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<document id="DAE2FCBF52DD08053734272402F6A6C4" ID-CLB-Dataset="46496" ID-DOI="10.11646/zootaxa.3764.2.1" ID-GBIF-Dataset="7ec156cd-68b7-45e9-804a-f17481ae1b72" ID-ISSN="1175-5326" ID-Zenodo-Dep="227668" ID-ZooBank="707FAA19-12B5-4870-A22C-0859A058A73A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460380742660" checkinUser="plazi" docAuthor="Papp, László" docDate="2014" docId="061487A2FFDFD14AFF58F90BFB94FADC" docLanguage="en" docName="zt03764p130.pdf" docOrigin="Zootaxa 3764 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Biphallapodema Papp, 2014, gen. n." docType="treatment" docVersion="9" lastPageNumber="107" masterDocId="FA2DFFDAFFDBD14CFFCFFFE9FFE7FFD3" masterDocTitle="New genera of Afrotropical limosinine sphaerocerids (Diptera: Sphaeroceridae)" masterLastPageNumber="130" masterPageNumber="101" pageNumber="105" updateTime="1698296837072" updateUser="ExternalLinkService">
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<mods:title id="2945067A76D4F924C896BF83688615C7">New genera of Afrotropical limosinine sphaerocerids (Diptera: Sphaeroceridae)</mods:title>
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<mods:namePart id="5A29CAAE53EE77FADD5CA4A41B351BBA">Papp, László</mods:namePart>
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<mods:title id="9ABEFBC7F803257A17ACD9A5A8707834">Zootaxa</mods:title>
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<mods:date id="230EE4A80B3063965150E73A030F5174">2014</mods:date>
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<mods:number id="0E7E06B086F67B813E971897A00D535F">3764</mods:number>
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<mods:identifier id="F7896DB73A9BBAD37ED4E57E84782319" type="DOI">10.11646/zootaxa.3764.2.1</mods:identifier>
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<treatment id="061487A2FFDFD14AFF58F90BFB94FADC" ID-DOI="http://doi.org/10.5281/zenodo.6125147" ID-GBIF-Taxon="119414142" ID-Zenodo-Dep="6125147" LSID="urn:lsid:plazi:treatment:061487A2FFDFD14AFF58F90BFB94FADC" httpUri="http://treatment.plazi.org/id/061487A2FFDFD14AFF58F90BFB94FADC" lastPageId="6" lastPageNumber="107" pageId="4" pageNumber="105">
<subSubSection id="C6A7653FFFDFD148FF58F90BFEC0F8CC" pageId="4" pageNumber="105" type="nomenclature">
<paragraph id="8E0236B4FFDFD148FF58F90BFE4CF928" blockId="4.[151,427,1762,1823]" box="[151,427,1762,1787]" pageId="4" pageNumber="105">
<heading id="D54A81D8FFDFD148FF58F90BFE4CF928" bold="true" box="[151,427,1762,1787]" fontSize="10" level="2" pageId="4" pageNumber="105" reason="2">
<emphasis id="BCC9EAA6FFDFD148FF58F90BFE4CF928" bold="true" box="[151,427,1762,1787]" pageId="4" pageNumber="105">
<taxonomicName id="49BD4D37FFDFD148FF58F90BFEB2F928" box="[151,341,1762,1787]" class="Insecta" family="Sphaeroceridae" genus="Biphallapodema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BCC9EAA6FFDFD148FF58F90BFEB2F928" bold="true" box="[151,341,1762,1787]" italics="true" pageId="4" pageNumber="105">Biphallapodema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7FA57DDFFDFD148FE93F90BFE4CF928" box="[348,427,1762,1787]" pageId="4" pageNumber="105" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8E0236B4FFDFD148FF58F8EFFEC0F8CC" blockId="4.[151,427,1762,1823]" box="[151,295,1798,1823]" pageId="4" pageNumber="105">
(
<figureCitation id="16862A31FFDFD148FF50F8EFFEF8F8CC" box="[159,287,1798,1823]" captionStart-0="FIGURES 13 – 20" captionStart-1="FIGURES 21 – 27" captionStart-2="FIGURES 28 – 32" captionStart-3="FIGURES 33 – 38" captionStartId-0="5.[151,264,1921,1943]" captionStartId-1="7.[151,264,1728,1750]" captionStartId-2="8.[151,264,1678,1700]" captionStartId-3="9.[151,264,1764,1786]" captionTargetBox-0="[324,1260,545,1903]" captionTargetBox-1="[329,1258,596,1701]" captionTargetBox-2="[340,1253,560,1653]" captionTargetBox-3="[334,1256,595,1741]" captionTargetId-0="figure@5.[324,1264,541,1908]" captionTargetId-1="figure@7.[325,1261,591,1706]" captionTargetId-2="figure@8.[330,1256,553,1657]" captionTargetId-3="figure@9.[329,1257,591,1741]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURES 13 – 20. Female postabdomen and genitalia. 13 – 16 = Afrolimosina albitarsis sp. n. 13 = sternite 8, ventral view, 14 = tergite 8 with contours of sternite 8, lateral view, 15 = tergite 8, epiproct and cerci, dorsal view, 16 = spermathecae in glycerol with contours of one spermatheca in water. 17 – 20 = Biphallapodema oligodentata sp. n.. 17 = sternite 8, ventral view, 18 = hypoproct, ventral view, 19 = epiproct and cerci, dorsal view, 20 = spermathecae. Scale bar: Figs 13 – 20 = 0.1 mm." captionText-1="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." captionText-2="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." captionText-3="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri-0="https://zenodo.org/record/227671/files/figure.png" httpUri-1="https://zenodo.org/record/227672/files/figure.png" httpUri-2="https://zenodo.org/record/227673/files/figure.png" httpUri-3="https://zenodo.org/record/227674/files/figure.png" pageId="4" pageNumber="105">Figs 17–38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C6A7653FFFDFD148FF58F8A7FE8CF879" pageId="4" pageNumber="105" type="materials_examined">
<paragraph id="8E0236B4FFDFD148FF58F8A7FABEF8B7" blockId="4.[151,1369,1870,1892]" box="[151,1369,1870,1892]" pageId="4" pageNumber="105">
<emphasis id="BCC9EAA6FFDFD148FF58F8A7FEFCF8B7" bold="true" box="[151,283,1870,1892]" pageId="4" pageNumber="105">
<typeStatus id="51068816FFDFD148FF58F8A7FF2DF8B7" box="[151,202,1870,1892]" pageId="4" pageNumber="105">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="49BD4D37FFDFD148FEE5F8A7FDB7F8B7" box="[298,592,1870,1892]" class="Insecta" family="Sphaeroceridae" genus="Biphallapodema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="polydentata" status="sp. nov.">
<emphasis id="BCC9EAA6FFDFD148FEE5F8A7FDB7F8B7" box="[298,592,1870,1892]" italics="true" pageId="4" pageNumber="105">Biphallapodema polydentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7FA57DDFFDFD148FD9AF8A7FD41F8B7" box="[597,678,1870,1892]" pageId="4" pageNumber="105" rank="species">
<emphasis id="BCC9EAA6FFDFD148FD9AF8A7FD41F8B7" bold="true" box="[597,678,1870,1892]" pageId="4" pageNumber="105">sp. nov.</emphasis>
</taxonomicNameLabel>
, by present designation; additional species:
<taxonomicName id="49BD4D37FFDFD148FBA9F8A6FAE5F8B7" box="[1126,1282,1870,1892]" class="Insecta" family="Sphaeroceridae" genus="Biphallapodema" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="oligodentata" status="sp. nov.">
<emphasis id="BCC9EAA6FFDFD148FBA9F8A6FAE5F8B7" box="[1126,1282,1870,1892]" italics="true" pageId="4" pageNumber="105">B. oligodentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7FA57DDFFDFD148FAC6F8A7FABEF8B7" box="[1289,1369,1870,1892]" pageId="4" pageNumber="105" rank="species">
<emphasis id="BCC9EAA6FFDFD148FAC6F8A7FABEF8B7" bold="true" box="[1289,1369,1870,1892]" pageId="4" pageNumber="105">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8E0236B4FFDFD148FF58F878FE8CF879" blockId="4.[151,1436,1937,2034]" box="[151,363,1937,1962]" pageId="4" pageNumber="105">
<emphasis id="BCC9EAA6FFDFD148FF58F878FF16F879" bold="true" box="[151,241,1937,1962]" pageId="4" pageNumber="105">Gender</emphasis>
: feminine.
</paragraph>
</subSubSection>
<subSubSection id="C6A7653FFFDFD148FF08F85DFAFDF821" pageId="4" pageNumber="105" type="etymology">
<paragraph id="8E0236B4FFDFD148FF08F85DFAFDF821" blockId="4.[151,1436,1937,2034]" pageId="4" pageNumber="105">
<emphasis id="BCC9EAA6FFDFD148FF08F85DFEA9F81E" bold="true" box="[199,334,1972,1997]" pageId="4" pageNumber="105">Etymology.</emphasis>
The name of the new genus refers to the extremely long connections of postgonites to the phallapodeme, which look like a second shorter apodeme at a first glance under the true phallapodeme.
</paragraph>
</subSubSection>
<subSubSection id="C6A7653FFFDED149FF08FF7EFEABFE27" pageId="5" pageNumber="106" type="diagnosis">
<paragraph id="8E0236B4FFDED149FF08FF7EFEABFE27" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="106">
<emphasis id="BCC9EAA6FFDED149FF08FF7EFEA6FF63" bold="true" box="[199,321,151,176]" pageId="5" pageNumber="106">Diagnosis.</emphasis>
Small terricolous flies with peculiar male genitalia. Base of postgonite connected to phallapodeme by extremely long hairpin-like flexible black connections, in profile they look like a second phallapodeme. Epandrial complex strongly fused ventrally in the cercal part, most ventrally with a pair of strong caniniform processes, hypandrium strongly fused to epandrium, lateral parts (arms) asymmetrical, medial part short broad and left-side curved, basiphallus ventrally directed, hook-like in profile with an upcurved thinner epiphallus. Sclerotised parts of ducts of spermathecae short, each with a bulbous distal end; spermathecae are not dish-shaped. Body less sclerotised and not granulose. This new genus keys out to
<taxonomicName id="49BD4D37FFDED149FC2BFE86FB5FFE54" authority="L.Papp" authorityName="L.Papp" box="[996,1208,367,392]" class="Insecta" family="Sphaeroceridae" genus="Rohacekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="BCC9EAA6FFDED149FC2BFE86FBBDFE5B" box="[996,1114,367,392]" italics="true" pageId="5" pageNumber="106">Rohacekia</emphasis>
L.Papp
</taxonomicName>
(couplet 72) or to
<taxonomicName id="49BD4D37FFDED149FF58FE7DFED6FE7E" box="[151,305,404,429]" class="Insecta" family="Sphaeroceridae" genus="Spinilimosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="BCC9EAA6FFDED149FF58FE7DFED6FE7E" box="[151,305,404,429]" italics="true" pageId="5" pageNumber="106">Spinilimosina</emphasis>
</taxonomicName>
(couplet 77) in Papp’s (2008) key. The habitus and male genitalia are much different from
<taxonomicName id="49BD4D37FFDED149FAE9FE7DFA7BFE7E" box="[1318,1436,404,429]" class="Insecta" family="Sphaeroceridae" genus="Rohacekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="BCC9EAA6FFDED149FAE9FE7DFA7BFE7E" box="[1318,1436,404,429]" italics="true" pageId="5" pageNumber="106">Rohacekia</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EA2C4B45FFDED149FF08FE51FEA2FE03" author="Papp" box="[199,325,440,464]" pageId="5" pageNumber="106" refString="Papp, L. (2008) New genera of the Old World Limosininae (Diptera, Sphaeroceridae). Acta Zoologica Academiae Scientiarum Hungaricae, 54 (Suppl. 2), 47 - 209." type="journal article" year="2008">Papp 2008</bibRefCitation>
: figs 281–285). Contrary to
<taxonomicName id="49BD4D37FFDED149FD40FE5EFCCEFE03" box="[655,809,439,464]" class="Insecta" family="Sphaeroceridae" genus="Spinilimosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="BCC9EAA6FFDED149FD40FE5EFCCEFE03" box="[655,809,439,464]" italics="true" pageId="5" pageNumber="106">Spinilimosina</emphasis>
</taxonomicName>
the epandrium of the new genus does not bear robust spine-like setae.
</paragraph>
</subSubSection>
<caption id="DAC2663CFFDED149FF58F868FAF8F820" httpUri="https://zenodo.org/record/227671/files/figure.png" pageId="5" pageNumber="106" targetBox="[324,1260,545,1903]" targetPageId="5">
<paragraph id="8E0236B4FFDED149FF58F868FAF8F820" blockId="5.[151,1436,1921,2035]" pageId="5" pageNumber="106">
<emphasis id="BCC9EAA6FFDED149FF58F868FEB4F844" bold="true" box="[151,339,1921,1943]" pageId="5" pageNumber="106">FIGURES 13–20.</emphasis>
Female postabdomen and genitalia. 13–16 =
<taxonomicName id="49BD4D37FFDED149FCD2F868FBE1F844" box="[797,1030,1921,1943]" class="Insecta" family="Sphaeroceridae" genus="Afrolimosina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="albitarsis" status="sp. nov.">
<emphasis id="BCC9EAA6FFDED149FCD2F868FBE1F844" box="[797,1030,1921,1943]" italics="true" pageId="5" pageNumber="106">Afrolimosina albitarsis</emphasis>
</taxonomicName>
<emphasis id="BCC9EAA6FFDED149FBC3F868FBA3F844" bold="true" box="[1036,1092,1921,1943]" pageId="5" pageNumber="106">
<taxonomicNameLabel id="A7FA57DDFFDED149FBC3F868FBA3F844" box="[1036,1092,1921,1943]" pageId="5" pageNumber="106" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
13 = sternite 8, ventral view, 14 = tergite 8 with contours of sternite 8, lateral view, 15 = tergite 8, epiproct and cerci, dorsal view, 16 = spermathecae in glycerol with contours of one spermatheca in water. 17–20 =
<taxonomicName id="49BD4D37FFDED149FD75F857FC0EF807" box="[698,1001,1982,2004]" class="Insecta" family="Sphaeroceridae" genus="Biphallapodema" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="oligodentata" status="sp. nov.">
<emphasis id="BCC9EAA6FFDED149FD75F857FC0EF807" box="[698,1001,1982,2004]" italics="true" pageId="5" pageNumber="106">Biphallapodema oligodentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7FA57DDFFDED149FC3EF857FBCBF807" box="[1009,1068,1982,2004]" pageId="5" pageNumber="106" rank="species">
<emphasis id="BCC9EAA6FFDED149FC3EF857FBCBF807" bold="true" box="[1009,1068,1982,2004]" pageId="5" pageNumber="106">sp. n.</emphasis>
</taxonomicNameLabel>
. 17 = sternite 8, ventral view, 18 = hypoproct, ventral view, 19 = epiproct and cerci, dorsal view, 20 = spermathecae. Scale bar: Figs 13–20 = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C6A7653FFFDDD14AFF08FF7EFB94FADC" pageId="6" pageNumber="107" type="description">
<paragraph id="8E0236B4FFDDD14AFF08FF7EFD4EFECF" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FF7EFEBEFF63" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="107">Description.</emphasis>
<emphasis id="BCC9EAA6FFDDD14AFEADFF7EFE78FF63" box="[354,415,151,176]" italics="true" pageId="6" pageNumber="107">Head</emphasis>
semiglobular. Inner and outer occipitals particularly strong. Postocellar seta distinct. No inner orbital setae. 2 pairs of fronto-orbitals. 3–4 medium, equally long interfrontal pairs present. Facial plate shiny, oral margin protruding, a flat carina between antennal bases well developed. No upcurved genal seta. First flagellomere rounded with a broad dorsal apex.
</paragraph>
<paragraph id="8E0236B4FFDDD14AFF08FECEFB37FE54" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FECEFEC7FE93" box="[199,288,295,320]" italics="true" pageId="6" pageNumber="107">Thorax.</emphasis>
Scutellum concolorous with the brown mesonotum, without additional scutellars. 1 pair of posterior dorsocentral setae, prescutellar acrostichal distinct though only slightly longer than other acrostichal setae. Acrostichal setulae sparse but comparatively long. 2 subequal pair of katepisternal setae present.
</paragraph>
<paragraph id="8E0236B4FFDDD14AFF08FE7CFD8BFDCB" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FE7CFEE4FE7F" box="[199,259,405,428]" italics="true" pageId="6" pageNumber="107">Legs.</emphasis>
Fore femur with comparatively weak posteroventral setae. Mid coxa with a rather long ventrally directed seta. Mid tibia without a ventral preapical seta, ventro-apical seta strong and with a distinct mid ventral seta. 1 longer seta in the posteroventral row on mid basitarsus. Ventro-apical part of hind tibia indistinct, hind tibia without long setae at all. Claws and pulvilli small.
</paragraph>
<paragraph id="8E0236B4FFDDD14AFF08FDCCFBFEFD5A" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FDCCFF18FDEF" box="[199,255,549,572]" italics="true" pageId="6" pageNumber="107">Wing</emphasis>
unicolorous and without microtrichia. Costa sub-basally with a pair of long setae,
<quantity id="49459B51FFDDD14AFB0AFDCDFA8CFDEF" box="[1221,1387,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="6" pageNumber="107" unit="mm" value="0.105" valueMax="0.11" valueMin="0.1">0.10–0.11 mm</quantity>
,
<emphasis id="BCC9EAA6FFDDD14AFAB5FDCCFA7BFDEF" box="[1402,1436,549,572]" italics="true" pageId="6" pageNumber="107">i.e.</emphasis>
much shorter than in
<taxonomicName id="49BD4D37FFDDD14AFE5FFDAEFE19FDB3" box="[400,510,583,608]" class="Insecta" family="Sphaeroceridae" genus="Rudolfina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="BCC9EAA6FFDDD14AFE5FFDAEFE19FDB3" box="[400,510,583,608]" italics="true" pageId="6" pageNumber="107">Rudolfina</emphasis>
</taxonomicName>
; first costal section otherwise with short setae only. Costa not produced beyond apex of R4+5. Radial vein R4+5 strongly bent up, costa reaches just to apex of R4+5.
</paragraph>
<paragraph id="8E0236B4FFDDD14AFF08FD7BFEA6FD20" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FD7BFEDFFD78" box="[199,312,658,683]" italics="true" pageId="6" pageNumber="107">Abdomen.</emphasis>
Abdominal tergite 2 membranous centrally. Preabdominal sternites various but not small: sternite 2 half as broad as corresponding tergite, sternite 4 2/3 as broad as tergite 4. Preabdomen (except for sternite 5) not modified at all.
</paragraph>
<paragraph id="8E0236B4FFDDD14AFF08FD17FE45FB14" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FD17FEE6FCC4" box="[199,257,766,791]" italics="true" pageId="6" pageNumber="107">Male</emphasis>
sternite 5 (
<figureCitation id="16862A31FFDDD14AFE4DFD17FDE1FCC4" box="[386,518,766,791]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 21–22</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFDDCFD16FDBDFCC4" box="[531,602,767,791]" captionStart="FIGURES 28 – 32" captionStartId="8.[151,264,1678,1700]" captionTargetBox="[340,1253,560,1653]" captionTargetId="figure@8.[330,1256,553,1657]" captionTargetPageId="8" captionText="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." httpUri="https://zenodo.org/record/227673/files/figure.png" pageId="6" pageNumber="107">28–29</figureCitation>
) rather large with a pair of caudal processes (or a simple broad process), which bear a pair of groups of long parallel pegs (5 to 12 each). Anterior to those pegs a larger field of discoloured sharp setulae present. Synsternite (
<figureCitation id="16862A31FFDDD14AFDEBFCAFFD9FFC8C" box="[548,632,838,863]" captionStart="FIGURES 28 – 32" captionStartId="8.[151,264,1678,1700]" captionTargetBox="[340,1253,560,1653]" captionTargetId="figure@8.[330,1256,553,1657]" captionTargetPageId="8" captionText="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." httpUri="https://zenodo.org/record/227673/files/figure.png" pageId="6" pageNumber="107">Fig. 30</figureCitation>
) with a robust sternite 6 part and a broad but not long sternite 8 part. Epandrial complex (
<figureCitation id="16862A31FFDDD14AFEB2FC83FE18FC50" box="[381,511,874,899]" captionStart="FIGURES 28 – 32" captionStartId="8.[151,264,1678,1700]" captionTargetBox="[340,1253,560,1653]" captionTargetId="figure@8.[330,1256,553,1657]" captionTargetPageId="8" captionText="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." httpUri="https://zenodo.org/record/227673/files/figure.png" pageId="6" pageNumber="107">Figs 31–32</figureCitation>
) strongly fused ventrally (in the cercal part), most ventrally with a pair of strong caniniform processes (
<figureCitation id="16862A31FFDDD14AFE69FC67FDE0FC74" box="[422,519,910,935]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 25</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFDD3FC66FDDBFC74" box="[540,572,911,935]" captionStart="FIGURES 28 – 32" captionStartId="8.[151,264,1678,1700]" captionTargetBox="[340,1253,560,1653]" captionTargetId="figure@8.[330,1256,553,1657]" captionTargetPageId="8" captionText="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." httpUri="https://zenodo.org/record/227673/files/figure.png" pageId="6" pageNumber="107">32</figureCitation>
). Epandrium with only one connection to hypandrium on both sides. Hypandrium (
<figureCitation id="16862A31FFDDD14AFEF8FC5BFE6CFC18" box="[311,395,946,971]" captionStart="FIGURES 28 – 32" captionStartId="8.[151,264,1678,1700]" captionTargetBox="[340,1253,560,1653]" captionTargetId="figure@8.[330,1256,553,1657]" captionTargetPageId="8" captionText="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." httpUri="https://zenodo.org/record/227673/files/figure.png" pageId="6" pageNumber="107">Fig. 31</figureCitation>
) strongly fused to epandrium, lateral parts (arms) asymmetrical, medial part (“rod”) short broad and left-side curved. Subepandrial sclerite rather weakly sclerotised (
<figureCitation id="16862A31FFDDD14AFBDFFC3FFB7FFC3C" box="[1040,1176,982,1007]" captionStart="FIGURES 28 – 32" captionStartId="8.[151,264,1678,1700]" captionTargetBox="[340,1253,560,1653]" captionTargetId="figure@8.[330,1256,553,1657]" captionTargetPageId="8" captionText="FIGURES 28 – 32. Biphallapodema polydentata sp. n., male postabdomen. 28 = sternite 5, ventral view, 29 = medio-caudal part of sternite 5 in higher magnification, 30 = synsternite, ventral view, 31 = epandrium and hypandrium, subanteriorsubventral view, 32 = ventral part of epandrium, caudal view. Scale bars: Figs 28, 30 – 31 = 0.2 mm, Figs 29, 32 = 0.1 mm." httpUri="https://zenodo.org/record/227673/files/figure.png" pageId="6" pageNumber="107">Figs 31–32</figureCitation>
), except for a sagittal narrow sub-quadratic part. Surstylus (
<figureCitation id="16862A31FFDDD14AFDF1FC13FD72FBC0" box="[574,661,1018,1043]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 25</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFD6DFC13FD58FBC0" box="[674,703,1018,1043]" captionStart="FIGURES 33 – 38" captionStartId="9.[151,264,1764,1786]" captionTargetBox="[334,1256,595,1741]" captionTargetId="figure@9.[329,1257,591,1741]" captionTargetPageId="9" captionText="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri="https://zenodo.org/record/227674/files/figure.png" pageId="6" pageNumber="107">35</figureCitation>
) comparatively small, compact, without any teeth but with dense medium-long setae. Basiphallus (
<figureCitation id="16862A31FFDDD14AFDC1FBF7FD76FBE4" box="[526,657,1054,1079]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 24, 26</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFD52FBF6FD5DFBE4" box="[669,698,1055,1079]" captionStart="FIGURES 33 – 38" captionStartId="9.[151,264,1764,1786]" captionTargetBox="[334,1256,595,1741]" captionTargetId="figure@9.[329,1257,591,1741]" captionTargetPageId="9" captionText="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri="https://zenodo.org/record/227674/files/figure.png" pageId="6" pageNumber="107">34</figureCitation>
) ventrally directed, hook-like in profile with an upcurved thinner epiphallus. Distiphallus (
<figureCitation id="16862A31FFDDD14AFE73FBABFD92FB88" box="[444,629,1090,1115]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 24, 26–27</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFD47FBAAFD42FB88" box="[648,677,1091,1115]" captionStart="FIGURES 33 – 38" captionStartId="9.[151,264,1764,1786]" captionTargetBox="[334,1256,595,1741]" captionTargetId="figure@9.[329,1257,591,1741]" captionTargetPageId="9" captionText="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri="https://zenodo.org/record/227674/files/figure.png" pageId="6" pageNumber="107">34</figureCitation>
) with a pair of dorsal processes and distinct apical structures. Phallapodeme (
<figureCitation id="16862A31FFDDD14AFE87FB8FFE4EFBAC" box="[328,425,1126,1151]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs. 23</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFE77FB8EFE32FBAC" box="[440,469,1127,1151]" captionStart="FIGURES 33 – 38" captionStartId="9.[151,264,1764,1786]" captionTargetBox="[334,1256,595,1741]" captionTargetId="figure@9.[329,1257,591,1741]" captionTargetPageId="9" captionText="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri="https://zenodo.org/record/227674/files/figure.png" pageId="6" pageNumber="107">33</figureCitation>
) extremely long rod-like, its base connected to postgonites through extremely long hairpin-like flexible black connections, which has the appearance of a second phallapodeme. Postgonite (
<figureCitation id="16862A31FFDDD14AFAF6FB63FF51FB14" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 23– 24</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFF0EFB46FF39FB14" box="[193,222,1199,1223]" captionStart="FIGURES 33 – 38" captionStartId="9.[151,264,1764,1786]" captionTargetBox="[334,1256,595,1741]" captionTargetId="figure@9.[329,1257,591,1741]" captionTargetPageId="9" captionText="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri="https://zenodo.org/record/227674/files/figure.png" pageId="6" pageNumber="107">34</figureCitation>
) large and broad.
</paragraph>
<paragraph id="8E0236B4FFDDD14AFF08FB3BFB94FADC" blockId="6.[151,1437,151,1296]" pageId="6" pageNumber="107">
<emphasis id="BCC9EAA6FFDDD14AFF08FB3BFEFCFB38" box="[199,283,1234,1259]" italics="true" pageId="6" pageNumber="107">Female</emphasis>
epiproct large round, cercus normal or slightly elongated with some long setae; spermathecae (
<figureCitation id="16862A31FFDDD14AFA8FFB3BFA7FFB38" box="[1344,1432,1234,1259]" captionStart="FIGURES 21 – 27" captionStartId="7.[151,264,1728,1750]" captionTargetBox="[329,1258,596,1701]" captionTargetId="figure@7.[325,1261,591,1706]" captionTargetPageId="7" captionText="FIGURES 21 – 27. Biphallapodema oligodentata sp. n., male postabdomen and genitalia. 21 = sternite 5, ventral view, 22 = medio-caudal part of sternite 5 in higher magnification, 23 = phallus and phallapodeme, 24 = postgonite with connecting sclerite, 25 = surstylus with ventral part of epandrium, lateral view, 26 = distiphallus with contours of basiphallus, left lateral view, 27 = distiphallus, anterior view. (BPh: basiphallus, con: connecting sclerite of postgonite, PA: phallapodeme, PG: postgonite). Scale bars: Figs 21, 23 = 0.2 mm, Figs 22, 24 – 27 = 0.1 mm." httpUri="https://zenodo.org/record/227672/files/figure.png" pageId="6" pageNumber="107">Figs 27</figureCitation>
,
<figureCitation id="16862A31FFDDD14AFF58FB1EFF53FADC" box="[151,180,1271,1295]" captionStart="FIGURES 33 – 38" captionStartId="9.[151,264,1764,1786]" captionTargetBox="[334,1256,595,1741]" captionTargetId="figure@9.[329,1257,591,1741]" captionTargetPageId="9" captionText="FIGURES 33 – 38. Biphallapodema polydentata sp. n., male and female genitalia. 33 – 35 = paratype male. 33 = postgonite and phallapodeme with their connecting sclerite, left lateral view, 34 = phallus, left lateral view, 35 = surstylus with ventral part of epandrium, lateral view, 36 – 38: paratype female. 36 = epiproct and cerci, dorsal view, 37 = hypoproct, ventral view, 38 = spermathecae. (con: connecting sclerite, PG: postgonite). Scale bar: Figs 33 – 38 = 0.1 mm." httpUri="https://zenodo.org/record/227674/files/figure.png" pageId="6" pageNumber="107">38</figureCitation>
) various, but sclerotised ducts are very short with a comparatively large bulb distally.
</paragraph>
</subSubSection>
</treatment>
</document>
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