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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZBK="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel &amp; Njunjic, Iva" docDate="2018" docId="2AB68B28E8199BA4DA5006D53C6A25A8" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaginus muluensis Schilthuizen &amp; Perreau, sp. n." docType="treatment" docUuid="0A383A2B-9730-409A-98F0-7EAAF851B15F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="75" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="73" updateTime="1668166102793" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Njunjic, Iva</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>777</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>108</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.777.23212</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.777.23212</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-777-57</mods:identifier>
<mods:identifier type="ZBK">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
<mods:identifier type="ZooBank">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="146746176" LSID="urn:lsid:zoobank.org:act:0A383A2B-9730-409A-98F0-7EAAF851B15F" httpUri="http://treatment.plazi.org/id/2AB68B28E8199BA4DA5006D53C6A25A8" lastPageId="18" lastPageNumber="75" pageId="16" pageNumber="73">
<subSubSection pageId="16" pageNumber="73" type="nomenclature">
<paragraph pageId="16" pageNumber="73">
<taxonomicName LSID="http://zoobank.org/0A383A2B-9730-409A-98F0-7EAAF851B15F" authority="Schilthuizen &amp; Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus muluensis" order="Coleoptera" pageId="16" pageNumber="73" phylum="Arthropoda" rank="species" species="muluensis">Ptomaphaginus muluensis Schilthuizen &amp; Perreau</taxonomicName>
<taxonomicNameLabel pageId="16" pageNumber="73">sp. n.</taxonomicNameLabel>
Figures 5b, 7e, f, 9g, 10e
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="74" type="materials_examined">
<paragraph pageId="17" pageNumber="74">
<pageBreakToken pageId="17" pageNumber="74" start="start">Material</pageBreakToken>
.
</paragraph>
<paragraph pageId="17" pageNumber="74">
Holotype: Malaysia, Sarawak, Mulu National Park, Mixed dipterocarp forest, TPS 7-10,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, NHMUK, B.M. 1978-49), male. Paratypes: Sarawak. Mulu National Park, mixed dipterocarp forest, TPS 7-10,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, NHMUK, B.M. 1978-49), 4 males, 5 females; Mulu National Park, Slope, TPS 13-16,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, NHMUK, B.M. 1978-49), 9 males, 6 females; Mulu National Park, Slope, TPS 17-21,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, NHMUK, B.M. 1978-49), 4 males, 5 females; Mulu National Park, Slope, TPS 5-6,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, NHMUK, B.M. 1978-49), 1 male.
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="75" pageId="17" pageNumber="74" type="description">
<paragraph pageId="17" pageNumber="74">Description.</paragraph>
<paragraph lastPageId="18" lastPageNumber="75" pageId="17" pageNumber="74">
Length 2.5-3.1 mm. Habitus: reddish brown; relatively parallel-sided and somewhat flattened, head broad, pronotum 1.6 times as wide as long, as wide as the elytra, caudal angles clearly extended. Elytra of moderate length, gently convex, in the caudal one-third gently rounded towards the apex, jointly ca. 1.4 times as long as wide (length measured from the caudal tip of the scutellum to the apex of the elytra). Body entirely covered in dense golden-yellow setation. Wings present. Antennae slender, 4th, 9th, and 10th antennomeres slightly wider than long. Male protarsi slightly dilated, the first four tarsomeres jointly ca. three times as long as wide. Aedeagus gently bent ventrad, in lateral view apically flattened and ending in a bulbous upturned tip, in dorsal view gradually narrowing towards the apex. Stylet long and thin, hair-like.
<pageBreakToken pageId="18" pageNumber="75" start="start">Spiculum</pageBreakToken>
gastrale long-ovoid, with the caudal part truncated. Spermatheca J-shaped, with 7-10 narrow rings on the long shaft of the
<normalizedToken originalValue="“J”">&quot;J&quot;</normalizedToken>
, and 5-7 broader rings on the curved part. Spermiduct long, thin, consisting of at least ten 360° coils.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="75" type="differential diagnosis">
<paragraph pageId="18" pageNumber="75">Differential diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="75">
Very similar to
<taxonomicName lsidName="P. louis" pageId="18" pageNumber="75" rank="species" species="louis">P. louis</taxonomicName>
, but externally distinguished by the larger size and more slender habitus, with longer elytra, broader pronotum and distinctly drawn-out caudal pronotal angles. Aedeagus in dorsal view tapering gradually towards the apex, not as abruptly as in
<taxonomicName lsidName="P. louis" pageId="18" pageNumber="75" rank="species" species="louis">P. louis</taxonomicName>
; in lateral view, the apex is not as convex. Spermatheca distinguished from
<taxonomicName lsidName="P. louis" pageId="18" pageNumber="75" rank="species" species="louis">P. louis</taxonomicName>
by the J-shape with distinctive rings.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="75" type="habitat">
<paragraph pageId="18" pageNumber="75">Habitat and distribution.</paragraph>
<paragraph pageId="18" pageNumber="75">
Only known from Mulu National Park, Sarawak. Several specimens infected with black
<taxonomicName lsidName="Staphylinoidea" order="Laboulbeniales" pageId="18" pageNumber="75" rank="order">Laboulbeniales</taxonomicName>
on the elytra and the pygidium.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="75" type="etymology">
<paragraph pageId="18" pageNumber="75">Etymology.</paragraph>
<paragraph pageId="18" pageNumber="75">Named after Mulu National Park, to date the only locality from which this species is known.</paragraph>
</subSubSection>
</treatment>
</document>