642 lines
57 KiB
XML
642 lines
57 KiB
XML
<document id="9EEE0BF5C1442631BC75E2C2129E5D02" ID-DOI="10.3897/zookeys.1223.138220" ID-ZooBank="3557A991-63DF-42D8-B8B1-F310CD52FD00" ID-publisher-id="138220" URI-arpha="7352E8C9-65CA-546F-BE46-EF55E52C51A6" URI-zoobank="http://zoobank.org/3557A991-63DF-42D8-B8B1-F310CD52FD00" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736187739607" checkinUser="pensoft" docAuthor="Hendrich, Lars & Brancucci, Michel" docDate="2025" docId="1113CD6291B65D00858C62DFB6C9FB47" docLanguage="en" docName="ZooKeys 1223: 87-147" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/138220/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sandracottus insignis Hendrich" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="147" masterDocId="7352E8C965CA546FBE46EF55E52C51A6" masterDocTitle="Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)" masterLastPageNumber="147" masterPageNumber="87" pageNumber="87" updateTime="1736200074934" updateUser="admin">
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<mods:title id="016ECA4E6D38A7D4041BAC08192E3BE7">Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)</mods:title>
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<mods:name id="2A444F53F1D3E4862AC62EBF653AFEE2" type="personal">
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<mods:namePart id="1B5AEF452D7F01B23DF0B5A14E7704F1">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="9E3C0D52F645A99C039740B89AF86A09">SNSB - Zoologische Staatssammlung München, Münchhausenstraße 21, D – 81247 München, Germany</mods:affiliation>
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<mods:namePart id="734F18C1F3E48B050280B7C96B78D559">Brancucci, Michel</mods:namePart>
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<mods:affiliation id="98BDFE6C5704E9BC3FC64EDAAC123522">Naturhistorisches Museum Basel, Basel, Switzerland</mods:affiliation>
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<mods:typeOfResource id="B23462B93207DFAACAE1B361E34482E9">text</mods:typeOfResource>
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<mods:title id="C1D71E466C69B2CC391B0EC47683B6CE">ZooKeys</mods:title>
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<mods:part id="E6EFC722CC576A7CF08486BF303F8F82">
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<mods:date id="35615294A36D8D15ED09E34F91FB001E">2025</mods:date>
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<mods:number id="DF1FC0FA2FA016CEFEFC766781F14535">2025-01-06</mods:number>
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<mods:number id="CB92224C704B810D33B91AFCF59955A4">1223</mods:number>
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<mods:start id="5660F3B52F957E1F44FDC8C2C8BC7779">87</mods:start>
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<mods:classification id="E5F7DB8F110F4FD887E80EA798AF59A2">journal article</mods:classification>
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<mods:identifier id="0A39596A0A297C9ED89DED41CF10B984" type="DOI">10.3897/zookeys.1223.138220</mods:identifier>
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<mods:identifier id="1BB6427CF25F47364C32C8B7C9B6B8F7" type="ZooBank">3557A991-63DF-42D8-B8B1-F310CD52FD00</mods:identifier>
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<treatment id="1113CD6291B65D00858C62DFB6C9FB47" ID-arpha="1113CD62-91B6-5D00-858C-62DFB6C9FB47" LSID="urn:lsid:plazi:treatment:1113CD6291B65D00858C62DFB6C9FB47" httpUri="http://treatment.plazi.org/id/1113CD6291B65D00858C62DFB6C9FB47" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Dytiscinae">
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<subSubSection id="39C0B2DEE36959E25D57DBEFFB04DEAF" type="nomenclature">
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<paragraph id="AE903A684DAEC31196F34D700103F83D">
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<taxonomicName id="7BBF9388BC09A48E6516A09BDA366CA6" ID-CoL="6XHNJ" authority="(Wehncke, 1876)" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
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<emphasis id="F7A2DB88EBCC22BDBEE537E2AFBE9558" italics="true">Sandracottus insignis</emphasis>
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(
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<bibRefCitation id="21358DC9B75659817C70C6E115C250CA" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="5C7EEA78A895C6E959262DE876CCB05E" type="description">
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<paragraph id="38BD376683D2E38ACC9FD08AE53FC465">
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<figureCitation id="52EAB1D64809B5238FABF04E8AC21C45" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8–11. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">Figs 8</figureCitation>
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,
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||
<figureCitation id="426FE6F2611224E204385DA184351E97" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">19</figureCitation>
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,
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<figureCitation id="A542E94E9CF363407DD28FA8B07DCBB5" captionStart="Figure 28" captionStartId="F12" captionText="Figure 28. Distribution of Sandracottus insignis in the Philippines." figureDoi="10.3897/zookeys.1223.138220.figure28" httpUri="https://binary.pensoft.net/fig/1214419">28</figureCitation>
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,
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<figureCitation id="F3FF50FAE134CB46A55260CCE65DB8F2" captionStart="Figures 37–42" captionStartId="F16" captionText="Figures 37–42. Habitats of Sandracottus 37, 38 Laos, Attapeu prov., Annam Highlands Mts., Dong Amphan NBCA, Nong Fa [puddles near lake shore] and lake, habitat of S. hunteri 39 Laos, Katamtok, rest pools along a rocky river bed, habitat of S. maculatus (photos: J. Hájek) 40 Thailand, near Salawin NP, open and muddy puddle made by elephants (photo: J. Šťastný), and 41 Sakaerat Biosphere Reserve, rocky pool in an otherwise dry stream bed, habitats of S. hunteri and S. maculatus (photo: W. Atthakor) 42 Philippines, Luzon, Pampanga Prov., Bano San Juan, Mt. Arayat NP, spring creek, habitat of S. insignis (photo: H. Freitag)." figureDoi="10.3897/zookeys.1223.138220.figures37-42" httpUri="https://binary.pensoft.net/fig/1214423">42</figureCitation>
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||
,
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||
<figureCitation id="81DE007B9AF87E19FA44F5387D54EA96" captionStart="Figures 54–57" captionStartId="F20" captionText="Figures 54–57. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">55</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E9CCE45EFB61192B0184949B885BDE94" type="reference_group">
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||
<paragraph id="46777D61321D9F69A49B86070E18D698">
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||
<treatmentCitationGroup id="A4C234DA19208CB10039D9D8B52030FE">
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||
<treatmentCitation id="909A3CF514D5633FC165DCDF01C609CA" author="Wehncke E" page="194" year="1876">
|
||
<taxonomicName id="5977C65A54CE486B5BB4DFD85CE6AA8E" authority="Wehncke, 1876: 194" authorityName="Wehncke Hendrich & Brancucci" authorityPageNumber="194" authorityYear="1876" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
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||
<emphasis id="CFCF18A1EC1A30F4A2C9463A424209A9" italics="true">Hydaticus insignis</emphasis>
|
||
<comment id="8C7ADA1140CAC4347C68B78B56BCB133">
|
||
<bibRefCitation id="69647B1A92226F84DE7403DDCFBCD5CF" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876: 194</bibRefCitation>
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||
(type locality
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||
<collectingCountry id="D219B95D28BDD101414BB229BB08E932" name="Philippines">Philippines</collectingCountry>
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||
, “
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||
<collectingCountry id="AF307EABD07AB1D902E0C266164385F2" name="Iceland">Insel</collectingCountry>
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Luzon ”);
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||
<treatmentCitation id="222CA14A0FFDDD0C802B84AEED9C7A0E" author="Sharp D" page="687" year="1882">
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||
<bibRefCitation id="25920A137F6A9DCA968C811F12E92B93" DOI="10.5962/bhl.title.9530" author="Sharp D" firstAuthor="Sharp" page="687" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179–1003. [+ pls. 7–18] https://doi.org/10.5962/bhl.title.9530" year="1882">Sharp 1882: 687</bibRefCitation>
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||
</treatmentCitation>
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||
(descr.).
|
||
</comment>
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||
</taxonomicName>
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||
</treatmentCitation>
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="D42F68EBA639929EC3747B3514B33C51">
|
||
<treatmentCitationGroup id="1D2C06C44FB3601DB4A5B1395EA654F2">
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||
<treatmentCitation id="A42076C8C391362206DF0FE10375C64A">
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||
<taxonomicName id="B85F27AD6C64899EAB28424D863F1E29" ID-CoL="6XHNJ" authority="(Wehncke, 1876)" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="8465FBF36C1393C04853AF93F90C9077" italics="true">Sandracottus insignis</emphasis>
|
||
<comment id="2FCC7CCC05DBDB253914C36F19449A48">
|
||
(
|
||
<bibRefCitation id="7C7C80071AFF3274F0605B386A5FF4C4" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
|
||
);
|
||
<treatmentCitation id="ADDF801E20116A66BFB73A3AC1138235" author="Régimbart M" page="338" year="1899">
|
||
<bibRefCitation id="E3F94FA2132407E6A75F71B16BA472A3" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" firstAuthor="Regimbart" journalOrPublisher="Annales de la Société Entomologique de France" page="338" pagination="186-367" refId="B42" refString="Régimbart M (1899) Révision des Dytiscidae de la région Indo-Sino-Malaise. Annales de la Société Entomologique de France 68: 186–367. https://doi.org/10.1080/21686351.1899.12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart 1899: 338</bibRefCitation>
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||
</treatmentCitation>
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||
(tax., descr.);
|
||
<treatmentCitation id="F8EB058395E650A082A07772B6912441" author="Freitag H & Jäch MA & Wewalka G" page="8" year="2016">
|
||
<bibRefCitation id="4A9AAC8AF69BB700C44F4A520E7B073A" DOI="10.1080/01650424.2016.1210814" author="Freitag H & Jäch MA & Wewalka G" etAl="et al." firstAuthor="Freitag" issue="3" journalOrPublisher="Aquatic Insects" page="8" pagination="177-213" refId="B13" refString="Freitag H, Jäch MA, Wewalka G (2016) Diversity of aquatic and riparian Coleoptera of the Philippines: checklist, state of knowledge, priorities for future research and conservation. Aquatic Insects 37 (3): 177–213. https://doi.org/10.1080/01650424.2016.1210814" title="Diversity of aquatic and riparian Coleoptera of the Philippines: checklist, state of knowledge, priorities for future research and conservation." volume="37" year="2016">Freitag et al. 2016: 8</bibRefCitation>
|
||
</treatmentCitation>
|
||
(check list); Hájek and
|
||
<treatmentCitation id="C3500985E44BCC78A23FF523A81AD432" author="Nilsson A & Hájek J" page="91" year="2024">
|
||
<bibRefCitation id="2C6D7F452351019F4BEB8B34DF50797D" author="Nilsson A & Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
|
||
</treatmentCitation>
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||
(cat.).
|
||
</comment>
|
||
</taxonomicName>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="12CFEF5B8E9DB5DB6D2E94A0E4AF5CA7">
|
||
<treatmentCitationGroup id="5AAF6B28A125CFF3E8C8D728FE6D5193">
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||
<treatmentCitation id="E76A82ABAE83B3DB0FFFCEB42F51D84A" author="Régimbart M" page="99" year="1877">
|
||
<taxonomicName id="5C6AB4E45F42F5382F34C803957CB490" authority="Regimbart, 1877: 99" authorityName="Regimbart" authorityPageNumber="99" authorityYear="1877" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="baeri">
|
||
<emphasis id="757796583B679DBCC2CE940EA1D6067C" italics="true">Hydaticus baeri</emphasis>
|
||
<comment id="959E09E306C0918CBAA089461812043A">
|
||
<bibRefCitation id="74A83D33B57113547FFCA0AA2AC27F6C" author="Régimbart M" refId="B41" refString="Regimbart M (1877) Description de douze nouvelles especes d'hydrocanthares recueillis a Manille par M. Baer [sic!]. Bulletin Bimensuel des Seances de la Societe Entomologique de France 1877 (8) (Seance du 25 Avril): 99 - 100." year="1877">Régimbart, 1877: 99</bibRefCitation>
|
||
(type locality
|
||
<collectingCountry id="448F205C79ACD2DA12FE438442C8916B" name="Philippines">Philippines</collectingCountry>
|
||
, “
|
||
<collectingRegion id="6C275ADC5EF93D8B232DD248C02FDD69" country="Philippines" name="Manila">Manille</collectingRegion>
|
||
”);
|
||
<treatmentCitation id="AA6F7647D1323D7E2CE2FEFA28D08C92" author="Zimmermann A" page="234" year="1920">
|
||
<bibRefCitation id="B2B1618FF684D6095C741D8D1B80E053" author="Zimmermann A" firstAuthor="Zimmermann" journalOrPublisher="In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71" page="234" refId="B60" refString="Zimmermann A (1920) Dytiscidae. In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71, 326 pp." title="Dytiscidae." year="1920">Zimmermann 1920: 234</bibRefCitation>
|
||
</treatmentCitation>
|
||
(syn., cat.).
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="82557806C1D5E8ABA71EB51C322F082B">
|
||
<treatmentCitationGroup id="DE11324D47418F19095B077EE0994949">
|
||
<treatmentCitation id="E7572411BB8CAE2197DE6CD50DB3D1F6" author="Sharp D" page="689" year="1882">
|
||
<taxonomicName id="D8BF73871763BAC4CB7181E0B09F87CF" authority="Sharp, 1882: 689" authorityName="Sharp" authorityPageNumber="689" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="6F3DB4748F5ED9CCE58E4B34ADB2F38F" italics="true">Sandracottus ornatus</emphasis>
|
||
<comment id="7D76E827B3B5C6E1C3E5302E3B21757C">
|
||
<bibRefCitation id="FD00A117F5D9E855A8C1F950C04E1465" DOI="10.5962/bhl.title.9530" author="Sharp D" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003. [+ pls. 7 - 18] https: // doi. org / 10.5962 / bhl. title. 9530" year="1882">Sharp, 1882: 689</bibRefCitation>
|
||
(type locality Borneo);
|
||
<treatmentCitation id="338856ED5C48BCE78179AF2A1E9A0CEA" author="Vazirani TG" page="84" year="1977">
|
||
<bibRefCitation id="3CE3203EA4174CF6C587938360BA9897" author="Vazirani TG" firstAuthor="Vazirani" issue="1976" journalOrPublisher="Records of the Zoological Survey of India Miscellaneous Publication Occasional Paper" page="84" pagination="1-111" refId="B54" refString="Vazirani TG (1977) Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India Miscellaneous Publication Occasional Paper 6 (1976): 1–111." title="Catalogue of Oriental Dytiscidae." volume="6" year="1977">Vazirani 1977: 84</bibRefCitation>
|
||
</treatmentCitation>
|
||
(stat. nov.).
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D08ABC695EED1864AF7BB0E11BDA64DA">
|
||
<treatmentCitationGroup id="72C68883CE21C61D33E449377E5CA8C5">
|
||
<treatmentCitation id="1321348A5E477EC46A34CAA13AF35927">
|
||
<taxonomicName id="E93F4EA9E80553CA6BEF1410ABC9BEDB" ID-CoL="5KZSW" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subSpecies" species="insignis" subSpecies="ornatus">
|
||
<emphasis id="B502A8291AC423896CE169FFAB6D9002" italics="true">Sandracottus insignis ssp. ornatus</emphasis>
|
||
<comment id="1A944623DDDE5D447DBA264B580EFE89">
|
||
<bibRefCitation id="78EBB5E084B66AF3BECA427A4FF00D36" DOI="10.5962/bhl.title.9530" author="Sharp D" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003. [+ pls. 7 - 18] https: // doi. org / 10.5962 / bhl. title. 9530" year="1882">Sharp, 1882</bibRefCitation>
|
||
;
|
||
<treatmentCitation id="E5255202C054746FA93639E752637186" author="Vazirani TG" page="84" year="1977">
|
||
<bibRefCitation id="293836A650B73A0A38D14CA4F90B9758" author="Vazirani TG" firstAuthor="Vazirani" issue="1976" journalOrPublisher="Records of the Zoological Survey of India Miscellaneous Publication Occasional Paper" page="84" pagination="1-111" refId="B54" refString="Vazirani TG (1977) Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India Miscellaneous Publication Occasional Paper 6 (1976): 1–111." title="Catalogue of Oriental Dytiscidae." volume="6" year="1977">Vazirani 1977: 84</bibRefCitation>
|
||
</treatmentCitation>
|
||
(cat.); Hájek and
|
||
<treatmentCitation id="02C7E9A9693C0533BEDEEA84A10123A1" author="Nilsson A & Hájek J" page="91" year="2024">
|
||
<bibRefCitation id="E1B6D6AA850C18A785C71D32D07CA339" author="Nilsson A & Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
|
||
</treatmentCitation>
|
||
(cat.) (syn. nov.).
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZOBK" type="type material">
|
||
<paragraph id="8E07A24DFA7084878A7A1D207693C466">
|
||
<heading id="D304C6525AF4BFBC16500C72A91975AD" reason="title">
|
||
Type material of
|
||
<taxonomicName id="DF8D23755E000A4F48968CA5147C8944" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="A8D580B4E66F03A8AF889B81F0249DAB" italics="true">Sandracottus insignis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="90F35F69A7A787270D25C01CD7AEF155">
|
||
<emphasis id="9301E077DB611871EAC2A7AC45DDA6E9" bold="true" italics="true">
|
||
<typeStatus id="59269C8C033123005AC20DA6478FB929" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
(herewith designated): Female: “ Semper ” [white handwritten label with black fram]; “ Luzon ” [handwritten yellowish label], Ex. Coll. E. WEHNCKE Acq. 1884 ” [printed white label], “
|
||
<typeStatus id="0A8BC01DE9AEA37DBC77921D06B46611" type="lectotype">Lectotype</typeStatus>
|
||
<taxonomicName id="CDCF44EE6E49504EED48A2ACD8066834" authority="Wehncke Hendrich & Brancucci" authorityName="Wehncke Hendrich & Brancucci" authorityYear="1876" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="93212A80B80BC1A8A8E332AB9979645A" italics="true">Hydaticus insignis</emphasis>
|
||
Wehncke Hendrich & Brancucci
|
||
</taxonomicName>
|
||
des. 2010 ” [red printed label] (
|
||
<abbrev id="ABBRID0E1PBK" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="0BA101B63B8BB1DBAD968D141D76A646" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
). Examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E5PBK" type="type material">
|
||
<paragraph id="AA7BB8E8211A5249A4E9BE7B3A8B3ADE">
|
||
<heading id="E22B58CBF3294176B23447CE8C658A75" reason="title">
|
||
Type material of
|
||
<taxonomicName id="BAC3CD25E8D3421C778FC9A1FE96D6A1" baseAuthorityName="Regimbart" baseAuthorityYear="1877" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="baeri">
|
||
<emphasis id="098FDD49EB3121EDF90E407F7898BBC4" italics="true">Sandracottus baeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="875A54E787B9C8DAA314A1DAB035C530">
|
||
<materialsCitation id="AB1FA8C058977B4EE08F367AC36C4184" collectionCode="MNHN" collectorName="L. Hendrich" determinerName="M. Brancucci" location="Manille Baer" specimenCount="1" stateProvince="Manille Baer" typeStatus="lectotype">
|
||
<emphasis id="342649D0E9AEE2B3CBDCEFA1047FC172" bold="true" italics="true">
|
||
<typeStatus id="E9F14E8C108F6B9429D2C0F57CEA5387" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
(herewith designated): Male: “
|
||
<collectingRegion id="D09C17FEF4A93EED59A3A653877D429C">Manille Baer</collectingRegion>
|
||
”, “ Typus ”, “ Type ”, “
|
||
<taxonomicName id="6924A1675DF5102CAF4DCA9E9969DFD1" authority="Reg." authorityName="Regimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="baeri">
|
||
<originalDetermination id="B894E9A5F2CBB3671D22319FD418697C">
|
||
<emphasis id="94DFCA8C88E3DC51CEC6BF830E1DF4C2" italics="true">Sandracottus baeri</emphasis>
|
||
Rég.
|
||
</originalDetermination>
|
||
</taxonomicName>
|
||
”, “ Lectotypus
|
||
<originalDetermination id="9C3843F048849885A7EFC6B982F7EC13">
|
||
<taxonomicName id="0EB76E356E86FEB8133BD617DC0DDA6F" authority="Brancucci & Hendrich" authorityName="Brancucci & Hendrich" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="baeri">
|
||
<emphasis id="2ED21A8DCE8524B22D5535B799AA2081" italics="true">Hydaticus baeri</emphasis>
|
||
Brancucci & Hendrich
|
||
</taxonomicName>
|
||
07
|
||
</originalDetermination>
|
||
” [red printed label], “
|
||
<taxonomicName id="DC49157580CCCFFE8665A515872884CE" authority="(Wehncke)" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<originalDetermination id="4F20E43F58DE8EA8E29DE30E8DB866D6">
|
||
<emphasis id="2EC03B6CD03E66BA5601B6E6E9C14946" italics="true">Sandracottus insignis</emphasis>
|
||
(Wehncke)
|
||
</originalDetermination>
|
||
</taxonomicName>
|
||
det.
|
||
<determinerName id="C9989BF05441FD52563C81201299F3DA">M. Brancucci</determinerName>
|
||
&
|
||
<collectorName id="BBC6AFDB01683CE9DACC5848F75C4AF3">L. Hendrich</collectorName>
|
||
07 ” [white printed label] (
|
||
<abbrev id="ABBRID0ETRBK" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="6288ADD104A2042B709FB3769BBFA07F" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="80388C4C08B5FEB64438FDEF80A067F5" collectionCode="MNHN" determinerName="M. Brancucci & L. Hendrich" location="Manille Baer" specimenCount="2" specimenCount-male="2" stateProvince="Manille Baer" typeStatus="paralectotype">
|
||
<emphasis id="FAD8D9503DCE2F294E50CE35807F8D89" bold="true" italics="true">
|
||
<typeStatus id="BFEB8D6D6E6FEFB8A450011D52A01306" type="paralectotype">Paralectotypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="86F8E42E5E7D9FAF8210D2742B03A92B" count="2" type="male">2 males</specimenCount>
|
||
, “
|
||
<collectingRegion id="22E44E51CEFC3F01DE3AD9290F52192F">Manille Baer</collectingRegion>
|
||
”, “ Typus ”, “ Type ”, “
|
||
<taxonomicName id="459E968BE6BA3E981A0272EB28D687B8" authority="Reg." authorityName="Regimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="baeri">
|
||
<originalDetermination id="59D2044339F83B9977BCF0523BA96692">
|
||
<emphasis id="D420F4626A044BA7381BB7FE59E0376C" italics="true">Sandracottus baeri</emphasis>
|
||
Rég.
|
||
</originalDetermination>
|
||
</taxonomicName>
|
||
”, “ Paralectotypus
|
||
<originalDetermination id="03A0A1D372BD3B46498A64B76AA34A8C">
|
||
<taxonomicName id="6EEE6FBF54E22E36D2A96F0AC2523DD8" authority="Brancucci & Hendrich" authorityName="Brancucci & Hendrich" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="baeri">
|
||
<emphasis id="6286046AAC438FD4D6DB4948125630A0" italics="true">Hydaticus baeri</emphasis>
|
||
Brancucci & Hendrich
|
||
</taxonomicName>
|
||
07
|
||
</originalDetermination>
|
||
” [red printed label], “
|
||
<taxonomicName id="4E87D5B598FCB773B8B872DE1B2E1182" authority="(Wehncke)" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<originalDetermination id="ABB0E11FBF00D2AF107A59DBDE20E9B1">
|
||
<emphasis id="237E3B9CAC32305BFEFE46F1C7039945" italics="true">Sandracottus insignis</emphasis>
|
||
(Wehncke)
|
||
</originalDetermination>
|
||
</taxonomicName>
|
||
det.
|
||
<determinerName id="8258379AB5F33368A0CDF6B509CCF06A">M. Brancucci & L. Hendrich</determinerName>
|
||
07 ” [white printed label] (
|
||
<abbrev id="ABBRID0EYSBK" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="5E72CCAC65B9C7AF981DF8D69D716FAB" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
)
|
||
</materialsCitation>
|
||
. Examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E3SBK" type="type material">
|
||
<paragraph id="84AB501D02BBC850481BB4DBF5B959DD">
|
||
<heading id="6809BC2F8C960309B5767C6502286454" reason="title">
|
||
Type material of
|
||
<taxonomicName id="8239E56C8F654B01A3C1BF506C20F663" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="724EEA727074DDD7498C58262F7E3ED3" italics="true">Sandracottus ornatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="650FFC7AD143DB534EB69F7A0D185AF6">
|
||
<materialsCitation id="A2F82BA5205D9D38762A9287101EC5D5" collectionCode="NHMUK" collectorName="Sharp" determinerName="Hendrich" location="Borneo" municipality="Borneo" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<emphasis id="93C1F1ADBBEBCB6335957B527601442C" bold="true" italics="true">
|
||
<typeStatus id="859B138330690AB1E66B1AED3CBE145A" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="AB5F952CBB0F97F21358C8E7879ADF4C" type="female">Female</specimenCount>
|
||
: “
|
||
<collectingMunicipality id="3DCB43B9E91D6025C18DAEEF8B24B4F7">Borneo</collectingMunicipality>
|
||
986
|
||
<taxonomicName id="0581B9A5C03673346BDBBD1A63B282ED" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="6E377FC287801844DDECA24485616336" italics="true">ornatus</emphasis>
|
||
</taxonomicName>
|
||
”, “ Type H. T. ”, “
|
||
<collectorName id="64C05547E965256F7FF7DAB8A9B438E7">Sharp</collectorName>
|
||
coll. 1905-313 ”, “
|
||
<taxonomicName id="66BDB6C3C0EB4D13AF7CB005EE5D9FC7" authority="(Wehncke) Hendrich" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<originalDetermination id="476943FABDD50E54A8BB10A31589213E">
|
||
<emphasis id="BBEB2856DA8F2543FE74A9E0E2E387BA" italics="true">Sandracottus insignis</emphasis>
|
||
(Wehncke)
|
||
</originalDetermination>
|
||
<determinerName id="5D52E7DB90AB9D4BDE2BF2075121269E">Hendrich</determinerName>
|
||
</taxonomicName>
|
||
det. 2008 ” (
|
||
<abbrev id="ABBRID0EGUBK" xlink_title="Natural History Museum, London">
|
||
<collectionCode id="7CA3A11141808085B1F1E54139599BDB" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</abbrev>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="24327B12D2D8F9456DCA23EB0F3CDB79" collectionCode="NHMUK" determinerName="Hendrich" location="Borneo" municipality="Borneo" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<emphasis id="0FD74CFCC993E54D9C42E95C6E4F54E0" bold="true" italics="true">
|
||
<typeStatus id="E38640015932CE0DF1D81FB17A5BCECA" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="BFA2D55C9163C8230F143AF105596C07" type="female">Female</specimenCount>
|
||
: “
|
||
<collectingMunicipality id="2C5D0A1FB0C5589599CB23FA1CEBAED6">Borneo</collectingMunicipality>
|
||
986 1905-313 ”, “
|
||
<taxonomicName id="82967DBAC6501193D74E354D03FE1A92" authority="(Wehncke) Hendrich" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<originalDetermination id="1F06204968D555C83A2FD3C75A75F4B4">
|
||
<emphasis id="46D15E5915E06EF49B97B97F38C7E7B9" italics="true">Sandracottus insignis</emphasis>
|
||
(Wehncke)
|
||
</originalDetermination>
|
||
<determinerName id="040F8851A2AB6C482307A1CBD5E90103">Hendrich</determinerName>
|
||
</taxonomicName>
|
||
det. 2008 ” (
|
||
<abbrev id="ABBRID0EXUBK" xlink_title="Natural History Museum, London">
|
||
<collectionCode id="F00B165B9352983B7702ACE95BD57EC2" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</abbrev>
|
||
)
|
||
</materialsCitation>
|
||
. Examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2UBK" type="remarks">
|
||
<paragraph id="AFDEB8D3D18D0EE73034A87628002700">
|
||
<heading id="9AD22FD5A4D9FE8EE683A5AEC4A2FA06" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="721D00B2A2784501CE7EA3AAF614AEAC">
|
||
The type locality of
|
||
<taxonomicName id="C0EB0D8A120B284263E93ADB6247BAF9" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="E2FFCCB97AC4BF6CCDFC73500F13391E" italics="true">S. ornatus</emphasis>
|
||
</taxonomicName>
|
||
, “ Borneo ”, needs to be confirmed.
|
||
<taxonomicName id="627C43D3CFFD24D94F979A509FF574B0" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="70BFF0D303FA85C760840AA29A7C464E" italics="true">Sandracottus insignis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="2201CD0F50E18C5A0BD886351715C1D6" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="210A18B239F513BACAFFC0EA6D99CD61" italics="true">ornatus</emphasis>
|
||
</taxonomicName>
|
||
has never been reported from Borneo again. However, it cannot be ruled out that the species occurs there, as some dytiscids that occur in the south of the
|
||
<collectingCountry id="7CB86BA5EC03EB7CB5CE32ACC07E2248" name="Philippines">Philippines</collectingCountry>
|
||
have also been reported from the north of Borneo. From what is known so far,
|
||
<taxonomicName id="C6F3BE29BB99C296732B4ECF1B98D627" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="958638F76215EFD877F6CA225CA64C67" italics="true">S. insignis</emphasis>
|
||
</taxonomicName>
|
||
appears to be endemic to the
|
||
<collectingCountry id="CD3088754666485DFD33AE922CE61B13" name="Philippines">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EMWBK" type="material">
|
||
<paragraph id="AF66AF474B25BA08CD5381AE188265FA">
|
||
<heading id="9083840AA3AA2CF750FC80D3B11F3AF7" reason="title">Additional material.</heading>
|
||
</paragraph>
|
||
<paragraph id="A0D7C30123FA50F49C169431B2F612E3">
|
||
<emphasis id="1586864248BF955D38B0EC916DA803C4" bold="true">
|
||
(
|
||
<specimenCount id="17480EFA65EDC009EDBB9E23AD0BE9C9" count="10" type="generic">10 specimens</specimenCount>
|
||
)
|
||
</emphasis>
|
||
: •
|
||
<emphasis id="7BA5A77B44ADE26C58336E1FD4543DC4" bold="true">
|
||
<collectingCountry id="CCDA8D0EF0D34044A003CE68C8189216" name="Philippines">Philippines</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="07F7695D37C1C82EE497F2D6996CF6C7" count="1" type="generic">1 ex.</specimenCount>
|
||
, “ Balabac ” [handwritten label],
|
||
<taxonomicName id="4D4683797C50F519A0DB5695AD7B11BE" authority="Shrp" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ornatus">
|
||
“
|
||
<emphasis id="515080ED935026F78375228F4EB4461E" italics="true">ornatus</emphasis>
|
||
Shrp
|
||
</taxonomicName>
|
||
” [handwritten label] (
|
||
<abbrev id="ABBRID0E6WBK" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="407D06A6B19DA326135CE7AF74D209FA" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="A827EF722450371F0D0EE5F5345F9F3F" count="1" type="generic">1 ex.</specimenCount>
|
||
, “ Montalban ” [Luzon], “ Nr. 6 Fld. - H. ” [printed label], “ baeri Régb. Insignis Sharp ” [handwritten label by Régimbart] (
|
||
<abbrev id="ABBRID0EDXBK" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="BEB955B06B9229238A9DC1169034181E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
),
|
||
<specimenCount id="EA87F9399FBFE1D7B252F92284AC3A9D" count="2" type="generic">2 exs.</specimenCount>
|
||
, “ Mindanao ” (
|
||
<abbrev id="ABBRID0EHXBK" xlink_title="Muséum national d´Histoire naturelle, Paris">
|
||
<collectionCode id="69CF571E76AD23969615D9144842B7C4" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="700D9F00C38AEE908F8408B704196C25" count="2" type="generic">2 exs.</specimenCount>
|
||
, “
|
||
<collectingCounty id="2D825F863D7C9C7FCD2C46021B47BF42">Luzon</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="3BA10FDF1E2E00884F198D84A19F6A04">Los Banos</collectingMunicipality>
|
||
,
|
||
<location id="C72EBC3DA58B27E4B35F0064792C9F89" LSID="urn:lsid:plazi:treatment:1113CD6291B65D00858C62DFB6C9FB47:C72EBC3DA58B27E4B35F0064792C9F89" country="Philippines" county="Luzon" latitude="10.833333" longLatPrecision="1297" longitude="119.5" municipality="Los Banos" name="Mt. Makiling" stateProvince="Pampanga">Mt. Makiling</location>
|
||
[above the Botanical Garden,
|
||
<quantity id="33AE5A06747E978C6E544950630E4186" metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">
|
||
<elevation id="1F5AD7F81C3A15DF3583384B0F182270" metricMagnitude="2" metricUnit="m" metricValue="1.8" unit="m" value="180.0">180 m</elevation>
|
||
</quantity>
|
||
] (3),
|
||
<date id="97FE0E783BAA20B680AE71E1F74A714A" value="1992-11-13">
|
||
<collectingDate id="F5FAA138D1DE0250940B9B3FB00AB662" value="1992-11-13">13. XI. 1992</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FDAA37D38C6F28205B49D70308EC69B3">H. Schillhammer</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0ELXBK" xlink_title="Naturhistorisches Museum Wien">
|
||
<collectionCode id="DFA5C8CE92CBFBD9297EDAC5B8F72470" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="36A61C4A4AD784613F829FE936765F0F" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<location id="33F9CBF1DBF8C3C277A1B0AA23DBB0E0" LSID="urn:lsid:plazi:treatment:1113CD6291B65D00858C62DFB6C9FB47:33F9CBF1DBF8C3C277A1B0AA23DBB0E0" country="Philippines" county="Luzon" latitude="10.833333" longLatPrecision="1297" longitude="119.5" municipality="Los Banos" name="Luzon" stateProvince="Pampanga">Luzon</location>
|
||
,
|
||
<collectingRegion id="5A90DCAA39774319F8F1887B074373FE" country="Philippines" name="Pampanga">Pampanga Province</collectingRegion>
|
||
,
|
||
<location id="6E64B3277796ECD02B2BE08BB7326A27" LSID="urn:lsid:plazi:treatment:1113CD6291B65D00858C62DFB6C9FB47:6E64B3277796ECD02B2BE08BB7326A27" country="Philippines" county="Luzon" latitude="10.833333" longLatPrecision="1297" longitude="119.5" municipality="Los Banos" name="Bano" stateProvince="Pampanga">Bano</location>
|
||
<collectingRegion id="75F7752538C2F17648A50AFE3CB824AD" country="Philippines" name="San Juan">San Juan</collectingRegion>
|
||
,
|
||
<collectorName id="8446B7A6171C638B22DD2BC91A996654">
|
||
<location id="CEE561EDC8C1B1AC9DB9E63ACC9B8FC1" LSID="urn:lsid:plazi:treatment:1113CD6291B65D00858C62DFB6C9FB47:CEE561EDC8C1B1AC9DB9E63ACC9B8FC1" country="Philippines" county="Luzon" latitude="10.833333" longLatPrecision="1297" longitude="119.5" municipality="Los Banos" name="Mt. Arayat" stateProvince="Pampanga">Mt. Arayat</location>
|
||
NP
|
||
</collectorName>
|
||
, spring creek, bolders, root packs, riffle,
|
||
<quantity id="B8EAB24AC727E2537FA6888F80E81FEF" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">
|
||
<elevation id="92F08F7C8974727C3D4A160D33AFC981" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">37 m</elevation>
|
||
</quantity>
|
||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESXBK" specific-use="{"type":"Point","coordinates":[120.763272,15.172672]}">
|
||
<geoCoordinate id="87C574DC5B3CA8DDF8DC777559EE2261" degrees="15" direction="north" minutes="10" orientation="latitude" precision="1" seconds="21.62" value="15.172672">15 ° 10 ' 21.62 " N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C3D929288FD110B3F7D740EECBDEE407" degrees="120" direction="east" minutes="45" orientation="longitude" precision="1" seconds="47.78" value="120.763275">120 ° 45 ' 47.78 " E</geoCoordinate>
|
||
</named-content>
|
||
,
|
||
<date id="AD3854F9A5C2AED104D73D0508D58DA4" value="2009-10-31">
|
||
<collectingDate id="1A0B21A4E6A6DCEE378CE283508FFB3D" value="2009-10-31">31. X. 2009</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="4FE5C41CF844793FB264B03463FA48FC">H. Freitag</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0EXXBK" xlink_title="Zoologische Staatssammlung, München">
|
||
<collectionCode id="1A1C00976407802611D2CC461BB5A233" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="E39FD54CC2F79EE92329114AD46F6B21" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectorName id="9108A8D1DCEBB8AA947608114253FF65">Polillo</collectorName>
|
||
[Polillo Islands,
|
||
<collectorName id="63B934A0A9865C7FBC9B2007225E29E2">Luzon</collectorName>
|
||
],
|
||
<collectorName id="D8C5EC49DE016558898B2F3442B6448F">Taylor</collectorName>
|
||
” (
|
||
<abbrev id="ABBRID0E2XBK" xlink_title="Zoologische Staatssammlung, München">
|
||
<collectionCode id="B6192F2A2FD279F1CB4F7F50BE48C6F0" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
|
||
</abbrev>
|
||
);
|
||
<specimenCount id="34DB9933C82BDF15EB932DA7C7E77DCF" count="1" type="generic">1 ex.</specimenCount>
|
||
, “
|
||
<collectingRegion id="F8E288F8C903D2EBC4DDF78BCF53CA23" country="Philippines" name="Palawan">Palawan</collectingRegion>
|
||
,
|
||
<collectorName id="9E5F71872B45437788191EB6D7B01A81">Taytay</collectorName>
|
||
[
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECYBK" specific-use="{"type":"Point","coordinates":[119.500000,10.833333]}">
|
||
<geoCoordinate id="4801C48A088C2B4276B7C07023C3434C" degrees="10" direction="north" minutes="50" orientation="latitude" precision="925" value="10.833333">10 ° 50 ' N</geoCoordinate>
|
||
<geoCoordinate id="39E3E871E6A5C919B4DE13599B281AD8" degrees="119" direction="east" minutes="30" orientation="longitude" precision="925" value="119.5">119 ° 30 ' E</geoCoordinate>
|
||
</named-content>
|
||
],
|
||
<collectorName id="54407C862A4AF36478E53555F37CCDE6">W. Schultze</collectorName>
|
||
leg. ” (
|
||
<abbrev id="ABBRID0EHYBK" xlink_title="Staatliches Museum für Tierkunde">
|
||
<collectionCode id="268E4EF83A3337F9D5FC25CF19D2852F">SMTD</collectionCode>
|
||
</abbrev>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELYBK" type="redescription">
|
||
<paragraph id="607EE566ABB39019F343EEC8F775180B">
|
||
<heading id="C1EB6C6C67C7717C3DA143A00854243D" reason="title">Redescription.</heading>
|
||
</paragraph>
|
||
<paragraph id="FF78348AADB7F27F9554513425225481">Body broadly oval, shiny, testaceous with black markings. Ventral side completely dark brown to black, fore and mid legs testaceous, hind legs ferrugineus brown.</paragraph>
|
||
<paragraph id="D9202701A3EB294987878EA67CC53C4A">
|
||
Head testaceous with posterior part dark brown as well as two dark brown, oblique, elongated patches on frons (Figs
|
||
<figureCitation id="0ADC457A395B5A96D14484FC59FABE32" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8–11. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">8</figureCitation>
|
||
,
|
||
<figureCitation id="A2F5954D260A8FEF3B57FE70B2A9E0AE" captionStart="Figures 54–57" captionStartId="F20" captionText="Figures 54–57. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">55</figureCitation>
|
||
). Surface finely microreticulate, particularly on anterior half, just shagreened on rest of surface. Punctation consisting of dense and very small punctures and of larger, much sparser ones, the latter particularly numerous on frons. Clypeal grooves, punctures alongside eyes and transverse depression beside eyes distinctly impressed, punctures large and coalescent. Antennae testaceous; antennomeres slender, antennomere V 4 × as long as broad.
|
||
</paragraph>
|
||
<paragraph id="400E403C174A57AD2ABFFCC62F013D4E">
|
||
Pronotum testaceous with long and broad dark brown band posteriorly and narrow dark brown band anteriorly (Figs
|
||
<figureCitation id="461C4847DDEE16C40B7772ABD53EB272" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8–11. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">8</figureCitation>
|
||
,
|
||
<figureCitation id="FC3C61B5FAF52A8A121E216643C12DC1" captionStart="Figures 54–57" captionStartId="F20" captionText="Figures 54–57. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">55</figureCitation>
|
||
). Surface sculpture shagreened with dense punctation; punctures medium-sized. Anterior rows of punctures with dense, strong, and coalescent punctures, shortly interrupted in middle. Lateral puncture lines broad; punctures medium-sized and coalescent. Posterior row limited to few punctures at latero-basal quarter.
|
||
</paragraph>
|
||
<paragraph id="E632C6965D392F6523737F7F889FAED0">
|
||
Elytra testaceous with black markings leaving four angulous basal testaceous spots, two large postmedian as well as a preapical and an apical testaceous patch (Figs
|
||
<figureCitation id="0109DA5786CB5522316EAFDBB1677CF5" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8–11. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">8</figureCitation>
|
||
,
|
||
<figureCitation id="8BFCDE26C4A43A783D111254A63B8CB2" captionStart="Figures 54–57" captionStartId="F20" captionText="Figures 54–57. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">55</figureCitation>
|
||
). Epipleura testaceous. Surface microreticulate, meshes very small but strongly impressed. Punctation consisting of numerous minute punctures mostly in intersection of meshes and of large sparse punctures regularly spread on whole surface. Puncture lines with groups of medium-sized punctures, discal row almost complete. Sutural puncture line with few small punctures along suture.
|
||
</paragraph>
|
||
<paragraph id="8AF245BF8B228D2F99733B87560D2524">Ventral side dark brown. Legs particularly fore and mid legs testaceous, hind legs ferrugineus brown to dark brown. Prosternal process very short and broad, only 1.4 × longer than broad, flattened and superficially sculptured; posterior border broadly rounded. Metacoxa very superficially shagreened and finely punctured; punctures small and sparse; Metatibia with numerous medium-sized punctures on whole outer half, somewhat larger proximally. Ventrites II – VI shagreened, densely covered with very small punctures and larger sparser ones. Posterior margins rounded, bordered with short row of coalescent punctures each side of middle.</paragraph>
|
||
<paragraph id="222A4DB1BD3428B403273E3CB033EB6E">
|
||
Measurements:
|
||
<abbrev id="ABBRID0ERZBK" xlink_title="total length">TL</abbrev>
|
||
=
|
||
<quantity id="3EA74AE9293F49F36AC1601A4288CB01" metricMagnitude="-2" metricUnit="m" metricValue="1.28" metricValueMax="1.31" metricValueMin="1.25" unit="mm" value="12.8" valueMax="13.1" valueMin="12.5">12.5–13.1 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EVZBK" xlink_title="total length minus head length">TL-h</abbrev>
|
||
=
|
||
<quantity id="1523988EB87B6C8CADD85D86515355F4" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.22" metricValueMin="1.14" unit="mm" value="11.8" valueMax="12.2" valueMin="11.4">11.4–12.2 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EZZBK" xlink_title="maximum width">TW</abbrev>
|
||
=
|
||
<quantity id="CF47C830B8F9933A5575383563F5F6EC" metricMagnitude="-3" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.1" metricValueMin="7.8" unit="mm" value="7.949999999999999" valueMax="8.1" valueMin="7.8">7.8–8.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4A51525D5A5E237C92A7D6AAC30DC4F5">
|
||
<specimenCount id="E27D1524560D1B0AE534D793137C20EE" count="1" type="male">♂</specimenCount>
|
||
. Protarsomeres I – III strongly enlarged with three larger suckers and numerous smaller ones. Mesotarsomeres I – III with two rows of small suckers. Median lobe of aedeagus, in ventral view, broad, apical half more or less parallel-sided, each part narrowly rounded apically (Fig.
|
||
<figureCitation id="FE25F4847ED0EC4DFFDEFCD7DE385762" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">19 a</figureCitation>
|
||
). Parameres also narrow, slightly longer than median lobe, and gently tapered on apical half (Fig.
|
||
<figureCitation id="4250C6A29F79356C43B38A0C7DE76932" captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16–19. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 16 Sandracottus festivus 17 S. femoralis 18 S. hunteri and 19 S. insignis." figureDoi="10.3897/zookeys.1223.138220.figures16-19" httpUri="https://binary.pensoft.net/fig/1214412">19 b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BD389036E7167F1107916A1CBF7A119D">
|
||
<specimenCount id="A6F570C8701070052FE6BD71DBA60A35" count="1" type="female">♀</specimenCount>
|
||
. Similar to male. Tarsi not enlarged. Microsculpture on ventrite VI as in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EI1BK" type="Differential diagnosis">
|
||
<paragraph id="47BA71DD5CB3029B8EC76FBA81870E3C">
|
||
<heading id="D8DDA6CE627F3393A9B7566D4EBF2F3C" reason="title">Differential diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="5FF1AD21EFE32A28DC43714CB0599A44">
|
||
The species is clearly distinguished from most other Oriental species of the genus by its characteristic colouration. From the dorsal colouration
|
||
<taxonomicName id="46507A09463ABC76C39BE35947C36EC3" authorityName="Hendrich" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="insignis">
|
||
<emphasis id="82474DB4CCCA4860C24D75464297C3C5" italics="true">S. insignis</emphasis>
|
||
</taxonomicName>
|
||
is near to the Indian
|
||
<taxonomicName id="61EF5884C56F042C117981AD889B6AA1" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="festivus">
|
||
<emphasis id="6FF88D6EA7323133BB6F938E0811BB3B" italics="true">S. festivus</emphasis>
|
||
</taxonomicName>
|
||
which is generally larger and more elongated. Furthermore, both species can be separated by the shapes of their median lobes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EE2BK" type="distribution">
|
||
<paragraph id="C8ED49C44A1CBAF1EACA3992313A5F36">
|
||
<heading id="444E372D8717BA08FA96EAF00220F88E" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="70157CAD9C5FD7F1D406966932EA2462">
|
||
<collectingCountry id="137E740D637D4C10D4AD3E8A94A1FF01" name="Philippines">Philippines</collectingCountry>
|
||
:
|
||
<collectingRegion id="EFBA704071830EB7BBDCE99B1F1463E5" country="Philippines" name="Palawan">Palawan</collectingRegion>
|
||
, Mindanao, Luzon. The record from Borneo seems to be doubtful (Fig.
|
||
<figureCitation id="924A0A3868E9B8A74E5D102371498BBF" captionStart="Figure 28" captionStartId="F12" captionText="Figure 28. Distribution of Sandracottus insignis in the Philippines." figureDoi="10.3897/zookeys.1223.138220.figure28" httpUri="https://binary.pensoft.net/fig/1214419">28</figureCitation>
|
||
). Specimens were collected between 40 and
|
||
<quantity id="3718077F2A19C0CCB6472D7DB781D6DC" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
|
||
above sea level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EO2BK" type="habitat">
|
||
<paragraph id="2DA651ECBEA6FF0CEDA72559257CB18F">
|
||
<heading id="82E843D298FD2F83B81DDE461339B536" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="15A52DA827B89BBDF99EE71BDD8DF6F3">
|
||
The
|
||
<specimenCount id="702218739214B6D215F349D2BAE3625F" count="3" type="generic">three specimens</specimenCount>
|
||
at Mt. Makiling were collected in an exposed roadside ditch surrounded by mature second growth lowland rainforest (H. Schillhammer pers. comm. 2005). The single specimen from Bano
|
||
<collectingRegion id="0A50F61F820B81BB66ABAC21BE778591" country="Philippines" name="San Juan">San Juan</collectingRegion>
|
||
at Mt. Arayat NP was collected among root packs in a shallow bay of a slow flowing creek, partly shaded by second growth vegetation (Freitag pers. comm. 2012) (Fig.
|
||
<figureCitation id="41AAAF84F1C853782247650D74375B17" captionStart="Figures 37–42" captionStartId="F16" captionText="Figures 37–42. Habitats of Sandracottus 37, 38 Laos, Attapeu prov., Annam Highlands Mts., Dong Amphan NBCA, Nong Fa [puddles near lake shore] and lake, habitat of S. hunteri 39 Laos, Katamtok, rest pools along a rocky river bed, habitat of S. maculatus (photos: J. Hájek) 40 Thailand, near Salawin NP, open and muddy puddle made by elephants (photo: J. Šťastný), and 41 Sakaerat Biosphere Reserve, rocky pool in an otherwise dry stream bed, habitats of S. hunteri and S. maculatus (photo: W. Atthakor) 42 Philippines, Luzon, Pampanga Prov., Bano San Juan, Mt. Arayat NP, spring creek, habitat of S. insignis (photo: H. Freitag)." figureDoi="10.3897/zookeys.1223.138220.figures37-42" httpUri="https://binary.pensoft.net/fig/1214423">42</figureCitation>
|
||
). The larvae are still unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EY2BK" type="Conservation">
|
||
<paragraph id="258834BD8E6BF4E2FA60221AA8C182F5">
|
||
<heading id="2D40ED42C5AD57DD4B983684FC88571B" reason="title">Conservation.</heading>
|
||
</paragraph>
|
||
<paragraph id="7965233AE0804060BC6912DE729DCE2C">
|
||
A rare species, probably associated with the declining primary lowland rainforests in the
|
||
<collectingCountry id="4A2879387D2ADB2F71CB9FBA0F2B8925" name="Philippines">Philippines</collectingCountry>
|
||
(see Indomalayan species
|
||
<taxonomicName id="D1831DF74703F655B04DDC621455EBAC" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bizonatus">
|
||
<emphasis id="B2FB30190D200B3126C6EEC67260DF83" italics="true">S. bizonatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3F6E973B2046009C19C94341199EB217" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
|
||
<emphasis id="8AC00C7179EE90D1097709CF798C839B" italics="true">S. femoralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C00B64F0C1A043C9C3EAAADD243EF300" authorityName="Hendrich & Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
|
||
<emphasis id="4B68E87DBA06521B61021DE4B5ECE42E" italics="true">S. maculatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B3882DEAFB48F14257AE56C7EA696A68" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">
|
||
<emphasis id="42C25BF3BFE9ACB94481C0C71EECC992" italics="true">S. rotundus</emphasis>
|
||
</taxonomicName>
|
||
). According to the present knowledge it is an endangered species (
|
||
<bibRefCitation id="1CDE9E708966FAE41D66412287253359" DOI="10.1080/01650424.2016.1210814" author="Freitag" etAl="et al." firstAuthor="Freitag" issue="3" journalOrPublisher="Aquatic Insects" pagination="177-213" refId="B13" refString="Freitag H, Jäch MA, Wewalka G (2016) Diversity of aquatic and riparian Coleoptera of the Philippines: checklist, state of knowledge, priorities for future research and conservation. Aquatic Insects 37 (3): 177–213. https://doi.org/10.1080/01650424.2016.1210814" title="Diversity of aquatic and riparian Coleoptera of the Philippines: checklist, state of knowledge, priorities for future research and conservation." volume="37" year="2016">Freitag et al. 2016</bibRefCitation>
|
||
). It is recommended to be listed in the next IUCN red list.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |