. Sculpticell shape and size similarly variable, with the shape ranging from irregularly isodiametric to elongate irregularly polygonal. Body surface shiny. Dorsum with sparse vestiture of short, fine setae about 0.12–0.25 length of long pronotal and elytral interstrial setae.</paragraph>
<caption id="8D85D986FFE4477CFC94B9B2FB85FC51" ID-DOI="http://doi.org/10.5281/zenodo.12612687" ID-Zenodo-Dep="12612687" httpUri="https://zenodo.org/record/12612687/files/figure.png" pageId="9" pageNumber="410" startId="9.[874,886,915,934]" targetBox="[865,1147,221,892]" targetPageId="9">
<paragraph id="D945890EFFE4477CFC94B9B2FB85FC51" pageId="9" pageNumber="410">
<superScript id="2E8F2446FFE4477CFC94B9B2FB21FC06" attach="right" box="[874,1172,914,934]" fontSize="8" pageId="9" pageNumber="410">
<smallCapsWord id="DFA31FD2FFE4477CFC94B9B2FC12FC06" baselines="929,929" box="[874,935,915,934]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="9" pageNumber="410">FIGURE</smallCapsWord>
3. Dorsal habitus of female
</superScript>
<taxonomicName id="1EFAF28DFFE4477CFCB7B98DFC4FFC1C" box="[841,1018,940,959]" class="Insecta" family="Carabidae" genus="Anillaspis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="explanata">
<emphasis id="EB8E551CFFE4477CFCB7B98DFC4FFC1C" box="[841,1018,940,959]" italics="true" pageId="9" pageNumber="410">Anillaspis explanata</emphasis>
</taxonomicName>
from
<collectingCountry id="A1EDC99EFFE4477CFBCFB98DFBEAFC1C" box="[1073,1119,940,959]" name="United States of America" pageId="9" pageNumber="410">USA</collectingCountry>
:
<subScript id="457E8B4BFFE4477CFCB7B9E4FB21FC7B" attach="right" box="[841,1172,965,984]" fontSize="8" pageId="9" pageNumber="410">California: El Dorado Co., Alabaster Cave</subScript>
(
<collectionCode id="BFEB11CBFFE4477CFCB1B9FEFC21FC51" box="[847,916,991,1010]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="9" pageNumber="410" type="Museum">CMNH</collectionCode>
). Scale bar
<quantity id="1E0224EBFFE4477CFC06B9FEFB99FC51" box="[1016,1068,991,1010]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="410" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="D945890EFFE4477FFF55B9DAFDC3FC91" blockId="9.[123,1174,1019,1107]" lastBlockId="10.[123,1173,763,819]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
<emphasis id="EB8E551CFFE4477CFF55B9DAFF50FBB0" box="[171,229,1019,1043]" italics="true" pageId="9" pageNumber="410">Head</emphasis>
(
<figureCitation id="41C1958BFFE4477CFF0BB9DAFE8CFBB0" box="[245,313,1019,1043]" captionStart="FIGURE" captionStartId="11.[155,167,793,812]" captionTargetBox="[182,1123,229,767]" captionTargetId="figure-340@11.[172,1132,216,777]" captionTargetPageId="11" captionText="FIGURE 7. Dorsal views of heads. (A) Medusapyga alsea. (B) M. chehalis. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067232" httpUri="https://zenodo.org/record/11067232/files/figure.png" pageId="9" pageNumber="410">Fig. 7</figureCitation>
). Frontal furrows broadly impressed, parallel, ending well anterior of occiput. Frontolateral carinae distinct and sharply defined, extending from a shelf over the antennal insertions posteriorly to the level of posterior supraorbital setae. A triangular protrusion extends anteriorly between the antennal carinal shelf and the clypeal seta on each side, protruding above the dorsal mandibular articulation and terminating in a small capitation. Using the terms of
<bibRefCitation id="BD6BF4FFFFE7477FFEABBB1AFDB7FEF0" author="SOKOLOV, I. M." box="[341,514,315,339]" pageId="10" pageNumber="411" pagination="1 - 37" refId="ref18438" refString="SOKOLOV, I. M. 2016. A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution. Zootaxa 4196 (1): 1 - 37. &lt;doi. org / 10.11646 / zootaxa. 4196.1.1&gt;" type="journal article" year="2016">Sokolov (2016)</bibRefCitation>
, there are a pair of clypeal setae, a pair of frontal setae, a pair of mediofrontal setae (sometmes represented by only a single seta), two pair of supraorbital setae (with the posterior pair set somewhat mediad of the anterior pair), and a pair of postorbital setae. Eyes absent. Genae with sparse setae of moderate length. Tempora shallowly convex and rounded to obtusely angulate. Occiput set off from the vertex by a shallow transverse depression just posterior of the posterior supraorbital setae. Mentum and submentum separated by distinct suture. Mentum with large, sharply pointed anteromedial tooth, with a pair of long setae near the apex of the medial tooth and lateral and paramedial pairs anterior of the posterior margin. Submentum trapezoidal, with a pair each of long setae anterolaterally near the anterior angles, paramedially, and medially (the shortest), and also with a row of short setae just posterior of the anterior margin. Gula broad, sides slightly convergent posteriorly, sparsely setose.
</paragraph>
<caption id="8D85D986FFE4477CFF63BC5BFE4EF904" ID-DOI="http://doi.org/10.5281/zenodo.11067228" ID-Zenodo-Dep="11067228" httpUri="https://zenodo.org/record/11067228/files/figure.png" pageId="9" pageNumber="410" startId="9.[157,169,1658,1677]" targetBox="[129,1168,1135,1647]" targetPageId="9" targetType="figure">
<paragraph id="D945890EFFE4477CFF63BC5BFB35F92E" blockId="9.[124,1152,1658,1703]" box="[157,1152,1658,1677]" pageId="9" pageNumber="410">
<smallCapsWord id="DFA31FD2FFE4477CFF63BC5BFF6FF92F" baselines="1672,1672" box="[157,218,1658,1677]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="9" pageNumber="410">FIGURE</smallCapsWord>
4. Left profemora, anterior view. (A)
<taxonomicName id="1EFAF28DFFE4477CFDDCBC5BFD77F92E" box="[546,706,1658,1677]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE4477CFDDCBC5BFD77F92E" box="[546,706,1658,1677]" italics="true" pageId="9" pageNumber="410">Medusapyga alsea</emphasis>
</taxonomicName>
, male. (B)
<taxonomicName id="1EFAF28DFFE4477CFCDABC5AFCDBF92E" box="[804,878,1658,1677]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE4477CFCDABC5AFCDBF92E" box="[804,878,1658,1677]" italics="true" pageId="9" pageNumber="410">M. alsea</emphasis>
</taxonomicName>
, female. (C)
<taxonomicName id="1EFAF28DFFE4477CFC1FBC5AFBF0F92E" box="[993,1093,1658,1677]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE4477CFC1FBC5AFBF0F92E" box="[993,1093,1658,1677]" italics="true" pageId="9" pageNumber="410">M. chehalis</emphasis>
</taxonomicName>
, male.
</paragraph>
<paragraph id="D945890EFFE4477CFF82BCB2FE4EF904" blockId="9.[124,1152,1658,1703]" box="[124,507,1683,1703]" pageId="9" pageNumber="410">
(D)
<taxonomicName id="1EFAF28DFFE4477CFF5EBCB5FEB1F905" box="[160,260,1683,1702]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE4477CFF5EBCB5FEB1F905" box="[160,260,1683,1702]" italics="true" pageId="9" pageNumber="410">M. chehalis</emphasis>
</taxonomicName>
, female. Scale bars 100 µm.
</paragraph>
</caption>
<caption id="8D85D986FFE7477FFC91B812FC5EFD43" ID-DOI="http://doi.org/10.5281/zenodo.12612690" ID-Zenodo-Dep="12612690" httpUri="https://zenodo.org/record/12612690/files/figure.png" pageId="10" pageNumber="411" startId="10.[879,935,563,582]" targetBox="[849,1170,220,542]" targetPageId="10">
<paragraph id="D945890EFFE7477FFC91B812FB22FD65" pageId="10" pageNumber="411">
<superScript id="2E8F2446FFE7477FFC91B812FB23FDE5" attach="right" box="[879,1174,563,582]" fontSize="8" pageId="10" pageNumber="411">Figure 5 (above). Male right protarsi</superScript>
, dorsal view. (A)
<taxonomicName id="1EFAF28DFFE7477FFBD6B86DFB20FDFC" box="[1064,1173,588,607]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="EB8E551CFFE7477FFBD6B86DFB20FDFC" box="[1064,1173,588,607]" italics="true" pageId="10" pageNumber="411">Medusapyga</emphasis>
</taxonomicName>
<emphasis id="EB8E551CFFE7477FFCB1B847FC37FDDA" box="[847,898,614,633]" italics="true" pageId="10" pageNumber="411">
<taxonomicName id="1EFAF28DFFE7477FFCB1B847FCCAFDDA" box="[847,895,614,633]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="alsea">alsea</taxonomicName>
.
</emphasis>
(B)
<taxonomicName id="1EFAF28DFFE7477FFC4CB846FBADFDDA" box="[946,1048,614,633]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE7477FFC4CB846FBADFDDA" box="[946,1048,614,633]" italics="true" pageId="10" pageNumber="411">M. chehalis</emphasis>
</taxonomicName>
. The view of the
<specimenCount id="CFFC4287FFE7477FFC89B8A1FBB7FD31" box="[887,1026,639,659]" count="2" pageId="10" pageNumber="411" type="generic">two specimens</specimenCount>
is not exactly equivalent, which accounts for some differences between the two images.
</paragraph>
<paragraph id="D945890EFFE7477FFCB1B8EDFC5EFD43" box="[847,1003,716,736]" pageId="10" pageNumber="411">Scale bars 50 µm.</paragraph>
</caption>
<paragraph id="D945890EFFE7477EFF55B91AFE6DFC67" blockId="10.[123,1174,827,1139]" lastBlockId="11.[123,1174,844,1700]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
Mandibles matching basic structure of
<taxonomicName id="1EFAF28DFFE7477FFD9EB91AFC2EFCF0" authority="(Maddison 1993)" baseAuthorityName="Maddison" baseAuthorityYear="1993" box="[608,923,827,851]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="EB8E551CFFE7477FFD9EB91AFD61FCF0" box="[608,724,827,851]" italics="true" pageId="10" pageNumber="411">Bembidion</emphasis>
(
<bibRefCitation id="BD6BF4FFFFE7477FFD1BB91AFC21FCF0" author="MADDISON, D. R." box="[741,916,827,851]" pageId="10" pageNumber="411" pagination="143 - 299" refId="ref16959" refString="MADDISON, D. R. 1993. Systematics of the Holarctic beetle subgenus Bracteon and related Bembidion (Coleoptera: Carabidae). Bulletin of the Museum of Comparative Zoology 153: 143 - 299." type="journal article" year="1993">Maddison 1993</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="1EFAF28DFFE7477FFC28B91AFB9CFCF0" box="[982,1065,827,851]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="403" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
(Sokolov 2013). Stout, with sharp, slightly ventrally curved apices extending well beyond labrum. Each mandible of similar size and shape, except for details of mandibular teeth. Left mandible has terebral (rounded), premolar, and molar teeth. Right mandible has anterior retinacular, terebral, posterior retinacular, premolar, and molar teeth. Labrum rectangular, anterior margin distinctly arcuately emarginate, three pairs of setae (sequentially decreasing in length mediad) along anterior margin. Clypeus trapezoidal, clypeolabral suture distinct, frontoclypeal suture present; with a pair of long posterolateral fixed setae. Subterminal maxillary palpomeres sparsely setose; inner lateral margin of penultimate palpomere strongly arcuate, outer lateral margin straight; terminal palpomere subulate, about one-third as long as penultimate palpomere. Subterminal labial palpomeres sparsely setose; inner lateral margin of penultimate palpomere straight, outer lateral margin strongly arcuate; terminal palpomere subulate, about half as long as penultimate palpomere. Glossa with two apical setae. Distinct, lobate paraglossae extending beyond apex of glossa, each densely setose in the apical half.
</paragraph>
<caption id="8D85D986FFE7477FFF65BCA3FB22F90C" ID-DOI="http://doi.org/10.5281/zenodo.11067230" ID-Zenodo-Dep="11067230" httpUri="https://zenodo.org/record/11067230/files/figure.png" pageId="10" pageNumber="411" startId="10.[155,211,1666,1685]" targetBox="[143,1176,1168,1651]" targetPageId="10" targetType="figure">
<paragraph id="D945890EFFE7477FFF65BCA3FB22F90C" blockId="10.[123,1175,1666,1711]" pageId="10" pageNumber="411">
Figure 6. Female genitalia of
<taxonomicName id="1EFAF28DFFE7477FFE51BCA3FDE6F936" box="[431,595,1666,1685]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE7477FFE51BCA3FDE6F936" box="[431,595,1666,1685]" italics="true" pageId="10" pageNumber="411">Medusapyga alsea</emphasis>
</taxonomicName>
from the type locality, ventral view. (A) Overview of voucher V101505. Scale bar 100 µm. (B) Closeup of voucher V101504, showing bifurcate setae on laterotergites. Scale bar 50 µm.
</paragraph>
</caption>
<caption id="8D85D986FFE6477EFF65B938FC71FC8F" ID-DOI="http://doi.org/10.5281/zenodo.11067232" ID-Zenodo-Dep="11067232" box="[155,964,792,812]" httpUri="https://zenodo.org/record/11067232/files/figure.png" pageId="11" pageNumber="412" startId="11.[155,167,793,812]" targetBox="[182,1123,229,767]" targetPageId="11" targetType="figure">
<paragraph id="D945890EFFE6477EFF65B938FC71FC8F" blockId="11.[155,964,792,812]" box="[155,964,792,812]" pageId="11" pageNumber="412">
<smallCapsWord id="DFA31FD2FFE6477EFF65B938FF6DFC88" baselines="807,807" box="[155,216,793,812]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="11" pageNumber="412">FIGURE</smallCapsWord>
7. Dorsal views of heads. (A)
<taxonomicName id="1EFAF28DFFE6477EFE1FB939FD34FC88" box="[481,641,792,811]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE6477EFE1FB939FD34FC88" box="[481,641,792,811]" italics="true" pageId="11" pageNumber="412">Medusapyga alsea</emphasis>
</taxonomicName>
. (B)
<taxonomicName id="1EFAF28DFFE6477EFD50B938FCA7FC88" box="[686,786,792,811]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE6477EFD50B938FCA7FC88" box="[686,786,792,811]" italics="true" pageId="11" pageNumber="412">M. chehalis</emphasis>
</taxonomicName>
. Scale bars 100 µm.
</paragraph>
</caption>
<paragraph id="D945890EFFE6477EFF55B9EDFEECFBC7" blockId="11.[123,1174,844,1700]" pageId="11" pageNumber="412">Antennae eleven-segmented, robust, antennomeres 4–10 more or less moniliform. Except for scape, all antennomeres at least partially pubescent. Scape stout, oblong, glabrous except for a few scattered setae and four long apical setae; antennomere 2 slightly pedunculate basally, otherwise ovate; antennomere 3 strongly pedunculate, very narrow basally; antennomere 11 elongate oval, narrowest anteriorly.</paragraph>
<paragraph id="D945890EFFE6477EFF55BE4CFF11F980" blockId="11.[123,1174,844,1700]" pageId="11" pageNumber="412">
<emphasis id="EB8E551CFFE6477EFF55BE4CFEA0FB27" box="[171,277,1133,1156]" italics="true" pageId="11" pageNumber="412">Pronotum</emphasis>
(
<figureCitation id="41C1958BFFE6477EFEDCBE4DFED7FB27" box="[290,354,1132,1156]" captionStart="FIGURE" captionStartId="12.[155,167,666,685]" captionTargetBox="[195,1128,229,656]" captionTargetId="figure-339@12.[181,1141,216,662]" captionTargetPageId="12" captionText="FIGURE 8. Dorsal views of pronota. (A) Medusapyga alsea. (B) M. chehalis. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067234" httpUri="https://zenodo.org/record/11067234/files/figure.png" pageId="11" pageNumber="412">Fig. 8</figureCitation>
). Outline cordate, broadest at about one-third total pronotal length from anterior margin and posterior of lateral primary setae, width at anterior slightly greater than at posterior angles. Lateral primary setae about one-quarter of total pronotal length from anterior margin, posterior pronotal setae just anterior of or well anterior of posterior angles. Dorsum with sparse, socketed, small setae. Numerous small setae regularly distributed along lateral margins. Anterior angles rounded. Lateral margins approximately parallel from posterior angles anteriorward, then convergently arcuate to anterior angles, with several small denticles anterior of posterior angles. Posterior margin slightly convex near posterior angles. Lateral explanations distinct and reflexed, widest at posterior angles and abruptly narrowing anterior of posteromedial pronotal impression. Posterolateral depressions inside hind angles deep, short, and ovate, delimiting lateral bounds of posteromedian impression. Disc convex with well defined median longitudinal impression, deepest adjoining anteromedian and posteromedian impressions. Posteromedian impression deep, extending to posterior pronotal margin, deepest at anterior delimitation, otherwise convex throughout.
</paragraph>
<paragraph id="D945890EFFE64779FF55BC0DFE8EFCBA" blockId="11.[123,1174,844,1700]" lastBlockId="12.[123,718,669,793]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
<emphasis id="EB8E551CFFE6477EFF55BC0DFEA1F9E7" box="[171,276,1580,1604]" italics="true" pageId="11" pageNumber="412">Scutellum</emphasis>
. Small, lateral and posterior margins evenly convex, apex obtusely rounded. Almost entirely within depression anterior of elytral suture. When the pronotum is extended away from the anterior margin of the elytra, as in active, live specimens (
<figureCitation id="41C1958BFFE6477EFD11BC4DFC86F927" box="[751,819,1644,1668]" captionStart="FIGURE" captionStartId="1.[155,167,1465,1484]" captionTargetBox="[197,1115,216,1451]" captionTargetId="figure-74@1.[197,1115,216,1451]" captionTargetPageId="1" captionText="FIGURE 1. (A) Female of Medusapyga alsea from the type locality. (B) Female of M. chehalis from USA, Washington, Thurston Co., 11 km NW of Littlerock, approximately 0.4 km SSW of the type locality, 46.9564°N 123.1366°W, 315 m elevation. Both specimens were freshly killed, and then posed and photographed on soil from their habitats. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067224" httpUri="https://zenodo.org/record/11067224/files/figure.png" pageId="11" pageNumber="412">Figs 1</figureCitation>
,
<figureCitation id="41C1958BFFE6477EFCBFBC4DFCD4F927" box="[833,865,1644,1668]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="11" pageNumber="412">2B</figureCitation>
) or in most preserved specimens, the scutellum is visible. In live specimens at rest or in some preserved specimens, the posterior margin of the pronotum fits over and against the anterior elytral declivity (
<figureCitation id="41C1958BFFE14779FE72B8C0FE57FD5A" box="[396,482,737,761]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="12" pageNumber="413">Fig. 2A</figureCitation>
), completely concealing the scutellum.
</paragraph>
<caption id="8D85D986FFE14779FF65B8BBFC60FD0E" ID-DOI="http://doi.org/10.5281/zenodo.11067234" ID-Zenodo-Dep="11067234" box="[155,981,665,685]" httpUri="https://zenodo.org/record/11067234/files/figure.png" pageId="12" pageNumber="413" startId="12.[155,167,666,685]" targetBox="[195,1128,229,656]" targetPageId="12" targetType="figure">
<paragraph id="D945890EFFE14779FF65B8BBFC60FD0E" blockId="12.[123,718,669,793]" box="[155,981,665,685]" lastBlockId="12.[155,981,665,685]" pageId="12" pageNumber="413">
<smallCapsWord id="DFA31FD2FFE14779FF65B8BBFF6DFD0F" baselines="680,680" box="[155,216,666,685]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="12" pageNumber="413">FIGURE</smallCapsWord>
8. Dorsal views of pronota. (A)
<taxonomicName id="1EFAF28DFFE14779FE0CB8B8FD27FD0F" box="[498,658,665,684]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE14779FE0CB8B8FD27FD0F" box="[498,658,665,684]" italics="true" pageId="12" pageNumber="413">Medusapyga alsea</emphasis>
</taxonomicName>
. (B)
<taxonomicName id="1EFAF28DFFE14779FD41B8BBFC96FD0F" box="[703,803,665,684]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="chehalis">
<emphasis id="EB8E551CFFE14779FD41B8BBFC96FD0F" box="[703,803,665,684]" italics="true" pageId="12" pageNumber="413">M. chehalis</emphasis>
</taxonomicName>
. Scale bars 100 µm.
</paragraph>
</caption>
<paragraph id="D945890EFFE14779FF55B900FCC6FA5A" blockId="12.[123,718,801,1497]" lastBlockId="12.[123,883,1505,1529]" pageId="12" pageNumber="413">
<emphasis id="EB8E551CFFE14779FF55B900FF5BFC9A" box="[171,238,801,825]" italics="true" pageId="12" pageNumber="413">Elytra</emphasis>
(
<figureCitation id="41C1958BFFE14779FEFCB900FEFBFC9A" box="[258,334,801,825]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="12" pageNumber="413">Figs 2</figureCitation>
,
<figureCitation id="41C1958BFFE14779FE9EB900FEC5FC9A" box="[352,368,801,825]" captionStart="FIGURE" captionStartId="12.[778,790,1434,1453]" captionTargetBox="[770,1170,733,1402]" captionTargetId="figure-360@12.[744,1176,710,1422]" captionText="FIGURE 9. Left elytron of female Medusapyga alsea, showing fixed setae. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12612692" httpUri="https://zenodo.org/record/12612692/files/figure.png" pageId="12" pageNumber="413">9</figureCitation>
). Entire, concealing pygidium. Together elongate oval in dorsal aspect. Each elytron with lateral margin serrulate from humerus to plical crossing. Each elytron lacks basal margination, i.e., no carina from the humerus to the parascutellar seta. Lateral explanation of each elytron distinctly reflexed, narrow, narrowest just posterior of humerus and just anterior of the plical crossing. Each elytron with elytral plica distinctly crossed. Each elytron with five shallow and indistinct striae, only stria 1 complete to apex (although extremely faint in the apical quarter), all others interrupted and ending before elytral apex. Intervals each with a row of sparse, short setae. Elytral apices separately rounded. The elytral lateral umbiculate series consists of nine large setae in “
<typeStatus id="064137ACFFE14779FE23BEC0FDA1FB5A" box="[477,532,1249,1273]" pageId="12" pageNumber="413">Type</typeStatus>
A” arrangement (sensu
<bibRefCitation id="BD6BF4FFFFE14779FF39BF20FED1FABA" author="JEANNEL, R." box="[199,356,1281,1305]" pageId="12" pageNumber="413" pagination="33 - 204" refId="ref16297" refString="JEANNEL, R. 1963 a. Monographie des &quot; Anillini &quot;, bembidiides endoges (Coleoptera: Trechidae). Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28 (2): 33 - 204." type="journal article" year="1963">Jeannel 1963a</bibRefCitation>
), with seta Eo9 mesad of margin and forming a geminate pair with seta Eo8 (
<figureCitation id="41C1958BFFE14779FD87BF00FD75FA9A" box="[633,704,1313,1337]" captionStart="FIGURE" captionStartId="12.[778,790,1434,1453]" captionTargetBox="[770,1170,733,1402]" captionTargetId="figure-360@12.[744,1176,710,1422]" captionText="FIGURE 9. Left elytron of female Medusapyga alsea, showing fixed setae. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12612692" httpUri="https://zenodo.org/record/12612692/files/figure.png" pageId="12" pageNumber="413">Fig. 9</figureCitation>
). Each elytron has a large parascutellar umbiculate seta at the base of stria 2 and a large fixed seta at the apex of stria 2, while stria 3 has three large, socketed setae: one at a level posterad of lateral umbiculate seta Eo3, one at a level anterior of lateral umbiculate seta Eo5, and the last at a level posterad of the geminate pair of lateral umbiculate setae.
</paragraph>
<caption id="8D85D986FFE14779FCF4BFBBFBC3FA65" ID-DOI="http://doi.org/10.5281/zenodo.12612692" ID-Zenodo-Dep="12612692" httpUri="https://zenodo.org/record/12612692/files/figure.png" pageId="12" pageNumber="413" startId="12.[778,790,1434,1453]" targetBox="[770,1170,733,1402]" targetPageId="12">
<paragraph id="D945890EFFE14779FCF4BFBBFBC3FA65" pageId="12" pageNumber="413">
<superScript id="2E8F2446FFE14779FCF4BFBBFB20FA0F" attach="right" box="[778,1173,1433,1453]" fontSize="8" pageId="12" pageNumber="413">
<smallCapsWord id="DFA31FD2FFE14779FCF4BFBBFCF2FA0F" baselines="1448,1448" box="[778,839,1434,1453]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="12" pageNumber="413">FIGURE</smallCapsWord>
9. Left elytron of female
<taxonomicName id="1EFAF28DFFE14779FBD6BFB8FB20FA0F" box="[1064,1173,1433,1452]" pageId="12" pageNumber="413">
<emphasis id="EB8E551CFFE14779FBD6BFB8FB20FA0F" box="[1064,1173,1433,1452]" italics="true" pageId="12" pageNumber="413">Medusapyga</emphasis>
</taxonomicName>
</superScript>
<emphasis id="EB8E551CFFE14779FD14BF93FCA2FA66" box="[746,791,1458,1477]" italics="true" pageId="12" pageNumber="413">alsea</emphasis>
, showing fixed setae. Scale bar 100 µm.
</paragraph>
</caption>
<paragraph id="D945890EFFE14779FF55BC20FE32F9BA" blockId="12.[123,1174,1537,1721]" box="[171,391,1537,1561]" pageId="12" pageNumber="413">
<emphasis id="EB8E551CFFE14779FF55BC20FE98F9BA" box="[171,301,1537,1561]" italics="true" pageId="12" pageNumber="413">Hind wings.</emphasis>
Absent.
</paragraph>
<paragraph id="D945890EFFE14779FF55BC03FB23F91A" blockId="12.[123,1174,1537,1721]" pageId="12" pageNumber="413">
<emphasis id="EB8E551CFFE14779FF55BC03FF51F99A" box="[171,228,1570,1593]" italics="true" pageId="12" pageNumber="413">Legs.</emphasis>
Male profemur with large, ventrally directed spine near the base (
<figureCitation id="41C1958BFFE14779FC59BC00FBA9F99A" box="[935,1052,1569,1593]" captionStart="FIGURE" captionStartId="9.[157,169,1658,1677]" captionTargetBox="[129,1168,1135,1647]" captionTargetId="figure-299@9.[120,1176,1127,1655]" captionTargetPageId="9" captionText="FIGURE 4. Left profemora, anterior view. (A) Medusapyga alsea, male. (B) M. alsea, female. (C) M. chehalis, male. (D) M. chehalis, female. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067228" httpUri="https://zenodo.org/record/11067228/files/figure.png" pageId="12" pageNumber="413">Fig. 4A, C</figureCitation>
); this spine projects well beyond the ventral face, with its length about two to three times its greatest diameter. Ventral face of male profemur carinate and obtusely angulate from femoral base to spine. Femora sparsely setose, more densely so near apices. Tibiae with basal half sparsely setose, apical half densely setose. Protibia with external apical angle obliquely truncate, apical one-fourth inflexed.
</paragraph>
<paragraph id="D945890EFFE04778FF85BAFAFE38FDF0" blockId="13.[123,622,219,1011]" pageId="13" pageNumber="414">
Protarsus with five tarsomeres; basal two tarsomeres of males strongly asymmetrically laterally expanded (both with proximal anterior angle distinctly protruding beyond the anteromedial margin) (
<figureCitation id="41C1958BFFE04778FECBBB7AFECFFED0" box="[309,378,347,371]" captionStart="Figure 5" captionStartId="10.[879,935,563,582]" captionTargetBox="[849,1170,220,542]" captionTargetId="figure-335@10.[844,1176,216,547]" captionText="Figure 5 (above). Male right pro- tarsi, dorsal view. (A) Medusapyga alsea. (B) M. chehalis. The view of the two specimens is not exactly equivalent, which accounts for some differences between the two images. Scale bars 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12612690" httpUri="https://zenodo.org/record/12612690/files/figure.png" pageId="13" pageNumber="414">Fig. 5</figureCitation>
); first tarsomere ventrally with two rows of between 4 and 7 adhesive articulo-setae, second tarsomere with one row of about 4 or 5 articulo-setae, both tarsomeres with some of the articulo-setae fused apically (
<figureCitation id="41C1958BFFE04778FF19BBDAFE8BFDB0" box="[231,318,507,531]" captionStart="FIGURE" captionStartId="13.[683,695,929,948]" captionTargetBox="[653,1171,220,909]" captionTargetId="figure-294@13.[648,1176,216,914]" captionText="FIGURE 10. Ventral surface of basal two protarsomeres of male Medusapyga alsea. Protarsomere 1 is to the left of center, and protarsomere 2 to the right of center. (A) Protar- somere 1 showing two rows of adhesive setae, and protar- somere 2 showing one row of adhesive setae with partially separated apices. (B) From a second specimen, showing the fusion of apices of both rows of adhesive setae on protar- somere 1, and the fusion of apices of all setae on protar- somere 2. Scale bars 25 µm." figureDoi="http://doi.org/10.5281/zenodo.12612694" httpUri="https://zenodo.org/record/12612694/files/figure.png" pageId="13" pageNumber="414">Fig. 10</figureCitation>
); female basal tarsomeres symmetrical, not laterally expanded, and without ventral articulo-setae.
</paragraph>
<paragraph id="D945890EFFE04778FF55B87AFEC9FC51" blockId="13.[123,622,219,1011]" pageId="13" pageNumber="414">
<emphasis id="EB8E551CFFE04778FF55B87AFE37FDD0" box="[171,386,603,627]" italics="true" pageId="13" pageNumber="414">Abdominal ventrites</emphasis>
. Intercoxal process of second visible abdominal ventrite triangular, apex obtuse to acute. Second visible abdominal ventrite longest, about as long as the following two. Apical margin of last visible abdominal ventrite entire, not serrate. Females with two pair of long paramedial fixed setae just anterior of apex of last visible ventrite, males with a single pair. A small shallow fovea, variable among species and sexes, is present on the second visible abdominal ventrite just posterior of the intercoxal process between the metacoxae.
</paragraph>
<caption id="8D85D986FFE04778FD55B980FC35FB22" ID-DOI="http://doi.org/10.5281/zenodo.12612694" ID-Zenodo-Dep="12612694" httpUri="https://zenodo.org/record/12612694/files/figure.png" pageId="13" pageNumber="414" startId="13.[683,695,929,948]" targetBox="[653,1171,220,909]" targetPageId="13">
<paragraph id="D945890EFFE04778FD55B980FC35FB22" pageId="13" pageNumber="414">
<subScript id="457E8B4BFFE04778FD55B980FB23FC10" attach="right" box="[683,1174,928,948]" fontSize="8" pageId="13" pageNumber="414">
<smallCapsWord id="DFA31FD2FFE04778FD55B980FD5DFC10" baselines="943,943" box="[683,744,929,948]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="13" pageNumber="414">FIGURE</smallCapsWord>
10. Ventral surface of basal two protarsomeres of
</subScript>
male
<taxonomicName id="1EFAF28DFFE04778FD44B99BFCEFFC6E" box="[698,858,954,973]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="alsea">
<emphasis id="EB8E551CFFE04778FD44B99BFCEFFC6E" box="[698,858,954,973]" italics="true" pageId="13" pageNumber="414">Medusapyga alsea</emphasis>
</taxonomicName>
. Protarsomere 1 is to the left of
<superScript id="2E8F2446FFE04778FD75B9F5FB20FC44" attach="right" box="[651,1173,980,999]" fontSize="8" pageId="13" pageNumber="414">center, and protarsomere 2 to the right of center. (A) Protarsomere</superScript>
1 showing two rows of adhesive setae, and protarsomere 2 showing one row of adhesive setae with partially separated apices. (B) From a second specimen, showing the fusion of apices of both rows of adhesive setae on protarsomere 1, and the fusion of apices of all setae on protarsomere 2. Scale bars 25 µm.
</paragraph>
</caption>
<paragraph id="D945890EFFE04778FF55B9DAFDD7FB70" blockId="13.[123,621,1019,1171]" lastBlockId="13.[123,1174,1179,1362]" pageId="13" pageNumber="414">
<emphasis id="EB8E551CFFE04778FF55B9DAFED5FBB0" box="[171,352,1019,1043]" italics="true" pageId="13" pageNumber="414">Female genitalia</emphasis>
(
<figureCitation id="41C1958BFFE04778FE90B9DAFE1AFBB0" box="[366,431,1019,1043]" captionStart="Figure 6" captionStartId="10.[155,211,1666,1685]" captionTargetBox="[143,1176,1168,1651]" captionTargetId="figure-375@10.[120,1176,1168,1651]" captionTargetPageId="10" captionText="Figure 6. Female genitalia of Medusapyga alsea from the type locality, ventral view. (A) Overview of voucher V101505. Scale bar 100 µm. (B) Closeup of voucher V101504, showing bifurcate setae on laterotergites. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.11067230" httpUri="https://zenodo.org/record/11067230/files/figure.png" pageId="13" pageNumber="414">Fig. 6</figureCitation>
). Laterotergite IX with many long, bifurcate setae covering much of the ventroapical region. Gonocoxite 2 with a minute ensiform seta and a pair of nematiform setae, one long and the other minute, with the minute seta adjacent to the base of the long seta. Ventrite X slightly sclerotized, forming a single glabrous lobe (visible centrally in
<figureCitation id="41C1958BFFE04778FDEDBE9AFDE0FB70" box="[531,597,1211,1235]" captionStart="Figure 6" captionStartId="10.[155,211,1666,1685]" captionTargetBox="[143,1176,1168,1651]" captionTargetId="figure-375@10.[120,1176,1168,1651]" captionTargetPageId="10" captionText="Figure 6. Female genitalia of Medusapyga alsea from the type locality, ventral view. (A) Overview of voucher V101505. Scale bar 100 µm. (B) Closeup of voucher V101504, showing bifurcate setae on laterotergites. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.11067230" httpUri="https://zenodo.org/record/11067230/files/figure.png" pageId="13" pageNumber="414">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="D945890EFFE04778FF55BEFAFF02FAF1" blockId="13.[123,1174,1179,1362]" pageId="13" pageNumber="414">
<emphasis id="EB8E551CFFE04778FF55BEFAFEF2FB50" box="[171,327,1243,1267]" italics="true" pageId="13" pageNumber="414">Male genitalia</emphasis>
(
<figureCitation id="41C1958BFFE04778FEA8BEFAFE12FB50" box="[342,423,1243,1267]" captionStart="FIGURE" captionStartId="14.[155,167,910,929]" captionTargetBox="[120,624,216,894]" captionTargetId="figure-377@14.[120,645,216,902]" captionTargetPageId="14" captionText="FIGURE 11. Male aedeagus, left lateral view. (A) Medusapyga alsea, voucher JRL002. (B) M. alsea, voucher JRL013. (C) M. chehalis, voucher JRL012. (D) M. chehalis, voucher JRL010. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12613593" httpUri="https://zenodo.org/record/12613593/files/figure.png" pageId="13" pageNumber="414">Figs 11</figureCitation>
,
<figureCitation id="41C1958BFFE04778FE4DBEFAFE64FB50" box="[435,465,1243,1267]" captionStart="FIGURE" captionStartId="14.[687,699,741,760]" captionTargetBox="[648,1176,242,730]" captionTargetId="figure-414@14.[652,1176,384,729]" captionTargetPageId="14" captionText="FIGURE 12. Parameres of Medusapyga chehalis, voucher JRL012. (A) left paramere, (B) right paramere. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067236" httpUri="https://zenodo.org/record/11067236/files/figure.png" pageId="13" pageNumber="414">12</figureCitation>
). Aedeagus of typical form for a carabid (
<figureCitation id="41C1958BFFE04778FC6FBEFAFC6BFB50" box="[913,990,1243,1267]" captionStart="FIGURE" captionStartId="14.[155,167,910,929]" captionTargetBox="[120,624,216,894]" captionTargetId="figure-377@14.[120,645,216,902]" captionTargetPageId="14" captionText="FIGURE 11. Male aedeagus, left lateral view. (A) Medusapyga alsea, voucher JRL002. (B) M. alsea, voucher JRL013. (C) M. chehalis, voucher JRL012. (D) M. chehalis, voucher JRL010. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12613593" httpUri="https://zenodo.org/record/12613593/files/figure.png" pageId="13" pageNumber="414">Fig. 11</figureCitation>
), with basal lobe open dorsally; lacking ventral setae. Parameres of typical form for a member of supertribe
<taxonomicName id="1EFAF28DFFE04778FBB6BEDAFF28FA91" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="superTribe" superTribe="Trechitae">Trechitae</taxonomicName>
, with left paramere larger (
<figureCitation id="41C1958BFFE04778FE43BF3AFDB9FA90" box="[445,524,1307,1331]" captionStart="FIGURE" captionStartId="14.[687,699,741,760]" captionTargetBox="[648,1176,242,730]" captionTargetId="figure-414@14.[652,1176,384,729]" captionTargetPageId="14" captionText="FIGURE 12. Parameres of Medusapyga chehalis, voucher JRL012. (A) left paramere, (B) right paramere. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067236" httpUri="https://zenodo.org/record/11067236/files/figure.png" pageId="13" pageNumber="414">Fig. 12</figureCitation>
); both parameres with four apical setae, both without ventral setae.
</paragraph>
</subSubSection>
</treatment>
</document>
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