258 lines
28 KiB
XML
258 lines
28 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E782DFFEDFF013DC2ACFAE996" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Flagellobolus pauliani Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="ADC7576D-F0BB-4D2B-B29C-23EFBA8679A3" docUuidSource="ZooBank" docVersion="10" lastPageNumber="44" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="43" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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||
</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791407" ID-GBIF-Taxon="163747646" ID-Zenodo-Dep="3791407" LSID="urn:lsid:zoobank.org:act:ADC7576D-F0BB-4D2B-B29C-23EFBA8679A3" httpUri="http://treatment.plazi.org/id/1772122E782DFFEDFF013DC2ACFAE996" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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||
<subSubSection box="[140,590,694,721]" pageId="42" pageNumber="43" type="nomenclature">
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||
<paragraph blockId="42.[140,995,694,757]" box="[140,590,694,721]" pageId="42" pageNumber="43">
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||
<heading box="[140,590,694,721]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="3">
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<emphasis bold="true" box="[140,590,694,721]" pageId="42" pageNumber="43">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,513,694,721]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,405,694,721]" italics="true" pageId="42" pageNumber="43">Flagellobolus pauliani</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[525,590,695,721]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,995,730,757]" pageId="42" pageNumber="43" type="description">
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||
<paragraph blockId="42.[140,995,694,757]" box="[140,995,730,757]" pageId="42" pageNumber="43">
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||
<uri box="[140,995,730,757]" pageId="42" pageNumber="43">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[415,995,730,757]" pageId="42" pageNumber="43">ADC7576D-F0BB-4D2B-B29C-23EFBA8679A3</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="42" pageNumber="43" type="materials_examined">
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis bold="true" box="[140,373,800,826]" pageId="42" pageNumber="43">
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<emphasis box="[140,241,800,826]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608818270" collectingDate="1955-12-20" collectionCode="MNHN" collectorName="R. Paulian" country="Madagascar" latitude="-18.95" location="NoNumber" longLatPrecision="1273" longitude="48.433334" municipality="Perinet" pageId="42" pageNumber="43" specimenCount="1" typeStatus="holotype">
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2 ♂
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<emphasis box="[436,539,800,826]" italics="true" pageId="42" pageNumber="43">
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<typeStatus box="[436,533,800,826]" pageId="42" pageNumber="43">Holotype</typeStatus>
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:
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</emphasis>
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1 ♂ (
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<quantity box="[611,696,800,827]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="42" pageNumber="43" unit="mm" value="31.0">31 mm</quantity>
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long),
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<collectionCode box="[781,877,801,827]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">MNHN</collectionCode>
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<location LSID="urn:lsid:plazi:treatment:1772122E782DFFEDFF013DC2ACFAE996:9A04F5E3782DFFECFCFB3C54ACA5E96B" box="[886,1020,800,826]" country="Madagascar" latitude="-18.95" longLatPrecision="1273" longitude="48.433334" municipality="Perinet" name="NoNumber" pageId="42" pageNumber="43">NoNumber</location>
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,
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<collectingCountry name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
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,
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<collectingMunicipality box="[220,303,835,861]" pageId="42" pageNumber="43">Périnet</collectingMunicipality>
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, rainforest,
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<geoCoordinate box="[436,535,835,862]" degrees="18" direction="south" minutes="57" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-18.95">18°57’ S</geoCoordinate>
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,
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<geoCoordinate box="[547,663,835,862]" degrees="048" direction="east" minutes="26" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="48.433334">048°26’ E</geoCoordinate>
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, leg.
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<collectorName box="[722,842,835,862]" pageId="42" pageNumber="43">R. Paulian</collectorName>
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,
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<date box="[854,998,835,862]" pageId="42" pageNumber="43" value="1955-12-20">
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<collectingDate box="[854,998,835,862]" pageId="42" pageNumber="43" value="1955-12-20">20.XII.1955</collectingDate>
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</date>
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, IRSM.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis bold="true" box="[185,510,870,896]" pageId="42" pageNumber="43">
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<emphasis box="[185,257,870,896]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Other</emphasis>
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material examined
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</emphasis>
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:
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<materialsCitation ID-GBIF-Occurrence="2620669304" collectingDate="2000-12-05" collectingDateMax="2000-12-13" collectingDateMin="2000-12-05" collectorName="Fisher & Griswold" elevation="1300" latitude="-18.473333" location="Andranomay" longLatPrecision="21" longitude="47.96" pageId="42" pageNumber="43" specimenCount="2" specimenCount-male="1" stateProvince="Antananarivo">
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<specimenCount box="[538,596,871,897]" pageId="42" pageNumber="43" type="male">1 ♂</specimenCount>
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,
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<specimenCount box="[622,721,871,897]" pageId="42" pageNumber="43" type="generic">1 imm.</specimenCount>
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, CASENT 9032806,
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<collectingRegion country="Madagascar" name="Antananarivo" pageId="42" pageNumber="43">Province d’Antananarivo</collectingRegion>
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,
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<quantity box="[330,391,906,932]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="km" value="3.0">3 km</quantity>
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41°NE
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<location LSID="urn:lsid:plazi:treatment:1772122E782DFFEDFF013DC2ACFAE996:9A04F5E3782DFFECFE6B3CFEAD23E9F5" box="[486,634,906,932]" latitude="-18.473333" longLatPrecision="21" longitude="47.96" name="Andranomay" pageId="42" pageNumber="43" stateProvince="Antananarivo">Andranomay</location>
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,
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<locationDeviation box="[647,994,906,933]" location="Anjozorobe" pageId="42" pageNumber="43">
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<quantity box="[647,744,906,933]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="42" pageNumber="43" unit="km" value="11.5">11.5 km</quantity>
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147°SSE Anjozorobe
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</locationDeviation>
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, montane rainforest,
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<quantity box="[261,352,941,968]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="42" pageNumber="43" unit="m" value="1300.0">
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<elevation box="[261,352,941,968]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="42" pageNumber="43" unit="m" value="1300.0">1300 m</elevation>
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</quantity>
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,
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<geoCoordinate box="[363,503,941,967]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="42" pageNumber="43" precision="15" seconds="24" value="-18.473333">18°28’24” S</geoCoordinate>
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,
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<geoCoordinate box="[514,656,941,968]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="42" pageNumber="43" precision="15" seconds="36" value="47.96">47°57’36” E</geoCoordinate>
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, leg.
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<collectorName box="[713,783,941,967]" pageId="42" pageNumber="43">Fisher</collectorName>
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,
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<collectorName box="[794,898,941,967]" pageId="42" pageNumber="43">Griswold</collectorName>
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et al.,
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<collectingMethod box="[972,1103,941,968]" pageId="42" pageNumber="43">pitfall traps</collectingMethod>
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,
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<date box="[140,310,976,1003]" pageId="42" pageNumber="43" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
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<collectingDate box="[140,310,976,1003]" pageId="42" pageNumber="43" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5–13.XII.2000</collectingDate>
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</date>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis bold="true" box="[185,335,1011,1037]" pageId="42" pageNumber="43">
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<emphasis box="[185,330,1011,1037]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Description</emphasis>
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.
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</emphasis>
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<emphasis box="[342,494,1012,1037]" italics="true" pageId="42" pageNumber="43">Measurements</emphasis>
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: male
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<typeStatus box="[569,668,1011,1037]" pageId="42" pageNumber="43">holotype</typeStatus>
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with 42 body rings, circa
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<quantity box="[961,1043,1012,1038]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="42" pageNumber="43" unit="mm" value="31.0">31 mm</quantity>
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long,
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<quantity box="[140,230,1047,1073]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="42" pageNumber="43" unit="mm" value="2.4">2.4 mm</quantity>
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wide.
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</paragraph>
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis box="[185,306,1082,1108]" italics="true" pageId="42" pageNumber="43">Coloration</emphasis>
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almost completely decolorized because of alcohol, light stripe visible dorsally.
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</paragraph>
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis box="[185,244,1152,1178]" italics="true" pageId="42" pageNumber="43">Head</emphasis>
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: each eye with circa 30 ocelli arranged in 5 or 6 vertical rows. Labrum with standard three irregular teeth and a single row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennal cavity absent. Antennae very long, protruding back to ring 7. Relative lengths of antennomeres: 1<2>3=4=5=6, antennomere 2 longest. Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 latero-apically four or five rows, antennomere 6 with two or three rows of sensilla basiconica.
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</paragraph>
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis box="[185,373,1399,1425]" italics="true" pageId="42" pageNumber="43">Gnathochilarium</emphasis>
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: lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Mentum basally subdivided by a prominent suture.
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<emphasis box="[352,458,1469,1495]" italics="true" pageId="42" pageNumber="43">Mandible</emphasis>
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not dissected.
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</paragraph>
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<paragraph blockId="42.[140,1108,800,1636]" box="[185,823,1504,1531]" pageId="42" pageNumber="43">
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<emphasis box="[185,274,1504,1530]" italics="true" pageId="42" pageNumber="43">Collum:</emphasis>
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smooth, laterally not protruding as far as ring 2.
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</paragraph>
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<paragraph blockId="42.[140,1108,800,1636]" pageId="42" pageNumber="43">
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<emphasis box="[185,309,1539,1565]" italics="true" pageId="42" pageNumber="43">Body rings:</emphasis>
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mostly smooth, with some irregular punctation on mesozona. Ozopores starting at ring 6, touching suture between mesozona and metazona. Ozopore on ring 6 located more ventrally than subsequent ozopores.
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</paragraph>
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<paragraph blockId="43.[140,1108,167,968]" pageId="43" pageNumber="44">
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<emphasis box="[185,252,167,193]" italics="true" pageId="43" pageNumber="44">Telson</emphasis>
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: anal valves with weakly developed lips, with neither grooves, nor setae (
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<figureCitation box="[148,257,202,228]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23A</figureCitation>
|
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). Preanal ring and subanal process well-rounded, not extending beyond anal valves.
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</paragraph>
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<paragraph blockId="43.[140,1108,167,968]" pageId="43" pageNumber="44">
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<emphasis box="[185,230,273,298]" italics="true" pageId="43" pageNumber="44">Legs</emphasis>
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: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs as long as body diameter in males. Each podomere with a single or pair of apical, ventral seta(e). Coxae 3 and beyond of cylindrical shape (
|
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<figureCitation box="[697,799,378,404]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23B</figureCitation>
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). Tarsus with a stout dorsoapical setae and three pairs of setae; apical pair stout, others successively much smaller.
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</paragraph>
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<paragraph blockId="43.[140,1108,167,968]" pageId="43" pageNumber="44">
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<emphasis box="[185,417,448,474]" italics="true" pageId="43" pageNumber="44">Male sexual characters</emphasis>
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: tarsal pads absent (
|
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<figureCitation box="[634,733,448,474]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23B</figureCitation>
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). Coxae 3 and 4 each with a short, conical process. Process protruding posteriorly, but never overlapping adjacent coxa.
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</paragraph>
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<paragraph blockId="43.[140,1108,167,968]" pageId="43" pageNumber="44">
|
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<emphasis box="[185,371,519,545]" italics="true" pageId="43" pageNumber="44">Anterior gonopod</emphasis>
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sternite elongated into a wide, triangular lobe with a well-round- ed tip (
|
||
<figureCitation box="[223,329,554,580]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23C</figureCitation>
|
||
). Mesal process of coxite wide, long and well-rounded, shorter than sternite lobe (
|
||
<figureCitation box="[297,403,589,615]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23C</figureCitation>
|
||
). Process of telopodite large and slender, retrorse part not located apically, but basally of process (
|
||
<figureCitation box="[584,696,624,650]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23D</figureCitation>
|
||
). Retrorse projection slender, short with a well-rounded tip (
|
||
<figureCitation box="[420,527,659,686]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23D</figureCitation>
|
||
).
|
||
<emphasis box="[547,743,659,685]" italics="true" pageId="43" pageNumber="44">Posterior gonopods</emphasis>
|
||
unique (
|
||
<figureCitation box="[844,977,659,686]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Figs 23E, F</figureCitation>
|
||
). Gonopod pair connected via a large, well-sclerotized sternite (
|
||
<figureCitation box="[727,830,695,721]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23E</figureCitation>
|
||
). Telopodites positioned parallel to one another. Coxite and telopodite not separated from one another, but dividing suture well-visible (
|
||
<figureCitation box="[464,568,765,791]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23E</figureCitation>
|
||
). Coxite wide, with a single groove and no appendage (
|
||
<figureCitation box="[250,351,800,826]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23F</figureCitation>
|
||
). Telopodite completely elongated into a process, apically tapering (
|
||
<figureCitation box="[148,249,835,862]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23F</figureCitation>
|
||
), very slender. Sperm canal running along mesal margin of telopodite, apically occupying whole width of joint (
|
||
<figureCitation box="[569,670,871,897]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="43" pageNumber="44">Fig. 23F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="44" type="etymology">
|
||
<paragraph blockId="43.[140,1108,167,968]" pageId="43" pageNumber="44">
|
||
<emphasis bold="true" box="[185,323,906,932]" pageId="43" pageNumber="44">
|
||
<emphasis box="[185,318,906,932]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<emphasis box="[334,434,906,932]" italics="true" pageId="43" pageNumber="44">
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[334,429,906,932]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species">pauliani</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
adjective, after the collector Dr. Renaud Paulian (
|
||
<date pageId="43" pageNumber="44" value="1913" valueMax="2003">1913– 2003</date>
|
||
), who collected numerous interesting millipedes on
|
||
<collectingCountry box="[792,927,941,967]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |