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<document id="BC3C876F946CB759C5E608F94FA47B35" ID-DOI="10.5281/zenodo.176773" ID-GBIF-Dataset="1a670eb6-5f50-42d6-ace0-3e52fd4f4c17" ID-ISSN="1175-5326" ID-Zenodo-Dep="176773" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459839632607" checkinUser="plazi" docAuthor="Golding, Rosemary E., Ponder, Winston F. &amp; Byrne, Maria" docDate="2007" docId="03F887BA99125421118FC0D4FC29FDE0" docLanguage="en" docName="zt01476p050.pdf" docOrigin="Zootaxa 1476" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Phallomedusa solida Martens 1878" docType="treatment" docVersion="10" lastPageNumber="21" masterDocId="FFC1FFC2990054351118C63EFFE7FFA2" masterDocTitle="Taxonomy and anatomy of Amphiboloidea (Gastropoda: Heterobranchia: Archaeopulmonata)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="19" updateTime="1698224205944" updateUser="plazi">
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<mods:title id="2E0AC759164189F55695ED40DF2C171D">Taxonomy and anatomy of Amphiboloidea (Gastropoda: Heterobranchia: Archaeopulmonata)</mods:title>
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<mods:namePart id="E56ED2C0E7E699E3D6B253F63665185B">Golding, Rosemary E.</mods:namePart>
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<mods:namePart id="B797920F5C2415E1485712509DDE90EB">Ponder, Winston F.</mods:namePart>
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<mods:namePart id="C24CB6D5939C2481C969F99BC033D8BF">Byrne, Maria</mods:namePart>
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<mods:date id="93F45E4775379EA3B450F0421D3C630A">2007</mods:date>
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<treatment id="03F887BA99125421118FC0D4FC29FDE0" ID-DOI="http://doi.org/10.5281/zenodo.6240292" ID-GBIF-Taxon="119359975" ID-Zenodo-Dep="6240292" LSID="urn:lsid:plazi:treatment:03F887BA99125421118FC0D4FC29FDE0" httpUri="http://treatment.plazi.org/id/03F887BA99125421118FC0D4FC29FDE0" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="C34B652799125427118FC0D4FD87F8A6" box="[151,608,1770,1796]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BEE36AC99125427118FC0D4FD87F8A6" blockId="18.[151,608,1770,1796]" box="[151,608,1770,1796]" pageId="18" pageNumber="19">
<heading id="D0A681C099125427118FC0D4FD87F8A6" bold="true" box="[151,608,1770,1796]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C514D2F99125427118FC0D4FD87F8A6" ID-CoL="4FRXV" authority="Martens, 1878" authorityName="Martens" authorityYear="1878" box="[151,608,1770,1796]" class="Gastropoda" family="Amphibolidae" genus="Phallomedusa" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B925EABE99125427118FC0D4FD87F8A6" bold="true" box="[151,608,1770,1796]" pageId="18" pageNumber="19">
<emphasis id="B925EABE99125427118FC0D4FE74F8A6" bold="true" box="[151,403,1770,1796]" italics="true" pageId="18" pageNumber="19">Phallomedusa solida</emphasis>
(Martens, 1878)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34B652799125427118FC10EFCE2F845" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BEE36AC99125427118FC10EFC57F8C5" blockId="18.[151,1312,1840,2023]" pageId="18" pageNumber="19">
<taxonomicName id="4C514D2F99125427118FC10EFE93F8E5" box="[151,372,1840,1863]" class="Gastropoda" family="Thiaridae" genus="Ampullacera" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="fragilis">
<emphasis id="B925EABE99125427118FC10EFE93F8E5" box="[151,372,1840,1863]" italics="true" pageId="18" pageNumber="19">Ampullacera fragilis</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFC04B5D99125427109AC10EFD98F8E5" author="Quoy" box="[386,639,1840,1863]" pageId="18" pageNumber="29" refString="Quoy, J. R. C. &amp; Gaimard, J. P. (1832) Voyages de decouvertes de l`Astrolabe pendant les annees 1826 - 1829 sous le commandement de M. J. Dumont d`Urville, Zoologie Vol. 2. J. Tastu, Paris, 686 pp." type="book chapter" year="1832">Quoy &amp; Gaimard, 1832</bibRefCitation>
(in part): 201203, pl. XV, figs. 1012 (not of
<bibRefCitation id="EFC04B5D99125427156DC10EFAF4F8E5" author="Lamarck" box="[1141,1299,1840,1863]" pageId="18" pageNumber="29" refString="Lamarck, J. B. (1822) Histoire Naturelle des Animaux sans Vertebres. Unknown publisher, Paris, 7 volumes." type="book chapter" year="1822">Lamarck 1822</bibRefCitation>
).
<taxonomicName id="4C514D2F99125427118FC16EFE67F8C5" box="[151,384,1872,1895]" class="Gastropoda" family="Amphibolidae" genus="Ampullarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="quoyana">
<emphasis id="B925EABE99125427118FC16EFE67F8C5" box="[151,384,1872,1895]" italics="true" pageId="18" pageNumber="19">Ampullarina quoyana</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFC04B5D991254271096C16EFDFFF8C5" author="Angas" box="[398,536,1872,1895]" pageId="18" pageNumber="28" refString="Angas, G. F. (1867) A list of species of marine Mollusca found in Port Jackson harbour, New South Wales. Proceedings of the Zoological Society of London, 1867, 185 - 233." type="proceedings paper" year="1867">Angas, 1867</bibRefCitation>
: 232 (not of
<bibRefCitation id="EFC04B5D9912542713B8C16EFC44F8C5" author="Potiez" box="[672,931,1872,1895]" pageId="18" pageNumber="29" refString="Potiez, V. L. &amp; Michaud, A. L. (1838) Galerie des mollusques, ou catalogue methodique, descriptif et raisonne des mollusques et coquilles du Museum de Douai. Douai, Paris, 560 pp." type="book" year="1838">Potiez &amp; Michaud 1838</bibRefCitation>
).
</paragraph>
<paragraph id="8BEE36AC99125427118FC14EFCD5F825" blockId="18.[151,1312,1840,2023]" box="[151,818,1904,1927]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB41118299125427118FC14EFCD5F825" box="[151,818,1904,1927]" pageId="18" pageNumber="19">
<taxonomicName id="4C514D2F99125427118FC14EFE53F825" ID-CoL="CZDP" authority="Martens" authorityName="Martens" box="[151,436,1904,1927]" class="Gastropoda" family="Amphibolidae" genus="Amphibola" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B925EABE99125427118FC14EFEB4F825" box="[151,339,1904,1927]" italics="true" pageId="18" pageNumber="19">Amphibola solida</emphasis>
Martens
</taxonomicName>
, in
<treatmentCitation id="0AF010BD9912542710C2C14EFD68F825" author="Schacko" box="[474,655,1904,1927]" page="1" pageId="18" pageNumber="19" year="1878">
<bibRefCitation id="EFC04B5D9912542710C2C14EFD9CF825" author="Schacko" box="[474,635,1904,1927]" pageId="18" pageNumber="29" refString="Schacko, G. (1878) Die Zungenbewaffnung der Gattung Amphibola. Jahrbuch der Deutschen Malakozoologischen Gesellschaft, 5, 1 - 9." type="journal article" year="1878">Schacko, 1878</bibRefCitation>
: 1
</treatmentCitation>
9, pl. I, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AC99125427118FC1AEFC8CF805" blockId="18.[151,1312,1840,2023]" box="[151,875,1936,1959]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB41118299125427118FC1AEFC8CF805" box="[151,875,1936,1959]" pageId="18" pageNumber="19">
<taxonomicName id="4C514D2F99125427118FC1AEFEB8F805" box="[151,351,1936,1959]" class="Gastropoda" family="Amphibolidae" genus="Amphibola" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="fragilis">
<emphasis id="B925EABE99125427118FC1AEFEB8F805" box="[151,351,1936,1959]" italics="true" pageId="18" pageNumber="19">Amphibola fragilis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AF010BD991254271075C1AEFDD2F805" author="Bouvier" box="[365,565,1936,1959]" page="146" pageId="18" pageNumber="19" year="1892">
<bibRefCitation id="EFC04B5D991254271075C1AEFDE4F805" author="Bouvier" box="[365,515,1936,1959]" pageId="18" pageNumber="28" refString="Bouvier, E. L. (1892) Sur l'organisation des Amphiboles. Bulletin de la Societe Philomathique de Paris, 4, 146 - 153." type="journal article" year="1892">Bouvier 1892</bibRefCitation>
: 146
</treatmentCitation>
153 (not of
<bibRefCitation id="EFC04B5D9912542713D8C1AEFCB8F805" author="Lamarck" box="[704,863,1936,1959]" pageId="18" pageNumber="29" refString="Lamarck, J. B. (1822) Histoire Naturelle des Animaux sans Vertebres. Unknown publisher, Paris, 7 volumes." type="book chapter" year="1822">Lamarck 1822</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AC99125427118FC18EFCE2F845" blockId="18.[151,1312,1840,2023]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB41118299125427118FC18EFCE2F845" pageId="18" pageNumber="19">
<taxonomicName id="4C514D2F99125427118FC18EFEBAF865" box="[151,349,1968,1991]" class="Gastropoda" family="Amphibolidae" genus="Salinator" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="quoyana">
<emphasis id="B925EABE99125427118FC18EFEBAF865" box="[151,349,1968,1991]" italics="true" pageId="18" pageNumber="19">Salinator quoyana</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AF010BD991254271073C18EFD47F865" author="Cotton" box="[363,672,1968,1991]" page="150" pageId="18" pageNumber="19" year="1932">
<bibRefCitation id="EFC04B5D991254271073C18EFD89F865" author="Cotton" box="[363,622,1968,1991]" pageId="18" pageNumber="28" refString="Cotton, B. C. &amp; Godfrey, F. K. (1932) South Australian shells. South Australian Naturalist, 13, 127 - 176." type="journal article" year="1932">Cotton &amp; Godfrey 1932</bibRefCitation>
: 150
</treatmentCitation>
(not of
<bibRefCitation id="EFC04B5D9912542713EDC18EFC1FF865" author="Potiez" box="[757,1016,1968,1991]" pageId="18" pageNumber="29" refString="Potiez, V. L. &amp; Michaud, A. L. (1838) Galerie des mollusques, ou catalogue methodique, descriptif et raisonne des mollusques et coquilles du Museum de Douai. Douai, Paris, 560 pp." type="book" year="1838">Potiez &amp; Michaud 1838</bibRefCitation>
).
<taxonomicName id="4C514D2F99125427118FC1EEFEA5F845" ID-CoL="79B84" box="[151,322,2000,2023]" class="Gastropoda" family="Amphibolidae" genus="Salinator" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B925EABE99125427118FC1EEFEA5F845" box="[151,322,2000,2023]" italics="true" pageId="18" pageNumber="19">Salinator solida</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AF010BD991254271048C1EEFDF1F845" author="Woolacott" box="[336,534,2000,2023]" page="2" pageId="18" pageNumber="19" year="1945">
<bibRefCitation id="EFC04B5D991254271048C1EEFE1AF845" author="Woolacott" box="[336,509,2000,2023]" pageId="18" pageNumber="30" refString="Woolacott, L. (1945) The establishment of Salinator fragilis Lamarck and Salinator solida von Marten as two distinct species in Australia. Proceedings of the Royal Zoological Society of N. S. W., 1944 - 45, 35 - 38." type="proceedings paper" year="1945">Woolacott 1945</bibRefCitation>
: 2
</treatmentCitation>
, pl. III, figs. 1, 2, 7, 9.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B652799135421118FC6A9FCB0FEA0" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BEE36AC99135426118FC6A9FBD3FF10" blockId="19.[151,1437,151,2020]" box="[151,1076,151,178]" pageId="19" pageNumber="20">
<emphasis id="B925EABE99135426118FC6A9FEA3FF13" bold="true" box="[151,324,151,177]" pageId="19" pageNumber="20">
<typeStatus id="54EA880E99135426118FC6A9FF35FF13" box="[151,210,151,177]" pageId="19" pageNumber="20">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="54EA880E99135426104DC6A6FE25FF10" box="[341,450,152,178]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
(
<figureCitation id="136A2A299913542610CAC6A6FDFFFF10" box="[466,536,152,178]" captionStart="FIGURE 2" captionStartId="31.[151,255,1555,1579]" captionTargetBox="[151,1436,167,1526]" captionTargetId="figure@31.[151,1436,167,1528]" captionTargetPageId="31" captionText="FIGURE 2. Shell morphology of species in Amphibolidae, Phallomedusidae fam. nov. and Maningrididae fam. nov .. A. Amphibola crenata (Martyn, 1786), (AMS C. 446522). B D. Salinator fragilis (Lamarck, 1822), B. Lectotype (MHN 1094 / 1), C. Albany, Western Australia (AMS C. 446486), D. Cambridge, Tasmania (AMS C. 442291). E. Salinator tecta sp. nov .. Holotype, Narrabeen Lakes, Sydney (AMS C. 446774). F. Salinator rhamphidia sp. nov .. Holotype, Botany Bay, Sydney (AMS C. 446516). G. Salinator rosacea sp. nov .. Holotype, Elizabeth River, Darwin (AMS C. 446519). H, I. Phallomedusa solida (Martens, 1878), H. Holotype (ZMB 22207), I. Lake Illawarra, New South Wales (AMS C. 442239). J. Phallomedusa austrina sp. nov .. Holotype, Stanley, Tasmania (AMS C. 446510). K. Lactiforis tropicalis sp. nov .. Holotype, Broome, Western Australia (AMS C. 446775). L. Lactiforis takii (Kuroda, 1928). Ariake Sea, Japan (AMS C. 446490). M. Naranjia cf. swatowensis (Yen, 1939). Tsim Bei Tsui, Hong Kong (AMS C. 446511). N. Maningrida arnhemensis sp. nov .. Holotype, Maningrida, Northern Territory (AMS C. 446485). Scale bar = 2 mm." httpUri="https://zenodo.org/record/176775/files/figure.png" pageId="19" pageNumber="20">Fig. 2</figureCitation>
H) from Australia, Robillard (
<collectionCode id="ED40AE69991354261291C6A6FC2AFF10" box="[905,973,152,178]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="19" pageNumber="20">ZMB</collectionCode>
, 22207).
</paragraph>
<paragraph id="8BEE36AC9913542611DEC681FAB7FDCE" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
<emphasis id="B925EABE9913542611DEC681FDE0FF7B" bold="true" box="[198,519,191,217]" pageId="19" pageNumber="20">Other material examined</emphasis>
:
<collectingCountry id="F346763C991354261302C6FEFD6EFF78" box="[538,649,192,218]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
, Queensland: Port Douglas, Dicksons Inlet, in mangroves,
<date id="FFEF106C991354261443C6FEFF33FEA0" pageId="19" pageNumber="20" value="1977-06-03">3 Jun 1977</date>
, I. Loch, P.H. Colman and R. Creese (AMS C.400788); Proserpine River, opposite Flying Fox Island, in mangroves,
<date id="FFEF106C991354261033C72EFE21FE88" box="[299,454,272,298]" pageId="19" pageNumber="20" value="1975-05-01">1 May 1975</date>
, W.F. Ponder (AMS C.446518).
<collectingCountry id="F346763C991354261247C72EFC29FE88" box="[863,974,272,298]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
, New South
<collectingCountry id="F346763C99135426156CC72EFB58FE88" box="[1140,1215,272,298]" name="United Kingdom" pageId="19" pageNumber="20">Wales</collectingCountry>
: Brisbane Waters, Fagan's Bay, on mud in saltmarsh,
<date id="FFEF106C991354261336C706FD20FEF0" box="[558,711,312,338]" pageId="19" pageNumber="20" value="2003-03-10">10 Mar 2003</date>
, R.E. Golding, W.F. Ponder and H. Fukuda (AMS C.442245); South side of Narrabeen Lakes, on sandy mud amongst reeds and mangroves,
<date id="FFEF106C991354261527C75EFB15FED8" box="[1087,1266,352,378]" pageId="19" pageNumber="20" value="2004-03" valueMax="2004-09-31" valueMin="2004-03-00">MarSep 2004</date>
, R.E. Golding and M. Hill (AMS C.442241, C.442277, C.442286); Lane Cove River, Boronia Park, on mud in open mangrove forest at low tide,
<date id="FFEF106C9913542610D2C78EFDD9FE68" box="[458,574,432,458]" pageId="19" pageNumber="20" value="2003-12">Dec 2003</date>
<date id="FFEF106C991354261356C78EFD24FE68" box="[590,707,432,458]" pageId="19" pageNumber="20" value="2004-09">Sep 2004</date>
, R.E. Golding, M. Hill and P.M. Golding (AMS C.442276, C.442284, C.442266); Parramatta River, Homebush Bay, on mud amongst dense mangroves,
<date id="FFEF106C9913542615F7C7E6FA86FE50" box="[1263,1377,472,498]" pageId="19" pageNumber="20" value="2003-12">Dec 2003</date>
<date id="FFEF106C991354261468C7E6FF32FDB8" pageId="19" pageNumber="20" value="2004-02">Feb 2004</date>
, R.E. Golding (AMS C.442247, C.442259, C.442262); Lake Illawarra, Kully Bay, creek near lake with sparse saltmarsh vegetation,
<date id="FFEF106C9913542610FFC414FD88FDE6" box="[487,623,554,580]" pageId="19" pageNumber="20" value="2004-03-03">3Mar 2004</date>
, R.E. Golding and P.M. Golding (AMS C.442239).
<collectingCountry id="F346763C9913542615D5C414FADCFDE6" box="[1229,1339,554,580]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
, Tasmania: Stanley, Sawyer Bay, East Inlet, damp mudflats at low tide,
<date id="FFEF106C991354261290C46CFBF8FDCE" box="[904,1055,594,620]" pageId="19" pageNumber="20" value="2004-04-20">20 Apr 2004</date>
, W.F. Ponder (C.446517).
</paragraph>
<paragraph id="8BEE36AC9913542611DEC444FB8EFD46" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
<emphasis id="B925EABE9913542611DEC444FE93FD36" bold="true" box="[198,372,634,660]" pageId="19" pageNumber="20">Redescription</emphasis>
: Shell (
<figureCitation id="136A2A299913542610CBC444FDC2FD36" box="[467,549,634,660]" captionStart="FIGURE 2" captionStartId="31.[151,255,1555,1579]" captionTargetBox="[151,1436,167,1526]" captionTargetId="figure@31.[151,1436,167,1528]" captionTargetPageId="31" captionText="FIGURE 2. Shell morphology of species in Amphibolidae, Phallomedusidae fam. nov. and Maningrididae fam. nov .. A. Amphibola crenata (Martyn, 1786), (AMS C. 446522). B D. Salinator fragilis (Lamarck, 1822), B. Lectotype (MHN 1094 / 1), C. Albany, Western Australia (AMS C. 446486), D. Cambridge, Tasmania (AMS C. 442291). E. Salinator tecta sp. nov .. Holotype, Narrabeen Lakes, Sydney (AMS C. 446774). F. Salinator rhamphidia sp. nov .. Holotype, Botany Bay, Sydney (AMS C. 446516). G. Salinator rosacea sp. nov .. Holotype, Elizabeth River, Darwin (AMS C. 446519). H, I. Phallomedusa solida (Martens, 1878), H. Holotype (ZMB 22207), I. Lake Illawarra, New South Wales (AMS C. 442239). J. Phallomedusa austrina sp. nov .. Holotype, Stanley, Tasmania (AMS C. 446510). K. Lactiforis tropicalis sp. nov .. Holotype, Broome, Western Australia (AMS C. 446775). L. Lactiforis takii (Kuroda, 1928). Ariake Sea, Japan (AMS C. 446490). M. Naranjia cf. swatowensis (Yen, 1939). Tsim Bei Tsui, Hong Kong (AMS C. 446511). N. Maningrida arnhemensis sp. nov .. Holotype, Maningrida, Northern Territory (AMS C. 446485). Scale bar = 2 mm." httpUri="https://zenodo.org/record/176775/files/figure.png" pageId="19" pageNumber="20">Figs. 2</figureCitation>
H, I): Solid, opaque, spire relatively tall, conical, shell diameter to
<quantity id="4CA99B49991354261459C444FA70FD36" box="[1345,1431,634,660]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="mm" value="20.0">20 mm</quantity>
, axial striae indistinct. Whorls rounded, lacking shoulder. Outer lip of aperture slightly thickened at base. Exterior white with brown, narrow zigzag lines on all whorls. Interior of aperture white.
</paragraph>
<paragraph id="8BEE36AC9913542611DEC4CCFF29FC26" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
Operculum (
<figureCitation id="136A2A29991354261044C4CCFE49FCAE" box="[348,430,754,780]" captionStart="FIGURE 3" captionStartId="32.[151,255,1167,1191]" captionTargetBox="[151,1435,194,1143]" captionTargetId="figure@32.[151,1436,194,1143]" captionTargetPageId="32" captionText="FIGURE 3. Internal surfaces of opercula. A. Amphibola crenata (Martyn, 1786). B. Salinator fragilis (Lamarck, 1822). C. Lactiforis tropicalis sp. nov .. D. Phallomedusa solida (Martens, 1878). E. Maningrida arnhemensis sp. nov .. Scale bar = 2 mm." httpUri="https://zenodo.org/record/176776/files/figure.png" pageId="19" pageNumber="20">Figs. 3</figureCitation>
D, 4A): Thick, dark brown, corneous, elliptical, columellar edge straight, outer edge curved forming rounded point in upper left corner (viewed from exterior). Nucleus paucispiral, eccentric. Prominent hollow keel curving around nucleus on interior of operculum, containing muscular projection from foot.
</paragraph>
<paragraph id="8BEE36AC9913542611DEC5ACFE99FC5E" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
External morphology (
<figureCitation id="136A2A299913542610D7C5ACFDF8FC0E" box="[463,543,914,940]" captionStart="FIGURE 5" captionStartId="34.[151,255,790,814]" captionTargetBox="[184,1435,247,754]" captionTargetId="figure@34.[151,1436,197,764]" captionTargetPageId="34" captionText="FIGURE 5. Photographs of live animals. A. Phallomedusa solida (Martens, 1878). B. Two specimens of Salinator rhamphidia sp. nov .. Scale bar = 5 mm." httpUri="https://zenodo.org/record/176778/files/figure.png" pageId="19" pageNumber="20">Figs. 5</figureCitation>
A, 6C): Head-foot dark grey in preserved specimens, protruding only a short way beyond shell when animal is crawling; diffuse black pigment across snout. Eyes at central base of small, triangular tentacles.
</paragraph>
<paragraph id="8BEE36AC9913542611DEC234FF12FB3E" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
Mantle organs (
<figureCitation id="136A2A29991354261067C234FE21FB86" box="[383,454,1034,1060]" captionStart="FIGURE 7" captionStartId="36.[151,255,1775,1799]" captionTargetBox="[221,1371,204,1742]" captionTargetId="figure@36.[151,1436,194,1751]" captionTargetPageId="36" captionText="FIGURE 7. Morphology of organs in the mantle cavity. A. Amphibola crenata (Martyn, 1786). B. Salinator fragilis (Lamarck, 1822). C. Lactiforis tropicalis sp. nov .. D. Naranjia cf. swatowensis (Yen, 1939). E. Phallomedusa solida (Martens, 1878). F. Maningrida arnhemensis sp. nov .. Scale bar = 1 mm." httpUri="https://zenodo.org/record/176780/files/figure.png" pageId="19" pageNumber="20">Fig. 7</figureCitation>
E): Narrow, opposed ciliary tracts restricted to exhalant canal in mantle cavity; roof of mantle cavity with numerous blood vessels. Hypobranchial gland at right anterior mantle cavity roof pyriform, flat, light brown with dark brown flecks in fresh and formalin-preserved specimens, outer surface smooth.
</paragraph>
<paragraph id="8BEE36AC9913542611DEC294FD97FB66" blockId="19.[151,1437,151,2020]" box="[198,624,1194,1220]" pageId="19" pageNumber="20">
Digestive system: As for
<taxonomicName id="4C514D2F9913542610E8C295FD8EFB66" box="[496,617,1195,1220]" class="Gastropoda" family="Amphibolidae" genus="Amphibola" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="crenata">
<emphasis id="B925EABE9913542610E8C295FD8EFB66" box="[496,617,1195,1220]" italics="true" pageId="19" pageNumber="20">A. crenata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEE36AC9913542611DEC2ECFE6DFA2E" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
Radula (
<figureCitation id="136A2A29991354261036C2ECFE76FB4E" box="[302,401,1234,1260]" captionStart="FIGURE 11" captionStartId="40.[151,255,1454,1478]" captionTargetBox="[151,1435,194,1430]" captionTargetId="figure@40.[151,1436,194,1430]" captionTargetPageId="40" captionText="FIGURE 11. SEM photographs of radulae of Phallomedusa gen. nov. and Maningrida gen. nov. species, showing halfradula and detail of central and lateral teeth. A, B. Phallomedusa solida (Martens, 1878). C, D. Phallomedusa austrina sp. nov .. E, F. Maningrida arnhemensis sp. nov .. A, C, E. Scale bar = 50 µm, B, D, F. Scale bar = 20 µm." httpUri="https://zenodo.org/record/176784/files/figure.png" pageId="19" pageNumber="20">Figs. 11</figureCitation>
A, B): Each row with central tooth, inner and outer lateral teeth and, on each side, approximately 25 marginal teeth. Central tooth with five wide cusps, mesocone enlarged, rectangular; inner lateral teeth unicuspid, narrow; outer lateral teeth equal in size to central tooth, with three wide cusps increasing in size towards centre of radula; marginal teeth elongate, unicuspid, wide, increasing in length towards outer edge of radula.
</paragraph>
<paragraph id="8BEE36AC9913542611DEC3A4FF28F98E" blockId="19.[151,1437,151,2020]" pageId="19" pageNumber="20">
Central nervous system (
<figureCitation id="136A2A299913542610E9C3A4FDAEFA16" box="[497,585,1434,1460]" captionStart="FIGURE 12" captionStartId="41.[151,255,1235,1259]" captionTargetBox="[189,1399,186,1196]" captionTargetId="figure@41.[151,1436,160,1211]" captionTargetPageId="41" captionText="FIGURE 12. Central nervous systems. A. Amphibola crenata (Martyn, 1786). B. Salinator fragilis (Lamarck, 1822). C. Lactiforis tropicalis sp. nov .. D. Lactiforis takii (Kuroda, 1928). E. Naranjia cf. swatowensis (Yen, 1939). F. Phallomedusa solida (Martens, 1878). G. Maningrida arnhemensis sp. nov .. Scale bar = 500 µm. Subcerebral and parapedal commissures are indicated on the figures for A. crenata, S. fragilis and P. s o l i d a, but not on the figures for other species as the presence of these structures is unconfirmed." httpUri="https://zenodo.org/record/176785/files/figure.png" pageId="19" pageNumber="20">Fig. 12</figureCitation>
F): Pleural ganglia separated from adjacent cerebral, pedal and parietal ganglia by short connectives. Procerebrum, parapedal and subcerebral commissures present. Right parietal ganglion slightly larger than left, right parietal-visceral commissure shorter than left parietal-visceral commissure.
</paragraph>
<paragraph id="8BEE36AC9913542111DEC004FCB0FEA0" blockId="19.[151,1437,151,2020]" lastBlockId="20.[151,1437,152,578]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Reproductive system (
<figureCitation id="136A2A299913542610D3C004FDCCF9F6" box="[459,555,1594,1620]" captionStart="FIGURE 13" captionStartId="42.[151,244,1729,1750]" captionTargetBox="[191,1378,163,1695]" captionTargetId="figure@42.[182,1406,141,1711]" captionTargetPageId="42" captionText="FIGURE 13. Reproductive systems of Amphibolidae, Phallomedusidae fam. nov. and Maningrididae fam. nov .. These figures show the entire reproductive system and provide an anatomical map for the structures illustrated in Figs. 13 15. A. Amphibola crenata (Martyn, 1786), with spermovipositor sheath dissected open to show internal structures. Scale bar = 2 mm. B. Salinator tecta sp. nov., with spermovipositor sheath intact. Scale bar = 1 mm. C. Phallomedusa solida (Martens, 1878), with preputium dissected open to show penis. Scale bar = 2 mm. D. Maningrida arnhemensis sp. nov., with genital atrium dissected open to show internal structure. Parallel shaded lines indicate that ducts are embedded in the body wall. Scale bar = 1 mm." httpUri="https://zenodo.org/record/176786/files/figure.png" pageId="19" pageNumber="20">Figs. 13</figureCitation>
C, 15A, B, 17A, B): Ovotestis composed of numerous lobes uniting to form single acinule duct. Seminal vesicle diverticulum from upper hermaphrodite duct, posterior to junction with acinule duct. Hermaphrodite duct coiling anteriorly, swollen with pale pink, iridescent sperm. Numerous (&gt;10) seminal receptacles present as yellow, spherical projections from hermaphrodite duct. Spermoviduct embedded in right body wall distally, emerging to divide into muscular oviduct/vagina and long vas deferens. Muscular vagina long, with thick, folded walls and two lateral bulges proximally, narrowing distally. Small flagellum of vas deferens extending beyond junction with oviduct; vas deferens very long, coiled, yellow to orange in fresh and formalin-preserved specimens, muscular exterior lined internally with tall narrow prostatic cells containing spherical secretory vesicles. Complex penis with more spiral flange coiling more than three times around penis; spiral base with unciliated external sperm groove between spirally coiled lateral flanges. Penis attached near genital aperture, retractor muscle inserting into base of penis. Flange forming large shield distally and dividing into numerous moderately long tentacle-like appendages. Penial appendages and inner surface of spiral flange with long, forked, chitinous papilla-like microsculpture. Egg mass narrow, cylindrical string deposited in coil on surface of substratum.
</paragraph>
</subSubSection>
<subSubSection id="C34B65279914542111DEC731FCA3FED8" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BEE36AC9914542111DEC731FCA3FED8" blockId="20.[151,1437,152,578]" pageId="20" pageNumber="21">
<emphasis id="B925EABE9914542111DEC731FE18FE8B" bold="true" box="[198,511,271,297]" pageId="20" pageNumber="21">Distribution and habitat</emphasis>
(
<figureCitation id="136A2A2999145421130DC72EFDBAFE88" box="[533,605,272,298]" captionStart="FIGURE 1" captionStartId="30.[151,255,1168,1192]" captionTargetBox="[163,1431,197,1156]" captionTargetId="figure@30.[163,1448,197,1158]" captionTargetPageId="30" captionText="FIGURE 1. Distributions of amphiboloidean species determined from the present study and previous reports. Species in bold are examined in this paper." httpUri="https://zenodo.org/record/176774/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
): East and south-east coast of
<collectingCountry id="F346763C9914542112FAC72EFBB6FE88" box="[994,1105,272,298]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
, from north Queensland to South
<collectingCountry id="F346763C9914542111FEC706FEB5FEF0" box="[230,338,312,338]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
and the north-west of Tasmania (based on AMS collections). Found in upper littoral mangrove, saltmarsh and mudflat habitat, often very abundant.
</paragraph>
</subSubSection>
<subSubSection id="C34B65279914542111DEC7B9FC29FDE0" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BEE36AC9914542111DEC7B9FC29FDE0" blockId="20.[151,1437,152,578]" pageId="20" pageNumber="21">
<emphasis id="B925EABE9914542111DEC7B9FEA5FE03" bold="true" box="[198,322,391,417]" pageId="20" pageNumber="21">Remarks:</emphasis>
The shell of
<emphasis id="B925EABE9914542110F9C7B6FDA4FE03" box="[481,579,392,417]" italics="true" pageId="20" pageNumber="21">P. s o l i d a</emphasis>
was first figured by
<bibRefCitation id="EFC04B5D991454211258C7B6FB81FE00" author="Quoy" box="[832,1126,392,418]" pageId="20" pageNumber="29" refString="Quoy, J. R. C. &amp; Gaimard, J. P. (1832) Voyages de decouvertes de l`Astrolabe pendant les annees 1826 - 1829 sous le commandement de M. J. Dumont d`Urville, Zoologie Vol. 2. J. Tastu, Paris, 686 pp." type="book chapter" year="1832">Quoy &amp; Gaimard (1832)</bibRefCitation>
as a variety of
<taxonomicName id="4C514D2F991454211439C7B6FA74FE03" box="[1313,1427,392,417]" class="Gastropoda" family="Amphibolidae" genus="Amphibola" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="fragilis">
<emphasis id="B925EABE991454211439C7B6FA74FE03" box="[1313,1427,392,417]" italics="true" pageId="20" pageNumber="21">S. fragilis</emphasis>
</taxonomicName>
; although it is not clear where their figured specimen was collected, it appears to be similar to material from south-east
<collectingCountry id="F346763C99145421100DC7E6FE63FE50" box="[277,388,472,498]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
. The species has also been incorrectly attributed to
<taxonomicName id="4C514D2F9914542112F8C7E6FB85FE53" box="[992,1122,472,497]" class="Gastropoda" family="Amphibolidae" genus="Salinator" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="quoyana">
<emphasis id="B925EABE9914542112F8C7E6FB85FE53" box="[992,1122,472,497]" italics="true" pageId="20" pageNumber="21">S. quoyana</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B5D991454211596C7E6FAC8FE50" author="Angas" box="[1166,1327,472,498]" pageId="20" pageNumber="28" refString="Angas, G. F. (1867) A list of species of marine Mollusca found in Port Jackson harbour, New South Wales. Proceedings of the Zoological Society of London, 1867, 185 - 233." type="proceedings paper" year="1867">Angas (1867)</bibRefCitation>
and
<bibRefCitation id="EFC04B5D991454211470C7E6FE7DFDB8" author="Cotton" pageId="20" pageNumber="28" refString="Cotton, B. C. &amp; Godfrey, F. K. (1932) South Australian shells. South Australian Naturalist, 13, 127 - 176." type="journal article" year="1932">Cotton &amp; Godfrey (1932)</bibRefCitation>
from Adelaide (South
<collectingCountry id="F346763C9914542113B4C43EFCFAFDB8" box="[684,797,512,538]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
).
<taxonomicName id="4C514D2F991454211228C43EFBCFFDBB" box="[816,1064,512,537]" class="Gastropoda" family="Amphibolidae" genus="Phallomedusa" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B925EABE991454211228C43EFBCFFDBB" box="[816,1064,512,537]" italics="true" pageId="20" pageNumber="21">Phallomedusa solida</emphasis>
</taxonomicName>
is dominant in saltmarshes and at the back edge of mangrove habitat along the east coast of
<collectingCountry id="F346763C991454211242C416FC2EFDE0" box="[858,969,552,578]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>