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<document ID-DOI="http://doi.org/10.5281/zenodo.6870843" ID-GBIF-Dataset="8a6c6dcf-1911-46fc-8ea0-48fe55c5ed11" ID-ISBN="978-84-16728-08-4" ID-Zenodo-Dep="6870843" checkinTime="1658335596803" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2018" docId="3D474A54A0548739FAE2AD971601FDCB" docLanguage="en" docName="hbmw_8_Soricidae_0332.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos, Barcelona: Lynx Edicions" docTitle="Crocidura roosevelti" docType="treatment" docVersion="4" lastPageNumber="522" masterDocId="C17E322CA0288744FF8DAB47125EFFF9" masterDocTitle="Soricidae" masterLastPageNumber="551" masterPageNumber="332" pageNumber="521" updateTime="1658509882670" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Soricidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2018</mods:dateIssued>
<mods:dateOther type="pubDate">2018-07-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>332</mods:start>
<mods:end>551</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6870843</mods:identifier>
<mods:identifier type="GBIF-Dataset">8a6c6dcf-1911-46fc-8ea0-48fe55c5ed11</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-08-4</mods:identifier>
<mods:identifier type="Zenodo-Dep">6870843</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6870379" ID-Zenodo-Dep="6870379" LSID="urn:lsid:plazi:treatment:3D474A54A0548739FAE2AD971601FDCB" httpUri="http://treatment.plazi.org/id/3D474A54A0548739FAE2AD971601FDCB" lastPageId="125" lastPageNumber="522" pageId="124" pageNumber="521">
<heading pageId="124" pageNumber="521">
<subSubSection box="[1391,1473,1744,1786]" pageId="124" pageNumber="521" type="multiple">
<paragraph blockId="124.[1389,2593,1744,1865]" box="[1391,1473,1744,1786]" pageId="124" pageNumber="521">
<figureCitation box="[1391,1473,1744,1786]" captionStart="Plate 23: Soricidae" captionStartId="122.[130,160,3095,3120]" captionTargetBox="[13,2739,15,3635]" captionTargetPageId="121" captionText="346. Mdumas White-toothed Shrew (Crocidura mdumaz), 347. Montane White-toothed Shrew (Crocidura montis), 348. Newmarks White-toothed Shrew (Crocidura newmarki), 349. Munissis White-toothed Shrew (Crocidura munissi), 350. Tanzanian White-toothed Shrew (Crocidura tansaniana), 351. Usambara White-toothed Shrew (Crocidura usambarae), 352. Kivu White-toothed Shrew (Crocidura kivuana), 353. Niobes White-toothed Shrew (Crocidura niobe), 354. Desert White-toothed Shrew (Crocidura smith), 355. Sahelian Tiny White-toothed Shrew (Crocidura pasha), 356. Roosevelt's White-toothed Shrew (Crocidura roosevelti), 357. Long-footed White-toothed Shrew (Crocidura crenata), 358. Grasses White-toothed Shrew (Crocidura grasset), 359. Bicolored African White-toothed Shrew (Crocidura fuscomurina), 360. Flat-headed White-toothed Shrew (Crocidura planiceps), 361. Crosses White-toothed Shrew (Crocidura crosser), 362. Jouvenets White-toothed Shrew (Crocidura jouvenetae), 363. Mauritanian White-toothed Shrew (Crocidura lusitania), 364. Naked-tailed White-toothed Shrew (Crocidura lttoralis), 365. Gracile White-toothed Shrew (Crocidura maurisca), 366. Reddish-gray White-toothed Shrew (Crocidura cyanea), 367. Swamp White-toothed Shrew (Crocidura mariquensis), 368. Lesser Gray-brown White-toothed Shrew (Crocidura silacea), 369. Dent's White-toothed Shrew (Crocidura denti), 370. Hildegardes White-toothed Shrew (Crocidura hildegardeae), 371. Mamfe White-toothed Shrew (Crocidura virgata), 372. Batess White-toothed Shrew (Crocidura batesi), 373. Sao Tome White-toothed Shrew (Crocidura thomensis), 374. Fingui White-toothed Shrew (Crocidura fingui), 375. Frasers White-toothed Shrew (Crocidura poensis), 376. Nigerian White-toothed Shrew (Crocidura nigeriae), 377. Hun White-toothed Shrew (Crocidura attila), 378. Foxs White-toothed Shrew (Crocidura foxi), 379. Bittikofers White-toothed Shrew (Crocidura buettikoferi), 380. Thereses White-toothed Shrew (Crocidura theresae), 381. Large-headed White-toothed Shrew (Crocidura grandiceps), 382. Wimmers White-toothed Shrew (Crocidura wimmenr), 383. African Black White-toothed Shrew (Crocidura nigrofusca), 384. Savanna Dwarf White-toothed Shrew (Crocidura nanilla), 385. Nimba White-toothed Shrew (Crocidura nimbae)" figureDoi="http://doi.org/10.5281/zenodo.6871925" httpUri="https://zenodo.org/record/6871925/files/figure.png" pageId="124" pageNumber="521">356.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[1490,2176,1744,1786]" pageId="124" pageNumber="521" type="vernacular_names">
<paragraph blockId="124.[1389,2593,1744,1865]" box="[1490,2176,1744,1786]" pageId="124" pageNumber="521">
<vernacularName box="[1490,2176,1744,1786]" pageId="124" pageNumber="521">Roosevelt's White-toothed Shrew</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[2248,2592,1744,1786]" pageId="124" pageNumber="521" type="nomenclature">
<paragraph blockId="124.[1389,2593,1744,1865]" box="[2248,2592,1744,1786]" pageId="124" pageNumber="521">
<taxonomicName baseAuthorityName="Heller" baseAuthorityYear="1910" box="[2248,2592,1744,1786]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="124" pageNumber="521" phylum="Chordata" rank="species" species="roosevelti">
<emphasis box="[2248,2592,1744,1786]" italics="true" pageId="124" pageNumber="521">Crocidura roosevelti</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="vernacular_names">
<paragraph blockId="124.[1389,2593,1744,1865]" box="[1390,2494,1804,1825]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1390,1466,1804,1825]" pageId="124" pageNumber="521">French:</emphasis>
<vernacularName box="[1475,1708,1804,1825]" pageId="124" pageNumber="521">Crocidure de Roosevelt</vernacularName>
/
<emphasis bold="true" box="[1729,1820,1804,1825]" pageId="124" pageNumber="521">German:</emphasis>
<vernacularName box="[1830,2136,1804,1825]" pageId="124" pageNumber="521">Roosevelt-WeilRzahnspitzmaus</vernacularName>
/
<emphasis bold="true" box="[2157,2248,1804,1825]" pageId="124" pageNumber="521">Spanish:</emphasis>
<vernacularName box="[2258,2494,1804,1825]" pageId="124" pageNumber="521">Musarana de Roosevelt</vernacularName>
</paragraph>
<paragraph blockId="124.[1389,2593,1744,1865]" box="[1390,1827,1844,1865]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1390,1637,1844,1865]" pageId="124" pageNumber="521">Other common names:</emphasis>
<vernacularName box="[1647,1827,1844,1865]" pageId="124" pageNumber="521">Roosevelts Shrew</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection pageId="124" pageNumber="521" type="reference_group">
<paragraph blockId="124.[2002,2157,1918,1944]" pageId="124" pageNumber="521">
Taxonomy.
<taxonomicName authority="Heller, 1910" authorityName="Heller" authorityYear="1910" class="Mammalia" family="Soricidae" genus="Heliosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="124" pageNumber="521" phylum="Chordata" rank="species" species="roosevelti">Heliosorex roosevelti Heller, 1910</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="materials_examined">
<paragraph pageId="124" pageNumber="521">
<materialsCitation country="Uganda" location="Rhino Camp" pageId="124" pageNumber="521" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:3D474A54A0548739FAE2AD971601FDCB:B031AD99A0548738F7A6ACDB1ABAF844" box="[2091,2276,1948,1981]" country="Uganda" name="Rhino Camp" pageId="124" pageNumber="521">Rhino Camp</location>
,
<location LSID="urn:lsid:plazi:treatment:3D474A54A0548739FAE2AD971601FDCB:B031AD99A0548738F775ACDB1BE5F844" box="[2296,2491,1948,1981]" country="Uganda" name="Lado Enclave" pageId="124" pageNumber="521">Lado Enclave</location>
,
<collectingCountry name="Uganda" pageId="124" pageNumber="521">Uganda</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="discussion">
<paragraph pageId="124" pageNumber="521">
<taxonomicName baseAuthorityName="Heller" baseAuthorityYear="1910" box="[2006,2256,2027,2060]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="124" pageNumber="521" phylum="Chordata" rank="species" species="roosevelti">Crocidura roosevelti</taxonomicName>
is the type species of subgenus Heliosorex, named by E. Heller in 1910. According to genetic studies by S. Quérouil and others in 2005, it does not cluster with
<taxonomicName authorityName="Peters" authorityYear="1876" box="[2236,2393,2185,2218]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="124" pageNumber="541" phylum="Chordata" rank="species" species="dolichura">C. dolichura</taxonomicName>
and
<taxonomicName authorityName="Brosset, Dubost &amp; Heim de Balsac" authorityYear="1965" box="[2468,2597,2185,2218]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="124" pageNumber="524" phylum="Chordata" rank="species" species="crenata">C. crenata</taxonomicName>
as suggested by R. Hutterer in 1981 and 1994. In a recent phylogenetic study by S
</paragraph>
<paragraph lastBlockId="124.[1390,2597,2343,3478]" pageId="124" pageNumber="521">Dubey and colleagues in 2008,it is closely related to West Palearctic C. russalaEe C. pachyura and in a basal position to all other Afrotropical Crocidurines. Monotypic.</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6871701" ID-Zenodo-Dep="6871701" httpUri="https://zenodo.org/record/6871701/files/figure.png" inLine="true" pageId="124" pageNumber="521" targetBox="[1388,1979,1918,2332]" targetPageId="124">
<paragraph blockId="124.[1390,2597,2343,3478]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1390,1566,2430,2455]" pageId="124" pageNumber="521">Distribution.</emphasis>
Forest-savanna margin of the African forest block; there are records from Cameroon, Central African Republic, DR Congo, Uganda, Rwanda, Tanzania, and NE Angola, and possibly present in South Sudan.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="description">
<paragraph blockId="124.[1390,2597,2343,3478]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1391,1644,2540,2573]" pageId="124" pageNumber="521">Descriptive notes.</emphasis>
Head-body 62-82 mm, tail 59-69 mm, ear 6 mm, hindfoot 14-15 mm; weight 9 g. Condylo-incisive lengths are 20-7-22 mm. Roosevelt's Whitetoothed Shrew is small and pale cinnamon-brown, with tail 85-96% of head-body length. It is similar to the Somali White-toothed Shrew (
<taxonomicName authorityName="Thomas" authorityYear="1895" box="[2204,2361,2662,2691]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="124" pageNumber="536" phylum="Chordata" rank="species" species="somalica">C. somalica</taxonomicName>
), except it has a more reddish hue and longertail, which is bicolored and nearly naked. Dorsal pelage is grayish brown, with gray-based hairs and cinnamon-brown tips. Ventral pelage is whitish gray, with gray-based hairs and white tips. Head is similar to dorsal color. Ears are relatively large. Forefeet and hindfeet are flesh-colored, with whitish hairs. Soles of hindfeet have granulae between plantar pads. Tail is distinctly bicolored, being cinnamon-brown above and white below. Tail has only a few short bristle hairs and appears naked (pilosity of short hairs ¢.70%). Tails of Angola specimens are 86-94% of head-body length. Skull is elongated and gracile. Interorbital region is narrow and almost parallel. Braincase is slightly arched, and dentition is weak. I' is moderately long and hooked. M? is narrow to medium-sized.
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="biology_ecology">
<paragraph blockId="124.[1390,2597,2343,3478]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1391,1502,3091,3124]" pageId="124" pageNumber="521">Habitat.</emphasis>
Moist savanna around the Congo Basin forest block. Roosevelt's Whitetoothed Shrew inhabits lowlands at elevations below 1200 m. In Rwenzori National Park, specimens were found in Imperata-Cymbopogon grass and Capparis-Euphorbia bushland mosaic.
</paragraph>
</subSubSection>
<subSubSection box="[1391,1892,3253,3282]" pageId="124" pageNumber="521" type="food_feeding">
<paragraph blockId="124.[1390,2597,2343,3478]" box="[1391,1892,3253,3282]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1391,1653,3253,3282]" pageId="124" pageNumber="521">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection box="[1391,1765,3289,3322]" pageId="124" pageNumber="521" type="breeding">
<paragraph blockId="124.[1390,2597,2343,3478]" box="[1391,1765,3289,3322]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1391,1526,3289,3322]" pageId="124" pageNumber="521">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection box="[1390,1864,3332,3361]" pageId="124" pageNumber="521" type="activity">
<paragraph blockId="124.[1390,2597,2343,3478]" box="[1390,1864,3332,3361]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1390,1625,3332,3361]" pageId="124" pageNumber="521">Activity patterns.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="521" type="biology_ecology">
<paragraph blockId="124.[1390,2597,2343,3478]" pageId="124" pageNumber="521">
<emphasis bold="true" box="[1391,2104,3367,3400]" pageId="124" pageNumber="521">Movements, Home range and Social organization.</emphasis>
Roosevelts White-toothed Shrews are occasionally found in pellets of common barn owls (7yto alba) in Garamba Na- tional Park, DR Congo (five of 172 shrews from 13 species).
</paragraph>
</subSubSection>
<subSubSection pageId="125" pageNumber="522" type="conservation">
<paragraph blockId="125.[164,1369,297,565]" pageId="125" pageNumber="522">
<emphasis bold="true" box="[165,512,297,326]" pageId="125" pageNumber="522">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Roosevelt's White-toothed Shrew is widespread, and it does not appear to face major threats. It is found in Queen Elizabeth National Park in Uganda and Garamba National Park in north-eastern DR Congo.
</paragraph>
</subSubSection>
<subSubSection pageId="125" pageNumber="522" type="bibRefCitation_list">
<paragraph blockId="125.[164,1369,297,565]" pageId="125" pageNumber="522">
<emphasis bold="true" box="[166,319,458,483]" pageId="125" pageNumber="522">Bibliography.</emphasis>
Dubey, Salamin et al. (2008), Happold (2013j), Heim de Balsac &amp; Meester (1977), Heim de Balsac &amp; Verschuren (1968), Heller (1910), Hutterer (1981b, 1993, 2005b), Hutterer &amp; Kerbis Peterhans (2016), Hutterer, Van der Straeten &amp; Verheyen (1987), Quérouil et al. (2005), Thorn et al. (2009).
</paragraph>
</subSubSection>
</treatment>
</document>