260 lines
33 KiB
XML
260 lines
33 KiB
XML
<document id="972960E40A4586F889CBD5F65EEEC045" ID-DOI="10.11646/zootaxa.5296.4.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984462" ID-ZooBank="B9A0584F-0C12-412C-B926-FF7780C38D3A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685428968832" checkinUser="plazi" docAuthor="Neethu, V. P. & Sabu, K. Thomas" docDate="2023" docId="03F887E9FFFC2605FF3121078E672D80" docLanguage="en" docName="zootaxa.5296.4.8.pdf" docOrigin="Zootaxa 5296 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.4.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clypeuspinus devagiriensis Neethu & Sabu 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="593" masterDocId="FFC1FF91FFFD2601FFA626128F612A38" masterDocTitle="A new species of the genus Clypeuspinus Balkenohl, 2021 (Coleoptera: Carabidae Scaritinae) from India" masterLastPageNumber="594" masterPageNumber="589" pageNumber="590" updateTime="1685430467600" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6C390849233EB62DE3AC375F1C9C22F7">A new species of the genus Clypeuspinus Balkenohl, 2021 (Coleoptera: Carabidae Scaritinae) from India</mods:title>
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<mods:name id="944AF1C023840A26E56524EA7A041593" type="personal">
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<mods:namePart id="8956942A7C79E0B4B16151D41915303F">Neethu, V. P.</mods:namePart>
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<mods:nameIdentifier id="3543C00ACCE33035AE153C6C1A528EF7" type="ORCID">0000-0002-7785-2397</mods:nameIdentifier>
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<mods:affiliation id="FAC267225BFA357B17A502A87EDA5C5F">Entomology Research Lab, Post Graduate & Research Department of Zoology, College Research Centre Affiliated to University of Calicut, St. Joseph’s College (Autonomous), Devagiri, Kozhikode, Kerala 673008, India & neethuvp 333 @ gmail. com; https: // orcid. org / 0000 - 0002 - 7785 - 2397</mods:affiliation>
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<mods:nameIdentifier id="11A489B7F9C86EFF3CF0D9E61F720B34" type="email">neethuvp333@gmail.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="A174604094EE3618A1C4891913D01A16">Sabu, K. Thomas</mods:namePart>
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<mods:nameIdentifier id="7226F96459754FF83AB224230DE823FC" type="ORCID">0000-0002-8701-0484</mods:nameIdentifier>
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<mods:affiliation id="212FB0C1B7E6175B014F61DCB9D9366D">Entomology Research Lab, Post Graduate & Research Department of Zoology, College Research Centre Affiliated to University of Calicut, St. Joseph’s College (Autonomous), Devagiri, Kozhikode, Kerala 673008, India & sabukthomas @ gmail. com; https: // orcid. org / 0000 - 0002 - 8701 - 0484</mods:affiliation>
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<mods:nameIdentifier id="0FB1DFD478D327AA3B1C56D314EEB622" type="email">sabukthomas@gmail.com</mods:nameIdentifier>
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<mods:title id="8BA20DF8350191E77EF098CFD165877D">Zootaxa</mods:title>
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<mods:part id="28F652EA5D9ADD4050B76D50706D3EE4">
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<mods:date id="9EFAA07DBA64398B2C645B1840AF4883">2023</mods:date>
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<mods:number id="14BE18A15D8ED7626748F7E786549109">2023-05-30</mods:number>
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<mods:url id="11A498753DEDBB98B0E611EE8CE209D0">http://dx.doi.org/10.11646/zootaxa.5296.4.8</mods:url>
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<mods:classification id="4191D97224D1122DF10579629B138752">journal article</mods:classification>
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<subSubSection id="C34B6574FFFC2600FF3121078D242D17" box="[151,581,1813,1840]" pageId="1" pageNumber="590" type="nomenclature">
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<paragraph id="8BEE36FFFFFC2600FF3121078D242D17" blockId="1.[151,581,1813,1875]" box="[151,581,1813,1840]" pageId="1" pageNumber="590">
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<heading id="D0A68193FFFC2600FF3121078D242D17" bold="true" box="[151,581,1813,1840]" fontSize="11" level="1" pageId="1" pageNumber="590" reason="1">
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<emphasis id="B925EAEDFFFC2600FF3121078D242D17" bold="true" box="[151,581,1813,1840]" pageId="1" pageNumber="590">
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<taxonomicName id="4C514D7CFFFC2600FF3121078E812D08" authority="Neethu & Sabu, 2023" authorityName="Neethu & Sabu" authorityYear="2023" box="[151,480,1813,1840]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
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<emphasis id="B925EAEDFFFC2600FF3121078E812D08" bold="true" box="[151,480,1813,1840]" italics="true" pageId="1" pageNumber="590">Clypeuspinus devagiriensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2165796FFFC2600FE4121078D242D17" box="[487,581,1813,1839]" pageId="1" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6574FFFC2600FF31212B8E7E2D6B" box="[151,287,1849,1875]" pageId="1" pageNumber="590" type="description">
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<paragraph id="8BEE36FFFFFC2600FF31212B8E7E2D6B" blockId="1.[151,581,1813,1875]" box="[151,287,1849,1875]" pageId="1" pageNumber="590">
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(
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<figureCitation id="136A2A7AFFFC2600FF39212B8E762D6B" box="[159,279,1849,1875]" captionStart="FIGURE 3" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[215,1383,203,1879]" captionTargetId="figure-17@3.[179,1407,181,1920]" captionTargetPageId="3" captionText="FIGURE 3. Clypeuspinus devagiriensis sp. nov. Holotype, male.A) habitus, dorsal view; B) habitus, ventral view; C) aedeagus, lateral view; D) aedeagus, dorsal view; E) aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7984472" httpUri="https://zenodo.org/record/7984472/files/figure.png" pageId="1" pageNumber="590">Fig. 3A–E</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6574FFFC2600FF3121938B882D87" pageId="1" pageNumber="590" type="materials_examined">
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<paragraph id="8BEE36FFFFFC2600FF3121938B882D87" blockId="1.[151,1436,1921,2020]" pageId="1" pageNumber="590">
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<emphasis id="B925EAEDFFFC2600FF3121938E262DA3" bold="true" box="[151,327,1921,1947]" pageId="1" pageNumber="590">Type material.</emphasis>
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<typeStatus id="54EA885DFFFC2600FEF621938ED92DA3" box="[336,440,1921,1947]" pageId="1" pageNumber="590">Holotype</typeStatus>
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male,
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<materialsCitation id="3B393CA2FFFC2600FDA121908B842D87" collectingDate="2021-09-21" collectionCode="V, ZSIK" collectorName="P. Neethu" country="India" latitude="11.2604" location="Labelled" longLatPrecision="7" longitude="75.9391" pageId="1" pageNumber="590" specimenCount="8" specimenCount-female="2" specimenCount-male="2" stateProvince="Kerala" typeStatus="paratype">
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<typeStatus id="54EA885DFFFC2600FDA121908D152DA4" box="[519,628,1922,1948]" pageId="1" pageNumber="590">Paratypes</typeStatus>
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<specimenCount id="9D57FD76FFFC2600FDDA21938DD42DA4" box="[636,693,1921,1948]" pageId="1" pageNumber="590" type="generic">4 ex.</specimenCount>
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(
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<specimenCount id="9D57FD76FFFC2600FD6021938C422DA3" box="[710,803,1921,1947]" pageId="1" pageNumber="590" type="male">2 males</specimenCount>
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;
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<specimenCount id="9D57FD76FFFC2600FC9621938CC22DA3" box="[816,931,1921,1947]" pageId="1" pageNumber="590" type="female">2 females</specimenCount>
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).
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<location id="8E8E6024FFFC2600FC1121938B7C2DA3" LSID="urn:lsid:plazi:treatment:03F887E9FFFC2605FF3121078E672D80:8E8E6024FFFC2600FC1121938B7C2DA3" box="[951,1053,1921,1947]" country="India" latitude="11.2604" longLatPrecision="7" longitude="75.9391" name="Labelled" pageId="1" pageNumber="590" stateProvince="Kerala">Labelled</location>
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: “
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<collectingCountry id="F346766FFFFC2600FB9221938B152DA3" box="[1076,1140,1921,1947]" name="India" pageId="1" pageNumber="590">India</collectingCountry>
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:
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<collectingRegion id="4995F81DFFFC2600FB2621938BAE2DA3" box="[1152,1231,1921,1947]" country="India" name="Kerala" pageId="1" pageNumber="590">Kerala</collectingRegion>
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: Mavoor wetland, Palliyol (
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<geoCoordinate id="EE655038FFFC2600FF5B21B78EE32DF8" box="[253,386,1957,1984]" degrees="11.2604" direction="north" orientation="latitude" pageId="1" pageNumber="590" precision="5" value="11.2604">11.2604° N</geoCoordinate>
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,
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<geoCoordinate id="EE655038FFFC2600FE2B21B78D6F2D87" box="[397,526,1957,1984]" degrees="75.9391" direction="east" orientation="longitude" pageId="1" pageNumber="590" precision="5" value="75.9391">75.9391° E</geoCoordinate>
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), ‘Light trap’,
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<date id="FFEF103FFFFC2600FD1421B78C4C2D87" box="[690,813,1957,1983]" pageId="1" pageNumber="590" value="2021-09-21">
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<collectingDate id="EFABE9D7FFFC2600FD1421B78C4C2D87" box="[690,813,1957,1983]" pageId="1" pageNumber="590" value="2021-09-21">21.ix.2021</collectingDate>
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</date>
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. coll.
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||
<collectionCode id="ED40AE3AFFFC2600FCD621B48CE52DF8" box="[880,900,1958,1984]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="590" type="Museum">V</collectionCode>
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||
.
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<collectorName id="26A45329FFFC2600FC2821B48C9E2D87" box="[910,1023,1957,1984]" pageId="1" pageNumber="590">P. Neethu</collectorName>
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”, deposited in
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<collectionCode id="ED40AE3AFFFC2600FB0521B78B842D87" box="[1187,1253,1957,1983]" pageId="1" pageNumber="590">ZSIK</collectionCode>
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</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C34B6574FFFC2605FF6121DB8E392C69" lastPageId="4" lastPageNumber="593" pageId="1" pageNumber="590" type="description">
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<paragraph id="8BEE36FFFFFC2600FF6121DB8C272DDC" blockId="1.[151,1436,1921,2020]" box="[199,838,1993,2020]" pageId="1" pageNumber="590">
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<emphasis id="B925EAEDFFFC2600FF6121DB8E382DDB" bold="true" box="[199,345,1993,2019]" pageId="1" pageNumber="590">Description.</emphasis>
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Dorsal and ventral habitus as in
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<figureCitation id="136A2A7AFFFC2600FD6E21DB8C232DDC" box="[712,834,1993,2020]" captionStart="FIGURE 3" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[215,1383,203,1879]" captionTargetId="figure-17@3.[179,1407,181,1920]" captionTargetPageId="3" captionText="FIGURE 3. Clypeuspinus devagiriensis sp. nov. Holotype, male.A) habitus, dorsal view; B) habitus, ventral view; C) aedeagus, lateral view; D) aedeagus, dorsal view; E) aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7984472" httpUri="https://zenodo.org/record/7984472/files/figure.png" pageId="1" pageNumber="590">Fig. 3A, B</figureCitation>
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.
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</paragraph>
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<caption id="DF2E6677FFFF2603FF3121938E142D85" ID-DOI="http://doi.org/10.5281/zenodo.7984466" ID-Zenodo-Dep="7984466" httpUri="https://zenodo.org/record/7984466/files/figure.png" pageId="2" pageNumber="591" startId="2.[151,250,1921,1945]" targetBox="[156,1431,186,1892]" targetPageId="2" targetType="figure">
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<paragraph id="8BEE36FFFFFF2603FF3121938E142D85" blockId="2.[151,1437,1921,1981]" pageId="2" pageNumber="591">
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<emphasis id="B925EAEDFFFF2603FF3121938E752DA1" bold="true" box="[151,276,1921,1945]" pageId="2" pageNumber="591">FIGURE 2.</emphasis>
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Geographic distribution of the genus
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<taxonomicName id="4C514D7CFFFF2603FD3F21938C432DA1" authorityName="Balkenohl" authorityYear="2021" box="[665,802,1921,1945]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="genus">
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<emphasis id="B925EAEDFFFF2603FD3F21938C432DA1" box="[665,802,1921,1945]" italics="true" pageId="2" pageNumber="591">Clypeuspinus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C514D7CFFFF2603FCEF21938B832DA1" authority="(Andrewes, 1936)" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[841,1250,1921,1945]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="validus">
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<emphasis id="B925EAEDFFFF2603FCEF21938B502DA0" box="[841,1073,1921,1945]" italics="true" pageId="2" pageNumber="591">Clypeuspinus validus (</emphasis>
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<bibRefCitation id="EFC04B0EFFFF2603FB9721938BBA2DA1" author="Andrewes, H. E." box="[1073,1243,1921,1945]" pageId="2" pageNumber="591" pagination="201 - 209" refId="ref2125" refString="Andrewes, H. E. (1936) Descriptions of some new Carabidae from Ceylon (Part III). Spolia Zeylanica, 19, 201 - 209." type="journal article" year="1936">Andrewes, 1936</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="4C514D7CFFFF2603FAB221938E7F2D85" authority="Neethu & Sabu, 2023" authorityName="Neethu & Sabu" authorityYear="2023" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
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<emphasis id="B925EAEDFFFF2603FAB221938E7F2D85" italics="true" pageId="2" pageNumber="591">Clypeuspinus devagiriensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2165796FFFF2603FE8321B78E142D85" box="[293,373,1957,1981]" pageId="2" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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||
<caption id="DF2E6677FFFE2602FF31218A8C4C2DEC" ID-DOI="http://doi.org/10.5281/zenodo.7984472" ID-Zenodo-Dep="7984472" httpUri="https://zenodo.org/record/7984472/files/figure.png" pageId="3" pageNumber="592" startId="3.[151,250,1944,1968]" targetBox="[215,1383,203,1879]" targetPageId="3" targetType="figure">
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<paragraph id="8BEE36FFFFFE2602FF31218A8C4C2DEC" blockId="3.[151,1436,1944,2004]" pageId="3" pageNumber="592">
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||
<emphasis id="B925EAEDFFFE2602FF31218A8E732D88" bold="true" box="[151,274,1944,1968]" pageId="3" pageNumber="592">FIGURE 3.</emphasis>
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<taxonomicName id="4C514D7CFFFE2602FEB1218A8D4A2D88" authority="Neethu & Sabu, 2023" authorityName="Neethu & Sabu" authorityYear="2023" box="[279,555,1944,1968]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
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<emphasis id="B925EAEDFFFE2602FEB1218A8D4A2D88" box="[279,555,1944,1968]" italics="true" pageId="3" pageNumber="592">Clypeuspinus devagiriensis</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B925EAEDFFFE2602FD97218A8DE12D88" bold="true" box="[561,640,1944,1968]" pageId="3" pageNumber="592">
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||
<taxonomicNameLabel id="A2165796FFFE2602FD97218A8DE12D88" box="[561,640,1944,1968]" pageId="3" pageNumber="592" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Holotype, male.A) habitus, dorsal view; B) habitus, ventral view; C) aedeagus, lateral view; D) aedeagus, dorsal view; E) aedeagus, lateral view.
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</paragraph>
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</caption>
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<paragraph id="8BEE36FFFFF92605FF6126858BB02AC1" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">Head, pronotum, elytral margin, elytral suture, profemurs and tibiae dark reddish brown. Meso and metacoxae, femurs and metatrochanters light reddish-brown. Antennae, labial palpi, maxillary palpi, protarsomers, meso and metatibiae and tarsomeres yellowish brown. Elytral disc black. Abdominal ventrites dark brown.</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6127118BD928F5" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">Head elongated, smaller than pronotum. Surface smooth, with scattered micropunctures. Labrum three setose with isodiametric meshes. Clypeus straight, margin not bordered, clypeal suture indistinct, wings extending forward and rounded at apex, obtuse emargination between clypeus and clypeal wings blunt, clypeal setigerous punctures situated at anterior part of frontal furrows. Supra antennal plates elongate with reflexed margin in posterior half. Frons elongated and convex, frontal furrows deep. Vertex convex with fine micropuntures. Supra orbital carinae sharp. Supra orbital furrows deep and wide, with two supra orbital setae. Eyes normally developed, genae as long as eyes. Neck constriction complete, composed of large punctures. Antennae reaching up to middle of pronotum, pubescent from third antennomere onwards, antennomeres four to eleven moniliform, pedicellus attached eccentrically to scapus, scapus with setae at apical quarter. Mandibles elongated and curved apically. Apical maxillary palpomeres elongated, slightly securiform, with slightly truncated end. Penultimate labial palpomeres bisetose. Mentum and submentum separated. Mentum wide with two pairs of setae (a pair towards base of median tooth, another pair towards base of mentum), median tooth moderately wide and truncate at apex, projecting as far as lateral lobes, lateral lobes wide with rugose surface. Submentum quadrisetose. Genae with dense rugosities.</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6124C58F9929B9" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">Pronotum pentagonal in shape, moderately convex, surface glabrous, glossy with scattered micropunctures and transverse wrinkles, front angles produced, hind angles with a small denticle breaking through the outline, anterior transverse line deep and well impressed with large punctures, median line deep and wide, joining with basal constriction, lateral channel widened between two lateral setigerous punctures. Reflexed lateral margins, flange carinate.</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6125998EF12E45" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">Elytra moderately convex, elongate, nearly parallel, well-marked shoulders with distinct humeral tooth, scutellar strioles moderately developed, striae deep and punctuated, stria one joining to setigerous basal tubercle, striae two and three free at base, striae four and five fused at base, striae one and seven reaching apex, interval eight carinate from humerus to apex, intervals six and seven carinate at humerus only, intervals two, three and four with distinct basal tubercle, interval three with two setigerous punctures close to third stria, with reflexed lateral margins, marginal channels with uninterrupted series of setigerous punctures, extreme apex of elytra with isodiametric meshes. Hind wings fully developed</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6122958C422F09" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">
|
||
Ventral side (
|
||
<figureCitation id="136A2A7AFFF92605FEFA22958ED72E99" box="[348,438,1159,1185]" captionStart="FIGURE 3" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[215,1383,203,1879]" captionTargetId="figure-17@3.[179,1407,181,1920]" captionTargetPageId="3" captionText="FIGURE 3. Clypeuspinus devagiriensis sp. nov. Holotype, male.A) habitus, dorsal view; B) habitus, ventral view; C) aedeagus, lateral view; D) aedeagus, dorsal view; E) aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7984472" httpUri="https://zenodo.org/record/7984472/files/figure.png" pageId="4" pageNumber="593">Fig. 3B</figureCitation>
|
||
): Prosternal surface rugose, with isodiametric meshes at disc. Proepisternum with coarse punctures and rugosities. Prosternal process wide. Metasternum glossy, with transverse wrinkles and micropunctures. Metepisternum longer than wide. Abdominal ventrites smooth and glossy, third to fifth ventrites with a pair of median setae, last ventrite with variable number of median setae (ranges from zero to three) and with a pair of widely separated apical setigerous punctures on each side.
|
||
</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6123298CB02F41" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">Protibiae with three terminal spines, slender, curved laterally. Metatibiae with long protuberance preapically furnished with seta. Claws simple. Tarsomeres not widened in both sexes.</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6123918D362FF9" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">
|
||
Male genitalia: Median lobe well curved, pointed towards apex, right paramere wider than left paramere, both parameres bisetose at apex (
|
||
<figureCitation id="136A2A7AFFF92605FE7423B58D2B2FF9" box="[466,586,1447,1473]" captionStart="FIGURE 3" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[215,1383,203,1879]" captionTargetId="figure-17@3.[179,1407,181,1920]" captionTargetPageId="3" captionText="FIGURE 3. Clypeuspinus devagiriensis sp. nov. Holotype, male.A) habitus, dorsal view; B) habitus, ventral view; C) aedeagus, lateral view; D) aedeagus, dorsal view; E) aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7984472" httpUri="https://zenodo.org/record/7984472/files/figure.png" pageId="4" pageNumber="593">Fig. 3C–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6123D98DCA2FDD" blockId="4.[151,1437,151,1977]" box="[199,683,1483,1509]" pageId="4" pageNumber="593">Sexual dimorphism: Not externally visible.</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6123FD8E392C69" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">
|
||
<emphasis id="B925EAEDFFF92605FF6123FD8E1F2C31" bold="true" box="[199,382,1519,1545]" pageId="4" pageNumber="593">Measurements.</emphasis>
|
||
<typeStatus id="54EA885DFFF92605FE2223FD8E8D2C31" box="[388,492,1519,1545]" pageId="4" pageNumber="593">Holotype</typeStatus>
|
||
(male), TLB =
|
||
<quantity id="4CA99B1AFFF92605FD3A23FD8C632C31" box="[668,770,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="6.06" pageId="4" pageNumber="593" unit="mm" value="6.06">6.06 mm</quantity>
|
||
, PL =
|
||
<quantity id="4CA99B1AFFF92605FCEC23FC8CD12C31" box="[842,944,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="4" pageNumber="593" unit="mm" value="1.52">1.52 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CA99B1AFFF92605FBA423FC8B092C31" box="[1026,1128,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="593" unit="mm" value="1.55">1.55 mm</quantity>
|
||
, EL =
|
||
<quantity id="4CA99B1AFFF92605FB1423FD8A792C31" box="[1202,1304,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="593" unit="mm" value="3.3">3.30 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CA99B1AFFF92605FACD23FD8FA72C15" metricMagnitude="-3" metricUnit="m" metricValue="1.73" pageId="4" pageNumber="593" unit="mm" value="1.73">1.73 mm</quantity>
|
||
;
|
||
<typeStatus id="54EA885DFFF92605FF7420018E552C15" box="[210,308,1555,1581]" pageId="4" pageNumber="593">Paratype</typeStatus>
|
||
<emphasis id="B925EAEDFFF92605FE9A20008E242C14" bold="true" box="[316,325,1554,1580]" pageId="4" pageNumber="593">(</emphasis>
|
||
n=4
|
||
<emphasis id="B925EAEDFFF92605FED720008EE32C14" bold="true" box="[369,386,1554,1580]" pageId="4" pageNumber="593">),</emphasis>
|
||
TLB =
|
||
<quantity id="4CA99B1AFFF92605FE7820008DE52C15" box="[478,644,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="6.035" metricValueMax="6.38" metricValueMin="5.69" pageId="4" pageNumber="593" unit="mm" value="6.035" valueMax="6.38" valueMin="5.69">5.69–6.38 mm</quantity>
|
||
, PL =
|
||
<quantity id="4CA99B1AFFF92605FD7620018C172C15" box="[720,886,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" metricValueMax="1.61" metricValueMin="1.41" pageId="4" pageNumber="593" unit="mm" value="1.51" valueMax="1.61" valueMin="1.41">1.41–1.61 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CA99B1AFFF92605FC6D20018B102C15" box="[971,1137,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.62" metricValueMin="1.43" pageId="4" pageNumber="593" unit="mm" value="1.525" valueMax="1.62" valueMin="1.43">1.43–1.62 mm</quantity>
|
||
, EL =
|
||
<quantity id="4CA99B1AFFF92605FB1920008A042C15" box="[1215,1381,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="3.3049999999999997" metricValueMax="3.53" metricValueMin="3.08" pageId="4" pageNumber="593" unit="mm" value="3.3049999999999997" valueMax="3.53" valueMin="3.08">3.08–3.53 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CA99B1AFFF92605FF0820258E352C69" box="[174,340,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.79" metricValueMin="1.62" pageId="4" pageNumber="593" unit="mm" value="1.705" valueMax="1.79" valueMin="1.62">1.62–1.79 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34B6574FFF92605FF6120498B7F2C4D" box="[199,1054,1627,1653]" pageId="4" pageNumber="593" type="etymology">
|
||
<paragraph id="8BEE36FFFFF92605FF6120498B7F2C4D" blockId="4.[151,1437,151,1977]" box="[199,1054,1627,1653]" pageId="4" pageNumber="593">
|
||
<emphasis id="B925EAEDFFF92605FF6120498E2F2C4D" bold="true" box="[199,334,1627,1653]" pageId="4" pageNumber="593">Etymology.</emphasis>
|
||
Named after the host institution's local name, Devagiri College.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34B6574FFF92605FF61206D8A0F2D75" pageId="4" pageNumber="593" type="discussion">
|
||
<paragraph id="8BEE36FFFFF92605FF61206D8A0F2D75" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">
|
||
<emphasis id="B925EAEDFFF92605FF61206D8E5A2CA1" bold="true" box="[199,315,1663,1689]" pageId="4" pageNumber="593">Remarks.</emphasis>
|
||
Similar to
|
||
<taxonomicName id="4C514D7CFFF92605FE66206D8DD02CA1" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[448,689,1663,1689]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FE66206D8DD02CA1" box="[448,689,1663,1689]" italics="true" pageId="4" pageNumber="593">Clypeuspinus validus</emphasis>
|
||
</taxonomicName>
|
||
but differs in the following characters: labrum three setose (five setose in
|
||
<taxonomicName id="4C514D7CFFF92605FEA620B18E132C85" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[256,370,1699,1725]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FEA620B18E132C85" box="[256,370,1699,1725]" italics="true" pageId="4" pageNumber="593">C. validus</emphasis>
|
||
</taxonomicName>
|
||
); obtuse emargination between clypeus and clypeal wings blunt (obtuse emargination between clypeus and clypeal wings sharp in
|
||
<taxonomicName id="4C514D7CFFF92605FD8E20D58DFA2CD9" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[552,667,1735,1761]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FD8E20D58DFA2CD9" box="[552,667,1735,1761]" italics="true" pageId="4" pageNumber="593">C. validus</emphasis>
|
||
</taxonomicName>
|
||
); supraorbital carinae sharp blunt in
|
||
<taxonomicName id="4C514D7CFFF92605FB9D20D58BCF2CD9" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[1083,1198,1735,1761]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FB9D20D58BCF2CD9" box="[1083,1198,1735,1761]" italics="true" pageId="4" pageNumber="593">C. validus</emphasis>
|
||
</taxonomicName>
|
||
); elytra elongate and nearly parallel (subelongate in
|
||
<taxonomicName id="4C514D7CFFF92605FE5220F98D072D3D" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[500,614,1771,1797]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FE5220F98D072D3D" box="[500,614,1771,1797]" italics="true" pageId="4" pageNumber="593">C. validus</emphasis>
|
||
</taxonomicName>
|
||
); third elytral interval with two setigerous punctures (in
|
||
<taxonomicName id="4C514D7CFFF92605FB4120F98A382D3D" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[1255,1369,1771,1797]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FB4120F98A382D3D" box="[1255,1369,1771,1797]" italics="true" pageId="4" pageNumber="593">C. validus</emphasis>
|
||
</taxonomicName>
|
||
, third interval of left elytron with three setigerous punctures and right elytron with four setigerous punctures); second, third and fourth intervals with distinct basal tubercle (only third interval with a distinct basal tubercle in
|
||
<taxonomicName id="4C514D7CFFF92605FB4A21218A3F2D75" baseAuthorityName="Andrewes" baseAuthorityYear="1936" box="[1260,1374,1843,1869]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="validus">
|
||
<emphasis id="B925EAEDFFF92605FB4A21218A3F2D75" box="[1260,1374,1843,1869]" italics="true" pageId="4" pageNumber="593">C. validus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34B6574FFF92605FF6121448E672D80" pageId="4" pageNumber="593" type="distribution">
|
||
<paragraph id="8BEE36FFFFF92605FF6121448C0C2D49" blockId="4.[151,1437,151,1977]" box="[199,877,1878,1905]" pageId="4" pageNumber="593">
|
||
<emphasis id="B925EAEDFFF92605FF6121448E002D48" bold="true" box="[199,353,1878,1904]" pageId="4" pageNumber="593">Distribution.</emphasis>
|
||
<collectingCountry id="F346766FFFF92605FEC121458EDA2D49" box="[359,443,1879,1905]" name="India" pageId="4" pageNumber="593">INDIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4995F81DFFF92605FE6021458D742D49" box="[454,533,1879,1905]" country="India" name="Kerala" pageId="4" pageNumber="593">Kerala</collectingRegion>
|
||
: Kozhikode: Mavoor, Palliyol.
|
||
</paragraph>
|
||
<paragraph id="8BEE36FFFFF92605FF6121698E672D80" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="593">
|
||
<taxonomicName id="4C514D7CFFF92605FF6121698E9F2DAD" authority="Neethu & Sabu, 2023" authorityName="Neethu & Sabu" authorityYear="2023" box="[199,510,1915,1941]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
|
||
<emphasis id="B925EAEDFFF92605FF6121698E9F2DAD" box="[199,510,1915,1941]" italics="true" pageId="4" pageNumber="593">Clypeuspinus devagiriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B925EAEDFFF92605FDAF21698D062DAD" bold="true" box="[521,615,1915,1941]" pageId="4" pageNumber="593">
|
||
<taxonomicNameLabel id="A2165796FFF92605FDAF21698D062DAD" box="[521,615,1915,1941]" pageId="4" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the first species of the genus
|
||
<taxonomicName id="4C514D7CFFF92605FC4B21698BE52DAD" authorityName="Balkenohl" authorityYear="2021" box="[1005,1156,1915,1941]" class="Insecta" family="Carabidae" genus="Clypeuspinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925EAEDFFF92605FC4B21698BE52DAD" box="[1005,1156,1915,1941]" italics="true" pageId="4" pageNumber="593">Clypeuspinus</emphasis>
|
||
</taxonomicName>
|
||
discovered from Indian mainland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |