290 lines
37 KiB
XML
290 lines
37 KiB
XML
<document id="7E7B242618118F5C2FBDD3F477D5C66C" ID-CLB-Dataset="27734" ID-DOI="10.11646/zootaxa.4532.4.4" ID-GBIF-Dataset="4809e341-99ce-476f-a3a8-5229de62f5eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615629" ID-ZooBank="A35A92D5-2628-4203-8C94-DDBFFFCDC79E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788686464" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. & Telnov, Dmitry" docDate="2018" docId="03F887BAFFA1FFDDFF37F9871B35FDF9" docLanguage="en" docName="zootaxa.4532.4.4.pdf" docOrigin="Zootaxa 4532 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Distylopyga Bey-Bienko 1965" docType="treatment" docVersion="8" lastPageNumber="531" masterDocId="FFC1FFC2FFA2FFD5FFA0FFB4184CFE79" masterDocTitle="New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)" masterLastPageNumber="538" masterPageNumber="523" pageNumber="526" updateTime="1698715127610" updateUser="ExternalLinkService">
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<mods:mods id="5DB5BFB0D0E7425A0D011C010CC2C1B4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5DA10DFCA46D4F78AC722E2416E1D76B">
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<mods:title id="8D667C8E730054657B3BCAB76AC68DFF">New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)</mods:title>
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<mods:namePart id="114839008EE60AC71705CA74F403C863">Anisyutkin, Leonid N.</mods:namePart>
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<mods:name id="A27124F04C5C758659697048C4AD27C5" type="personal">
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<mods:namePart id="55CDD30473A67720915F95393C9979B1">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="AD45422E78BEB3FA9EC28CD8045DE917">text</mods:typeOfResource>
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<mods:title id="5F9D817167D3AA8D781D9C4C88CA2870">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E2EF300E45B500AE4E272A508E8937BE">
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<mods:date id="95052B5B7B38572A66D88F4A0E1E0F40">2018</mods:date>
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<mods:number id="CCC601932026D59695B2F96DA52AADA2">2018-12-20</mods:number>
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<mods:detail id="CD7E9AC21675F90A70B1213D48CBA0CE" type="volume">
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<mods:number id="5810E6BDDCA7FBF3763AC3BF54470EE8">4532</mods:number>
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<mods:number id="3228994833FC7C247B3C3116E8BA75DA">4</mods:number>
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<treatment id="03F887BAFFA1FFDDFF37F9871B35FDF9" ID-DOI="http://doi.org/10.5281/zenodo.6494961" ID-GBIF-Taxon="154521470" ID-Zenodo-Dep="6494961" LSID="urn:lsid:plazi:treatment:03F887BAFFA1FFDDFF37F9871B35FDF9" httpUri="http://treatment.plazi.org/id/03F887BAFFA1FFDDFF37F9871B35FDF9" lastPageId="8" lastPageNumber="531" pageId="3" pageNumber="526">
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<subSubSection id="C34B6527FFA1FFD6FF37F9871A11F834" box="[151,605,1586,1613]" pageId="3" pageNumber="526" type="nomenclature">
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<paragraph id="8BEE36ACFFA1FFD6FF37F9871A11F834" blockId="3.[151,605,1586,1613]" box="[151,605,1586,1613]" pageId="3" pageNumber="526">
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<heading id="D0A681C0FFA1FFD6FF37F9871A11F834" bold="true" box="[151,605,1586,1613]" fontSize="11" level="1" pageId="3" pageNumber="526" reason="1">
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<emphasis id="B925EABEFFA1FFD6FF37F9871A11F834" bold="true" box="[151,605,1586,1613]" pageId="3" pageNumber="526">
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Genus
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<taxonomicName id="4C514D2FFFA1FFD6FF4EF9861A11F834" ID-CoL="8N6YK" authority="Bey-Bienko, 1965" authorityName="Bey-Bienko" authorityYear="1965" box="[238,605,1586,1613]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">
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<emphasis id="B925EABEFFA1FFD6FF4EF9861930F835" bold="true" box="[238,380,1586,1612]" italics="true" pageId="3" pageNumber="526">Distylopyga</emphasis>
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<bibRefCitation id="EFC04B5DFFA1FFD6FE23F9871A11F834" author="Bey-Bienko, G. Ya." box="[387,605,1587,1613]" pageId="3" pageNumber="526" pagination="1637 - 1650" refId="ref7068" refString="Bey-Bienko, G. Ya. (1965) Orthopteroid insects of the orders Blattoptera and Dermaptera from Komodo and adjacent islands in Indonesia. Zoologicheskiy Zhurnal, 44 (11), 1637 - 1650. [in Russian, English abstract]" type="journal article" year="1965">Bey-Bienko, 1965</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6527FFA1FFD6FF37F9CD1B2BF91C" pageId="3" pageNumber="526" type="materials_examined">
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<paragraph id="8BEE36ACFFA1FFD6FF37F9CD1B49F8F6" blockId="3.[151,773,1657,1679]" box="[151,773,1657,1679]" pageId="3" pageNumber="526">
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||
<emphasis id="B925EABEFFA1FFD6FF37F9CD1969F8F6" bold="true" box="[151,293,1657,1679]" pageId="3" pageNumber="526">
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<typeStatus id="54EA880EFFA1FFD6FF37F9CD1880F8F6" box="[151,204,1657,1679]" pageId="3" pageNumber="526">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C514D2FFFA1FFD6FE8BF9CD1A27F8F6" authority="Krauss, 1903" authorityName="Krauss" authorityYear="1903" box="[299,619,1657,1679]" class="Insecta" family="Blattidae" genus="Stylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="semoni">
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<emphasis id="B925EABEFFA1FFD6FE8BF9CD1997F8F6" box="[299,475,1657,1679]" italics="true" pageId="3" pageNumber="526">Stylopyga semoni</emphasis>
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<bibRefCitation id="EFC04B5DFFA1FFD6FE42F9CE1A27F8F6" author="Krauss, H." box="[482,619,1657,1679]" pageId="3" pageNumber="526" pagination="743 - 770" refId="ref7549" refString="Krauss, H. (1903) Orthopteren aus Australien und dem Malayishen Archipel gesammelt von Professor Dr. Richard Semon. Denkschriften der medizinisch-naturwissenschaftlichen Gesellschaft zu Jena, 8, 743 - 770." type="journal article" year="1903">Krauss, 1903</bibRefCitation>
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</taxonomicName>
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. By monotypy.
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</paragraph>
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<paragraph id="8BEE36ACFFA1FFD6FF37F9081B2BF91C" blockId="3.[151,1436,1724,2001]" pageId="3" pageNumber="526">
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<emphasis id="B925EABEFFA1FFD6FF37F9081883F8AC" bold="true" box="[151,207,1724,1749]" pageId="3" pageNumber="526">Note</emphasis>
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. The genus
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<taxonomicName id="4C514D2FFFA1FFD6FEC1F90819A4F8AD" authorityName="Bey-Bienko" authorityYear="1965" box="[353,488,1724,1748]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">
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<emphasis id="B925EABEFFA1FFD6FEC1F90819A4F8AD" box="[353,488,1724,1748]" italics="true" pageId="3" pageNumber="526">Distylopyga</emphasis>
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</taxonomicName>
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was described from a single female collected in Cibodas Botanical garden (
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<collectingRegion id="4995F84EFFA1FFD6FAC2F9081DD4F8AD" box="[1378,1432,1724,1748]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="526">Java</collectingRegion>
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,
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<collectingCountry id="F346763CFFA1FFD6FF37F96B194AF881" box="[151,262,1759,1784]" name="Indonesia" pageId="3" pageNumber="526">Indonesia</collectingCountry>
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) and initially determined by Bey-Bienko as
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<taxonomicName id="4C514D2FFFA1FFD6FC91F9551D11F881" authority="(Krauss, 1903) (Bey-Bienko 1965)" authorityName="Krauss" authorityYear="1903" baseAuthorityName="Krauss" baseAuthorityYear="1903" box="[817,1373,1759,1784]" class="Insecta" family="Blattidae" genus="Stylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="semoni">
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<emphasis id="B925EABEFFA1FFD6FC91F9551B09F881" box="[817,837,1761,1784]" italics="true" pageId="3" pageNumber="526">D</emphasis>
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.
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<emphasis id="B925EABEFFA1FFD6FCFCF9541BE7F88E" box="[860,939,1760,1783]" italics="true" pageId="3" pageNumber="526">semoni</emphasis>
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(
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<bibRefCitation id="EFC04B5DFFA1FFD6FC62F9541C2DF881" author="Krauss, H." box="[962,1121,1760,1784]" pageId="3" pageNumber="526" pagination="743 - 770" refId="ref7549" refString="Krauss, H. (1903) Orthopteren aus Australien und dem Malayishen Archipel gesammelt von Professor Dr. Richard Semon. Denkschriften der medizinisch-naturwissenschaftlichen Gesellschaft zu Jena, 8, 743 - 770." type="journal article" year="1903">Krauss, 1903</bibRefCitation>
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) (
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<bibRefCitation id="EFC04B5DFFA1FFD6FB20F96B1D19F881" author="Bey-Bienko, G. Ya." box="[1152,1365,1759,1784]" pageId="3" pageNumber="526" pagination="1637 - 1650" refId="ref7068" refString="Bey-Bienko, G. Ya. (1965) Orthopteroid insects of the orders Blattoptera and Dermaptera from Komodo and adjacent islands in Indonesia. Zoologicheskiy Zhurnal, 44 (11), 1637 - 1650. [in Russian, English abstract]" type="journal article" year="1965">Bey-Bienko 1965</bibRefCitation>
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)
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</taxonomicName>
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. The characters listed in the diagnosis of
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<taxonomicName id="4C514D2FFFA1FFD6FD8EF8B01AF9F965" authorityName="Bey-Bienko" authorityYear="1965" box="[558,693,1796,1820]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">
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<emphasis id="B925EABEFFA1FFD6FD8EF8B01AF9F965" box="[558,693,1796,1820]" italics="true" pageId="3" pageNumber="526">Distylopyga</emphasis>
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</taxonomicName>
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are insufficient to distinguish this genus from other short-winged
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<taxonomicName id="4C514D2FFFA1FFD6FF37F893194CF939" authorityName="Latreille" authorityYear="1810" box="[151,256,1831,1856]" class="Insecta" family="Blattidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="subFamily" subFamily="Blattinae">Blattinae</taxonomicName>
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. The specimen studied by Bey-Bienko was expected to be deposited in the collection of Zoological Institute RAS, but it could not be located there during our study.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6527FFA1FFDDFF67F8C419C0FFD4" lastPageId="8" lastPageNumber="531" pageId="3" pageNumber="526" type="reference_group">
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<paragraph id="8BEE36ACFFA1FFD6FF67F8C41BD6F9A8" blockId="3.[151,1436,1724,2001]" pageId="3" pageNumber="526">
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<taxonomicName id="4C514D2FFFA1FFD6FF67F8C419EBF9FE" ID-CoL="8MYVM" baseAuthorityName="Krauss" baseAuthorityYear="1903" box="[199,423,1904,1928]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="semoni">
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<emphasis id="B925EABEFFA1FFD6FF67F8C419EBF9FE" box="[199,423,1904,1928]" italics="true" pageId="3" pageNumber="526">Distylopyga semoni</emphasis>
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</taxonomicName>
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was described from Tjibodas (now Cibodas), W
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<collectingRegion id="4995F84EFFA1FFD6FC44F8C41C5AF9FE" box="[996,1046,1904,1927]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="526">Java</collectingRegion>
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(
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<bibRefCitation id="EFC04B5DFFA1FFD6FB87F8C41CF5F9F1" author="Krauss, H." box="[1063,1209,1904,1928]" pageId="3" pageNumber="526" pagination="743 - 770" refId="ref7549" refString="Krauss, H. (1903) Orthopteren aus Australien und dem Malayishen Archipel gesammelt von Professor Dr. Richard Semon. Denkschriften der medizinisch-naturwissenschaftlichen Gesellschaft zu Jena, 8, 743 - 770." type="journal article" year="1903">Krauss 1903</bibRefCitation>
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) and have complex styli, probably similar to those in
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<taxonomicName id="4C514D2FFFA1FFD6FD90F8201AFAF9D5" ID-CoL="8MRQP" authorityName="Rehn" authorityYear="1933" box="[560,694,1940,1964]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">
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<emphasis id="B925EABEFFA1FFD6FD90F8201AFAF9D5" box="[560,694,1940,1964]" italics="true" pageId="3" pageNumber="526">Duchailluia</emphasis>
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</taxonomicName>
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: "...stylis spiniformibus, incurvus" (
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<bibRefCitation id="EFC04B5DFFA1FFD6FBC1F8201D14F9D4" author="Krauss, H." box="[1121,1368,1940,1965]" pageId="3" pageNumber="526" pagination="743 - 770" refId="ref7549" refString="Krauss, H. (1903) Orthopteren aus Australien und dem Malayishen Archipel gesammelt von Professor Dr. Richard Semon. Denkschriften der medizinisch-naturwissenschaftlichen Gesellschaft zu Jena, 8, 743 - 770." type="journal article" year="1903">Krauss 1903, p. 752</bibRefCitation>
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). The original and subsequent descriptions of
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<taxonomicName id="4C514D2FFFA1FFD6FDF4F80D1A8AF9A9" ID-CoL="8MYVM" baseAuthorityName="Krauss" baseAuthorityYear="1903" box="[596,710,1977,2000]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="semoni">
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<emphasis id="B925EABEFFA1FFD6FDF4F80D1A24F9A9" box="[596,616,1977,2000]" italics="true" pageId="3" pageNumber="526">D</emphasis>
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.
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<emphasis id="B925EABEFFA1FFD6FDD6F80D1A8AF9A9" box="[630,710,1977,2000]" italics="true" pageId="3" pageNumber="526">semoni</emphasis>
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</taxonomicName>
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are too simplified.
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</paragraph>
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||
<caption id="DF2E6624FFA6FFD1FF37F8AB1BDFF994" httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="4" pageNumber="527" startId="4.[151,249,1823,1845]" targetBox="[188,1396,214,1803]" targetPageId="4">
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<paragraph id="8BEE36ACFFA6FFD1FF37F8AB1BDFF994" blockId="4.[151,1436,1823,2029]" pageId="4" pageNumber="527">
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<emphasis id="B925EABEFFA6FFD1FF37F8AB195BF94C" bold="true" box="[151,279,1823,1845]" pageId="4" pageNumber="527">FIGURE 1.</emphasis>
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<taxonomicName id="4C514D2FFFA6FFD1FE82F8AB1A28F94C" ID-CoL="8N97K" authority="Anisyutkin" authorityName="Anisyutkin & Telnov" authorityYear="2018" box="[290,612,1823,1845]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="ivindo" status="sp. nov.">
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<emphasis id="B925EABEFFA6FFD1FE82F8AB19AAF94C" box="[290,486,1823,1845]" italics="true" pageId="4" pageNumber="527">Duchailluia ivindo</emphasis>
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Anisyutkin
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</taxonomicName>
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,
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<emphasis id="B925EABEFFA6FFD1FDD3F8AB1A85F94C" bold="true" box="[627,713,1823,1845]" pageId="4" pageNumber="527">
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<taxonomicNameLabel id="A21657C5FFA6FFD1FDD3F8AB1A85F94C" box="[627,713,1823,1845]" pageId="4" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st–4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations:
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<emphasis id="B925EABEFFA6FFD1FDCAF80D1CB8F9B7" box="[618,1268,1976,1998]" italics="true" pageId="4" pageNumber="527">I–VII—1st–7th abdominal tergites respectively, a.s., par., s.lobe.</emphasis>
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—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K.
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</paragraph>
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</caption>
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<caption id="DF2E6624FFA7FFD0FF37F8241A66F99A" httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="5" pageNumber="528" startId="5.[151,249,1936,1958]" targetBox="[207,1407,214,1901]" targetPageId="5">
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<paragraph id="8BEE36ACFFA7FFD0FF37F8241A66F99A" blockId="5.[151,1436,1936,2019]" pageId="5" pageNumber="528">
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<emphasis id="B925EABEFFA7FFD0FF37F824195FF9DF" bold="true" box="[151,275,1936,1958]" pageId="5" pageNumber="528">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C514D2FFFA7FFD0FEB9F8241A1DF9DF" ID-CoL="8N97K" authority="Anisyutkin" authorityName="Anisyutkin & Telnov" authorityYear="2018" box="[281,593,1936,1958]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="528" phylum="Arthropoda" rank="species" species="ivindo" status="sp. nov.">
|
||
<emphasis id="B925EABEFFA7FFD0FEB9F8241994F9DF" box="[281,472,1936,1958]" italics="true" pageId="5" pageNumber="528">Duchailluia ivindo</emphasis>
|
||
Anisyutkin
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B925EABEFFA7FFD0FDFCF8241AE0F9DF" bold="true" box="[604,684,1936,1958]" pageId="5" pageNumber="528">
|
||
<taxonomicNameLabel id="A21657C5FFA7FFD0FDFCF8241AE0F9DF" box="[604,684,1936,1958]" pageId="5" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations:
|
||
<emphasis id="B925EABEFFA7FFD0FAA3F804195FF99A" italics="true" pageId="5" pageNumber="528">a.t., R1G, R1H, R2, R3, s.br.</emphasis>
|
||
—see text. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2E6624FFA4FFD3FF37F8E218E2F99E" httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="6" pageNumber="529" startId="6.[151,249,1878,1900]" targetBox="[230,1361,198,1854]" targetPageId="6">
|
||
<paragraph id="8BEE36ACFFA4FFD3FF37F8E218E2F99E" blockId="6.[151,1436,1878,2023]" pageId="6" pageNumber="529">
|
||
<emphasis id="B925EABEFFA4FFD3FF37F8E21958F915" bold="true" box="[151,276,1878,1900]" pageId="6" pageNumber="529">FIGURE 3.</emphasis>
|
||
Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—
|
||
<taxonomicName id="4C514D2FFFA4FFD3FCB5F8C11B9AF9F2" ID-CoL="8N97K" authority="Anisyutkin & Telnov, 2018" authorityName="Anisyutkin & Telnov" authorityYear="2018" box="[789,982,1909,1931]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="ivindo" status="sp. nov.">
|
||
<emphasis id="B925EABEFFA4FFD3FCB5F8C11B9AF9F2" box="[789,982,1909,1931]" italics="true" pageId="6" pageNumber="529">Duchailluia ivindo</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B925EABEFFA4FFD3FC7FF8C11C7DF9F2" bold="true" box="[991,1073,1909,1931]" pageId="6" pageNumber="529">
|
||
<taxonomicNameLabel id="A21657C5FFA4FFD3FC7FF8C11C7DF9F2" box="[991,1073,1909,1931]" pageId="6" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; E–H—
|
||
<taxonomicName id="4C514D2FFFA4FFD3FB29F8C11D39F9F2" ID-CoL="8N97J" authority="Anisyutkin & Telnov, 2018" authorityName="Anisyutkin & Telnov" authorityYear="2018" box="[1161,1397,1909,1931]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="beccalonii" status="sp. nov.">
|
||
<emphasis id="B925EABEFFA4FFD3FB29F8C11D39F9F2" box="[1161,1397,1909,1931]" italics="true" pageId="6" pageNumber="529">Distylopyga beccalonii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B925EABEFFA4FFD3FADEF8C1188FF9D3" bold="true" pageId="6" pageNumber="529">
|
||
<taxonomicNameLabel id="A21657C5FFA4FFD3FADEF8C1188FF9D3" pageId="6" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; I–L—
|
||
<taxonomicName id="4C514D2FFFA4FFD3FEB0F8201A94F9D3" ID-CoL="8N9LS" authority="(Bey-Bienko, 1969)" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[272,728,1940,1962]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="yemenica">
|
||
<emphasis id="B925EABEFFA4FFD3FEB0F8201A4EF9D3" box="[272,514,1940,1962]" italics="true" pageId="6" pageNumber="529">Neostylopyga yemenica</emphasis>
|
||
(Bey-Bienko, 1969)
|
||
</taxonomicName>
|
||
. Dotted areas shows membranous parts. Abbreviations:
|
||
<emphasis id="B925EABEFFA4FFD3FABBF8211B32F9B0" italics="true" pageId="6" pageNumber="529">a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.</emphasis>
|
||
—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2E6624FFA5FFD2FF37F8A71D68F98B" httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="7" pageNumber="530" startId="7.[151,249,1811,1833]" targetBox="[219,1368,193,1791]" targetPageId="7">
|
||
<paragraph id="8BEE36ACFFA5FFD2FF37F8A71D68F98B" blockId="7.[151,1436,1811,2035]" pageId="7" pageNumber="530">
|
||
<emphasis id="B925EABEFFA5FFD2FF37F8A7195EF950" bold="true" box="[151,274,1811,1833]" pageId="7" pageNumber="530">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C514D2FFFA5FFD2FEB8F8A71A30F950" ID-CoL="8N97J" authority="Anisyutkin" authorityName="Anisyutkin & Telnov" authorityYear="2018" box="[280,636,1811,1833]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="530" phylum="Arthropoda" rank="species" species="beccalonii" status="sp. nov.">
|
||
<emphasis id="B925EABEFFA5FFD2FEB8F8A71A4FF950" box="[280,515,1811,1833]" italics="true" pageId="7" pageNumber="530">Distylopyga beccalonii</emphasis>
|
||
Anisyutkin
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B925EABEFFA5FFD2FD26F8A71A9AF950" bold="true" box="[646,726,1811,1833]" pageId="7" pageNumber="530">
|
||
<taxonomicNameLabel id="A21657C5FFA5FFD2FD26F8A71A9AF950" box="[646,726,1811,1833]" pageId="7" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male, holotype (A–K) and female, paratype (L–N). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st–4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations:
|
||
<emphasis id="B925EABEFFA5FFD2FC1CF80E195BF98A" italics="true" pageId="7" pageNumber="530">I–X—1st–10th abdominal tergites respectively, a.s., par., pv.</emphasis>
|
||
– see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEE36ACFFAAFFDDFF67FF2C19C0FFD4" blockId="8.[151,1437,151,896]" pageId="8" pageNumber="531">
|
||
In 1988, Roth redescribed
|
||
<emphasis id="B925EABEFFAAFFDDFDA8FF2C1AABFED6" box="[520,743,152,176]" italics="true" pageId="8" pageNumber="531">
|
||
Distilopyga
|
||
<taxonomicName id="4C514D2FFFAAFFDDFD38FF2C1AABFED6" ID-CoL="8MYVM" baseAuthorityName="Krauss" baseAuthorityYear="1903" box="[664,743,152,175]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="species" species="semoni">semoni</taxonomicName>
|
||
</emphasis>
|
||
, transferred this species to
|
||
<taxonomicName id="4C514D2FFFAAFFDDFB97FF2C1CF1FEC9" ID-CoL="8MRQP" authorityName="Rehn" authorityYear="1933" box="[1079,1213,152,176]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925EABEFFAAFFDDFB97FF2C1CF1FEC9" box="[1079,1213,152,176]" italics="true" pageId="8" pageNumber="531">Duchailluia</emphasis>
|
||
</taxonomicName>
|
||
, and synonymized
|
||
<taxonomicName id="4C514D2FFFAAFFDDFF37FF081952FEAD" ID-CoL="8N6YK" authorityName="Bey-Bienko" authorityYear="1965" box="[151,286,188,212]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925EABEFFAAFFDDFF37FF081952FEAD" box="[151,286,188,212]" italics="true" pageId="8" pageNumber="531">Distylopyga</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C514D2FFFAAFFDDFEC7FF081ACBFEAD" ID-CoL="8MRQP" authority="(Roth 1988)" authorityName="Roth" authorityYear="1988" baseAuthorityName="Roth" baseAuthorityYear="1988" box="[359,647,188,213]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925EABEFFAAFFDDFEC7FF0819A1FEAD" box="[359,493,188,212]" italics="true" pageId="8" pageNumber="531">Duchailluia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC04B5DFFAAFFDDFDA0FF081A33FEAD" author="Roth, L. M." box="[512,639,188,213]" pageId="8" pageNumber="531" pagination="297 - 321" refId="ref7807" refString="Roth, L. M. (1988) Some cavernicolous and epigean cockroaches with six new species, and a discussion of the Nocticolidae. Revue suisse de Zoologie, 95, 297 - 321. https: // doi. org / 10.5962 / bhl. part. 79654" type="journal article" year="1988">Roth 1988</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. This redescription was based on specimens from Pak Thong Chai (Prov.
|
||
<collectingRegion id="4995F84EFFAAFFDDFF43FF6B1986FE81" box="[227,458,223,248]" country="Thailand" name="Nakhon Ratchasima" pageId="8" pageNumber="531">Nakhon Ratchasima</collectingRegion>
|
||
, NE
|
||
<collectingCountry id="F346763CFFAAFFDDFDA5FF6B1A22FE81" box="[517,622,223,248]" name="Thailand" pageId="8" pageNumber="531">Thailand</collectingCountry>
|
||
). The male genitalia were only depicted, not described (
|
||
<bibRefCitation id="EFC04B5DFFAAFFDDFB4EFF6B18F6FF65" author="Roth, L. M." pageId="8" pageNumber="531" pagination="297 - 321" refId="ref7807" refString="Roth, L. M. (1988) Some cavernicolous and epigean cockroaches with six new species, and a discussion of the Nocticolidae. Revue suisse de Zoologie, 95, 297 - 321. https: // doi. org / 10.5962 / bhl. part. 79654" type="journal article" year="1988">Roth 1988, fig. 8B</bibRefCitation>
|
||
). Unfortunately, the structures of the male genitalia, as shown in
|
||
<bibRefCitation id="EFC04B5DFFAAFFDDFC03FEB01C63FF65" author="Roth, L. M." box="[931,1071,260,285]" pageId="8" pageNumber="531" pagination="297 - 321" refId="ref7807" refString="Roth, L. M. (1988) Some cavernicolous and epigean cockroaches with six new species, and a discussion of the Nocticolidae. Revue suisse de Zoologie, 95, 297 - 321. https: // doi. org / 10.5962 / bhl. part. 79654" type="journal article" year="1988">Roth (1988)</bibRefCitation>
|
||
, are strongly deformed, so even the main sclerites are not recognizable. The deformation of structures is particularly clearly visible due to the position of apical hook of sclerite L3 of the left phallomere (hook-like left phallomere in term of
|
||
<bibRefCitation id="EFC04B5DFFAAFFDDFB47FEF81892FFF1" author="Roth, L. M." pageId="8" pageNumber="531" pagination="297 - 321" refId="ref7807" refString="Roth, L. M. (1988) Some cavernicolous and epigean cockroaches with six new species, and a discussion of the Nocticolidae. Revue suisse de Zoologie, 95, 297 - 321. https: // doi. org / 10.5962 / bhl. part. 79654" type="journal article" year="1988">Roth (1988, fig. 8B, a)</bibRefCitation>
|
||
directed outwards, whereas this structure is always directed inwards. Thus, the illustration is useless for species identification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34B6527FFAAFFDDFF67FE031B35FDF9" pageId="8" pageNumber="531" type="description">
|
||
<paragraph id="8BEE36ACFFAAFFDDFF67FE0319A2FC44" blockId="8.[151,1437,151,896]" pageId="8" pageNumber="531">The reason for this deformation is obviously the way male genitalia were mounted on slide. In our opinion, the male and female genitalia of cockroaches should not be mounted on slides, since diagnostically important structures become distorted or squashed, and further manipulation of the genitals (e.g. in order to change their position) become not possible.</paragraph>
|
||
<paragraph id="8BEE36ACFFAAFFDDFF67FDF31C3AFCD1" blockId="8.[151,1437,151,896]" pageId="8" pageNumber="531">
|
||
Thus, the available descriptions of
|
||
<taxonomicName id="4C514D2FFFAAFFDDFDEFFDFD1A8CFC26" baseAuthorityName="Krauss" baseAuthorityYear="1903" box="[591,704,584,608]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="species" species="semoni">
|
||
<emphasis id="B925EABEFFAAFFDDFDEFFDFD1A2FFC19" box="[591,611,585,608]" italics="true" pageId="8" pageNumber="531">D</emphasis>
|
||
.
|
||
<emphasis id="B925EABEFFAAFFDDFDD1FDFC1A8CFC26" box="[625,704,584,607]" italics="true" pageId="8" pageNumber="531">semoni</emphasis>
|
||
</taxonomicName>
|
||
do not provide an opportunity to clarify the relationships with its congeners. In order to solve this problem it is necessary to redescribe the
|
||
<typeStatus id="54EA880EFFAAFFDDFC7CFDD91C40FCFC" box="[988,1036,621,645]" pageId="8" pageNumber="531">type</typeStatus>
|
||
material and/or additional material from Cibodas (
|
||
<typeStatus id="54EA880EFFAAFFDDFE9FFD24193DFCD1" box="[319,369,656,680]" pageId="8" pageNumber="531">type</typeStatus>
|
||
locality), with special attention to the structure of the male genitalia.
|
||
</paragraph>
|
||
<paragraph id="8BEE36ACFFAAFFDDFF67FD001B35FDF9" blockId="8.[151,1437,151,896]" pageId="8" pageNumber="531">
|
||
The herewith described
|
||
<taxonomicName id="4C514D2FFFAAFFDDFE41FD001AA5FCB5" authority="Anisyutkin & Telnov, 2018" authorityName="Anisyutkin & Telnov" authorityYear="2018" box="[481,745,692,716]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="species" species="beccalonii" status="sp. nov.">
|
||
<emphasis id="B925EABEFFAAFFDDFE41FD001AA5FCB5" box="[481,745,692,716]" italics="true" pageId="8" pageNumber="531">Distylopyga beccalonii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B925EABEFFAAFFDDFD54FD001B1EFCB4" bold="true" box="[756,850,692,717]" pageId="8" pageNumber="531">
|
||
<taxonomicNameLabel id="A21657C5FFAAFFDDFD54FD001B1EFCB4" box="[756,850,692,717]" pageId="8" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
demonstrates strong differences from
|
||
<taxonomicName id="4C514D2FFFAAFFDDFAB6FD001891FC89" authority="Anisyutkin & Telnov, 2018" authorityName="Anisyutkin & Telnov" authorityYear="2018" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="species" species="ivindo" status="sp. nov.">
|
||
<emphasis id="B925EABEFFAAFFDDFAB6FD001891FC89" italics="true" pageId="8" pageNumber="531">Duchailluia ivindo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21657C5FFAAFFDDFF44FD631971FC96" box="[228,317,727,752]" pageId="8" pageNumber="531" rank="species">
|
||
<emphasis id="B925EABEFFAAFFDDFF44FD6318B2FC89" bold="true" box="[228,254,727,752]" pageId="8" pageNumber="531">sp</emphasis>
|
||
.
|
||
<emphasis id="B925EABEFFAAFFDDFEACFD63197AFC89" bold="true" box="[268,310,727,752]" pageId="8" pageNumber="531">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in the male genitalia. These species have a completely different structure of the following sclerites: L4C (compare
|
||
<figureCitation id="136A2A29FFAAFFDDFEE3FD4819C2FD6D" box="[323,398,764,789]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="8" pageNumber="531">Figs. 3</figureCitation>
|
||
A–C and 3E–G), L2 (compare
|
||
<figureCitation id="136A2A29FFAAFFDDFD50FD481BBEFD6D" box="[752,1010,764,789]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="8" pageNumber="531">Figs. 3B, D and 3G, H</figureCitation>
|
||
), caudal part of "additional elongated sclerite" (compare
|
||
<figureCitation id="136A2A29FFAAFFDDFECEFCAB1AF9FD41" box="[366,693,799,824]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="8" pageNumber="531">
|
||
Figs. 3C,
|
||
<emphasis id="B925EABEFFAAFFDDFE7FFC951A5EFD41" box="[479,530,801,824]" italics="true" pageId="8" pageNumber="531">a.e.s</emphasis>
|
||
. and 3G,
|
||
<emphasis id="B925EABEFFAAFFDDFD22FC951AF9FD41" box="[642,693,801,824]" italics="true" pageId="8" pageNumber="531">a.e.s</emphasis>
|
||
</figureCitation>
|
||
.) and R1G (compare
|
||
<figureCitation id="136A2A29FFAAFFDDFC10FCAB1BBEFD41" box="[944,1010,799,824]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="8" pageNumber="531">Fig. 2</figureCitation>
|
||
A–C and 5A–C). Subsequently, these two species cannot be considered congeneric. In order to avoid multiplication of entities, we prefer to restore the genus
|
||
<taxonomicName id="4C514D2FFFAAFFDDFF7FFCDC192AFDF9" authorityName="Bey-Bienko" authorityYear="1965" box="[223,358,872,896]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925EABEFFAAFFDDFF7FFCDC192AFDF9" box="[223,358,872,896]" italics="true" pageId="8" pageNumber="531">Distylopyga</emphasis>
|
||
</taxonomicName>
|
||
and provisionally ascribe the new species to it.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |