363 lines
57 KiB
XML
363 lines
57 KiB
XML
<document id="ACC59EB80E9F21CD6AA17F54971A49AE" ID-CLB-Dataset="45880" ID-DOI="10.5281/zenodo.207321" ID-GBIF-Dataset="514604fc-bca8-4ed7-8783-b1b38596c9ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="207321" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460415105389" checkinUser="plazi" docAuthor="Bonato, Lucio & Minelli, Alessandro" docDate="2011" docId="03F887ADFFAEFFF0CCDAB57D35C7FE00" docLanguage="en" docName="zt03114p049.pdf" docOrigin="Zootaxa 3114" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Geophilus arenarius Meinert 1870" docType="treatment" docVersion="10" lastPageNumber="44" masterDocId="FFC1FFD5FFAFFFF5CC4DB252346BFF87" masterDocTitle="Geophilus arenarius, a long-misunderstood species in the still unresolved carpophagus species-complex (Chilopoda: Geophilidae)" masterLastPageNumber="49" masterPageNumber="40" pageNumber="41" updateTime="1698328489088" updateUser="ExternalLinkService">
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<mods:title id="2A40867D7B2AA3F390C113484F001819">Geophilus arenarius, a long-misunderstood species in the still unresolved carpophagus species-complex (Chilopoda: Geophilidae)</mods:title>
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<mods:namePart id="6090B152081B964DD20BFF85E00B94A9">Bonato, Lucio</mods:namePart>
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<mods:namePart id="56BC2010E8E31CCB537C4C1F193151A6">Minelli, Alessandro</mods:namePart>
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<mods:date id="077263AA4DB4989AEA62E82B38CE6EE7">2011</mods:date>
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<mods:detail id="DB9CD8EE6001774324ED102E84481045" type="volume">
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<mods:number id="9D94E7F27E53E02B90A8DA639752915E">3114</mods:number>
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<mods:identifier id="E13F9F93393E071B807CD102B8E1887E" type="DOI">10.5281/zenodo.207321</mods:identifier>
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<treatment id="03F887ADFFAEFFF0CCDAB57D35C7FE00" ID-DOI="http://doi.org/10.5281/zenodo.5689533" ID-GBIF-Taxon="119424694" ID-Zenodo-Dep="5689533" LSID="urn:lsid:plazi:treatment:03F887ADFFAEFFF0CCDAB57D35C7FE00" httpUri="http://treatment.plazi.org/id/03F887ADFFAEFFF0CCDAB57D35C7FE00" lastPageId="5" lastPageNumber="44" pageId="1" pageNumber="41">
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<subSubSection id="C34B6530FFAEFFF7CCDAB57D377EFF53" lastPageId="2" lastPageNumber="42" pageId="1" pageNumber="41" type="nomenclature">
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<paragraph id="8BEE36BBFFAEFFF4CCDAB57D362EF8CE" blockId="1.[151,581,1839,1865]" box="[151,581,1839,1865]" pageId="1" pageNumber="41">
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<heading id="D0A681D7FFAEFFF4CCDAB57D362EF8CE" bold="true" box="[151,581,1839,1865]" fontSize="11" level="1" pageId="1" pageNumber="41" reason="1">
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<taxonomicName id="4C514D38FFAEFFF4CCDAB57D362EF8CE" ID-CoL="3FQZD" authority="Meinert, 1870" authorityName="Meinert" authorityYear="1870" box="[151,581,1839,1865]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="arenarius">
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<emphasis id="B925EAA9FFAEFFF4CCDAB57D362EF8CE" bold="true" box="[151,581,1839,1865]" pageId="1" pageNumber="41">
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<emphasis id="B925EAA9FFAEFFF4CCDAB57D35E4F8CE" bold="true" box="[151,399,1839,1865]" italics="true" pageId="1" pageNumber="41">Geophilus arenarius</emphasis>
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<bibRefCitation id="EFC04B4AFFAEFFF4CDDBB57D362EF8CE" author="Meinert" box="[406,581,1839,1865]" pageId="1" pageNumber="41" refString="Meinert, F. (1870) Myriapoda Musaei Hauniensis. Bitrag til Myriapodernes morphologi og systematik. I. Geophile. Naturhistorisk Tidsskrift, 7, 1 - 128." type="journal article" year="1870">Meinert, 1870</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BEE36BBFFAEFFF7CCDAB525377EFF53" blockId="1.[151,1436,1910,2007]" lastBlockId="2.[151,1436,151,679]" lastPageId="2" lastPageNumber="42" pageId="1" pageNumber="41">
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<emphasis id="B925EAA9FFAEFFF4CCDAB52535EEF808" bold="true" box="[151,389,1910,1935]" pageId="1" pageNumber="41">Full list of citations.</emphasis>
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<bibRefCitation id="EFC04B4AFFAEFFF4CDC0B5243642F809" author="Meinert" box="[397,553,1910,1934]" pageId="1" pageNumber="41" refString="Meinert, F. (1870) Myriapoda Musaei Hauniensis. Bitrag til Myriapodernes morphologi og systematik. I. Geophile. Naturhistorisk Tidsskrift, 7, 1 - 128." type="journal article" year="1870">Meinert 1870</bibRefCitation>
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: 78 (original description, and in key).
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<bibRefCitation id="EFC04B4AFFAEFFF4CF94B52430F6F809" author="Tomosvary" box="[985,1181,1910,1934]" pageId="1" pageNumber="41" refString="Tomosvary, E. (1880) Beitrag zur Kenntnis der Myriopoden Ungarns. I. Die Chilopoden. Zoologischer Anzeiger, 3, 617 - 619." type="journal article" year="1880">Tömösváry 1880</bibRefCitation>
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: 618.
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<bibRefCitation id="EFC04B4AFFAEFFF4C8ACB5243105F809" author="Daday" box="[1249,1390,1910,1934]" pageId="1" pageNumber="41" refString="Daday, E. (1889) Myriopoda regni Hungariae. Kiralyi Magyar Termeszettudomanyi Tarsulat, Budapest, 104 pp." type="book" year="1889">Daday 1889</bibRefCitation>
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: 24, 28, 30, 33, 37, 85 (in key), 86 (description, and new records).
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<bibRefCitation id="EFC04B4AFFAEFFF4CF17B5C8306BF835" author="Verhoeff" box="[858,1024,1946,1970]" pageId="1" pageNumber="41" refString="Verhoeff, C. (1896) Geophiliden und Scolopendriden aus Portugal und Tabelle europaischer Geophilus - Arten. Zoologischer Anzeiger, 19, 74 - 79, 81 - 89." type="journal article" year="1896">Verhoeff 1896</bibRefCitation>
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||
: 88 (in key).
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<bibRefCitation id="EFC04B4AFFAEFFF4C8D5B5C93146F835" author="Attems" box="[1176,1325,1946,1971]" pageId="1" pageNumber="41" refString="Attems, C. (1903) Synopsis der Geophiliden. Zoologische Jahrbucher, Abteilung fur Systematik, 18, 155 - 302." type="journal article" year="1903">Attems 1903</bibRefCitation>
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: 169, 221 (in key), 228.
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<bibRefCitation id="EFC04B4AFFAEFFF4CD78B5EC35AEF851" author="Depoli" box="[309,453,1982,2006]" pageId="1" pageNumber="41" refString="Depoli, G. (1907) Analisi dei miriapodi componenti la fauna fiumana. Rivista Italiana di Scienze Naturali e Bollettino del Naturalista, 27, 23 - 27, 85 - 93." type="journal article" year="1907">Depoli 1907</bibRefCitation>
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: 25, 85, 86, 89.
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<bibRefCitation id="EFC04B4AFFAEFFF4CE34B5EC3753F851" author="Brolemann" box="[633,824,1982,2006]" pageId="1" pageNumber="41" refString="Brolemann, H. W. (1921) Liste des myriapodes signales dans le nord de l'Afrique. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 1, 99 - 110." type="journal article" year="1921">Brolemann 1921</bibRefCitation>
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: 103.
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<bibRefCitation id="EFC04B4AFFAEFFF4CF31B5ED3077F851" author="Attems" box="[892,1052,1982,2007]" pageId="1" pageNumber="41" refString="Attems, C. (1929 a) Das Tierreich. 52 Myriapoda. I. Geophilomorpha. De Gruyter and Co., Berlin, Leipzig, 388 pp." type="book" year="1929" yearSuffix="a">Attems 1929a</bibRefCitation>
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: 182 (description).
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<bibRefCitation id="EFC04B4AFFAEFFF4C8BBB5ED31F2F851" author="Attems" box="[1270,1433,1982,2007]" pageId="1" pageNumber="41" refString="Attems, C. (1929 b) Die Myriopodenfauna von Albanien und Jugoslavien. Zoologische Jahrbucher, Abteilung fur Systematik, 56, 269 - 356." type="journal article" year="1929" yearSuffix="b">Attems 1929b</bibRefCitation>
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: 287, 296.
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<bibRefCitation id="EFC04B4AFFADFFF7CD44B2C535A1FF28" author="Brolemann" box="[265,458,151,175]" pageId="2" pageNumber="42" refString="Brolemann, H. W. (1932) Tableaux de determination des chilopodes signales en Afrique du Nord. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord, 23, 31 - 64." type="journal article" year="1932">Brolemann 1932</bibRefCitation>
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: 43 (in key).
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<bibRefCitation id="EFC04B4AFFADFFF7CE2FB2C53680FF28" author="Szabo" box="[610,747,151,175]" pageId="2" pageNumber="42" refString="Szabo, M. (1932) Die Myriopoden der Halbinsel Tihany. Arbeiten des Ungarischen Biologischen Forschungsinstitutes in Tihany, 5, 44 - 50." type="journal article" year="1932">Szabó 1932</bibRefCitation>
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: 48 (new record).
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<bibRefCitation id="EFC04B4AFFADFFF7CFF6B2C530E9FF28" author="Matic" box="[955,1154,151,176]" pageId="2" pageNumber="42" refString="Matic, Z., Negrea, S. & Prunescu, C. (1970) Considerations zoogeographiques sur la faune des Chilopodes de Roumanie. Bulletin du Museum d'Histoire Naturelle, Paris, 41 (suppl. 2), 96 - 101." type="journal article" year="1970">
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Matic
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<emphasis id="B925EAA9FFADFFF7C849B2CB3055FF37" box="[1028,1086,152,176]" italics="true" pageId="2" pageNumber="42">et al.</emphasis>
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1970
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</bibRefCitation>
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: 98.
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<bibRefCitation id="EFC04B4AFFADFFF7C8F4B2C53154FF28" author="Matic" box="[1209,1343,151,175]" pageId="2" pageNumber="42" refString="Matic, Z. (1972) Clasa Chilopoda, Subclasa Epimorpha. Fauna Republicii Socialiste Romania 6. Academiei Republicii Socialiste Romania, Bucuresti, 220 pp." type="book" year="1972">Matic 1972</bibRefCitation>
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||
: 107 (in key), 118 (description).
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<bibRefCitation id="EFC04B4AFFADFFF7CDE9B2EF367FFF53" author="Kos" box="[420,532,188,212]" pageId="2" pageNumber="42" refString="Kos, I. (1992) A review of the taxonomy, geographical distribution and ecology of the centipedes of Yugoslavia (Myriapoda Chilopoda). Bericht des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, suppl. 10, 353 - 360." type="journal article" year="1992">Kos 1992</bibRefCitation>
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: 354.
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<bibRefCitation id="EFC04B4AFFADFFF7CE1BB2EE36B2FF53" author="Stoev" box="[598,729,188,212]" pageId="2" pageNumber="42" refString="Stoev, P. (1997) A check - list of the centipedes of the Balkan peninsula with some taxonomic notes and a complete bibliography (Chilopoda). Entomologica Scandinavica, suppl. 51, 87 - 105." type="journal article" year="1997">Stoev 1997</bibRefCitation>
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: 102.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6530FFADFFF7CC88B2B23167FD20" pageId="2" pageNumber="42" type="materials_examined">
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<paragraph id="8BEE36BBFFADFFF7CC88B2B2304DFF70" blockId="2.[151,1436,151,679]" box="[197,1062,223,248]" pageId="2" pageNumber="42">
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<emphasis id="B925EAA9FFADFFF7CC88B2B23508FF7F" bold="true" box="[197,355,223,248]" pageId="2" pageNumber="42">
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<typeStatus id="54EA8819FFADFFF7CC88B2B2356BFF7F" box="[197,256,224,248]" pageId="2" pageNumber="42">Type</typeStatus>
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locality.
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</emphasis>
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“Østen for
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<collectingCountry id="F346762BFFADFFF7CDABB28D3648FF70" box="[486,547,223,247]" name="Algeria" pageId="2" pageNumber="42">Bona</collectingCountry>
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” [= east of Annaba (
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<collectingCountry id="F346762BFFADFFF7CF46B28D3709FF70" box="[779,866,223,247]" name="Algeria" pageId="2" pageNumber="42">Algeria</collectingCountry>
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)](
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<bibRefCitation id="EFC04B4AFFADFFF7CF35B28D3072FF70" author="Meinert" box="[888,1049,223,247]" pageId="2" pageNumber="42" refString="Meinert, F. (1870) Myriapoda Musaei Hauniensis. Bitrag til Myriapodernes morphologi og systematik. I. Geophile. Naturhistorisk Tidsskrift, 7, 1 - 128." type="journal article" year="1870">Meinert 1870</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BEE36BBFFADFFF7CC88B356311CFE00" blockId="2.[151,1436,151,679]" pageId="2" pageNumber="42">
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<emphasis id="B925EAA9FFADFFF7CC88B3563519FE9B" bold="true" box="[197,370,259,284]" pageId="2" pageNumber="42">
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<typeStatus id="54EA8819FFADFFF7CC88B356356BFE9B" box="[197,256,260,284]" pageId="2" pageNumber="42">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54EA8819FFADFFF7CD35B3563588FE9B" box="[376,483,260,284]" pageId="2" pageNumber="42" type="syntype">Syntypes</typeStatus>
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: a total of
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<specimenCount id="9D57FD32FFADFFF7CE1BB356369AFE9B" box="[598,753,260,284]" pageId="2" pageNumber="42" type="generic">36 specimens</specimenCount>
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, including
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<specimenCount id="9D57FD32FFADFFF7CF26B356378FFE9B" box="[875,996,260,284]" pageId="2" pageNumber="42" type="female">16 females</specimenCount>
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and
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<specimenCount id="9D57FD32FFADFFF7C857B35630EBFE9B" box="[1050,1152,260,284]" pageId="2" pageNumber="42" type="male">20 males</specimenCount>
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, according to the original description (
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<bibRefCitation id="EFC04B4AFFADFFF7CD6FB37535AAFEB8" author="Meinert" box="[290,449,295,319]" pageId="2" pageNumber="42" refString="Meinert, F. (1870) Myriapoda Musaei Hauniensis. Bitrag til Myriapodernes morphologi og systematik. I. Geophile. Naturhistorisk Tidsskrift, 7, 1 - 128." type="journal article" year="1870">Meinert 1870</bibRefCitation>
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); however, a total of
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<specimenCount id="9D57FD32FFADFFF7CEFCB3753726FEB8" box="[689,845,295,319]" pageId="2" pageNumber="42" type="generic">34 specimens</specimenCount>
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, including
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<specimenCount id="9D57FD32FFADFFF7CF87B3753023FEB8" box="[970,1096,295,319]" pageId="2" pageNumber="42" type="female">13 females</specimenCount>
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,
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<specimenCount id="9D57FD32FFADFFF7C819B37530D3FEB8" box="[1108,1208,295,319]" pageId="2" pageNumber="42" type="male">20 males</specimenCount>
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and another incomplete specimen of unknown sex, are currently preserved in the Zoological Museum, Natural History Museum of
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<collectingCountry id="F346762BFFADFFF7CCDAB33D3568FE00" box="[151,259,367,391]" name="Denmark" pageId="2" pageNumber="42">Denmark</collectingCountry>
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, University of Copenhagen, and confidently recognised as the original series examined by F. Meinert.
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</paragraph>
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<paragraph id="8BEE36BBFFADFFF7CC88B3C135F9FE2B" blockId="2.[151,1436,151,679]" box="[197,402,403,428]" pageId="2" pageNumber="42">
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<emphasis id="B925EAA9FFADFFF7CC88B3C1352CFE2B" bold="true" box="[197,327,403,428]" pageId="2" pageNumber="42">Synonyms.</emphasis>
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None.
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</paragraph>
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<paragraph id="8BEE36BBFFADFFF7CC88B3E53167FD20" blockId="2.[151,1436,151,679]" pageId="2" pageNumber="42">
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<emphasis id="B925EAA9FFADFFF7CC88B3E535A0FE57" bold="true" box="[197,459,439,464]" pageId="2" pageNumber="42">Differential diagnosis.</emphasis>
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With respect to the most similar species (
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<tableCitation id="C6D30300FFADFFF7CFD0B3E5379AFE48" box="[925,1009,439,463]" captionStart="TABLE 1" captionStartId="6.[151,235,544,565]" captionTargetBox="[151,1437,697,1408]" captionTargetPageId="6" captionText="TABLE 1. Taxonomic data and differences between G. carpophagus, G. e a s o n i, and G. arenarius. Data are exclusively from type material and specimens that have been identified confidently (see Material and methods). * Published counts of leg pairs from populations that have been assigned to G. carpophagus, but are actually of uncertain identity, have been ignored. See also Figs 10 – 11." httpUri="http://table.plazi.org/id/DF2E6633FFA9FFF3CCDAB0723565FD1B" pageId="2" pageNumber="42" tableUuid="DF2E6633FFA9FFF3CCDAB0723565FD1B">Table 1</tableCitation>
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,
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<figureCitation id="136A2A3EFFADFFF7CFB2B3E53004FE48" box="[1023,1135,439,463]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 9" captionStart-2="FIGURES 10 – 11" captionStartId-0="2.[151,264,1585,1608]" captionStartId-1="3.[151,264,1697,1720]" captionStartId-2="5.[151,264,1524,1547]" captionTargetBox-0="[167,1431,734,1563]" captionTargetBox-1="[151,1436,337,1675]" captionTargetBox-2="[158,1429,455,1503]" captionTargetId-0="figure@2.[151,1436,733,1564]" captionTargetId-1="figure@3.[151,1436,337,1676]" captionTargetId-2="figure@5.[151,1436,447,1503]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." captionText-1="FIGURES 7 – 9. Geophilus arenarius; photos taken from a syntype 3, 27 mm long, with 55 pairs of legs. 7 — anterior part of body, dorsal view; 8 — head and forcipular segment, ventral view; 9 — part of trunk at ca. 20 % of the series of leg-bearing segments (segments 9 – 12 out of a total of 55), ventral view." captionText-2="FIGURES 10 – 11. Comparison between G. carpophagus, G. e a s o n i, and G. arenarius; line-drawings from adult specimens, setae omitted. 10 — forcipular segment, ventral view; 11 — a trunk segment at about 20 % of the sequence of leg-bearing segments, ventral view." httpUri-0="https://zenodo.org/record/207322/files/figure.png" httpUri-1="https://zenodo.org/record/207323/files/figure.png" httpUri-2="https://zenodo.org/record/207324/files/figure.png" pageId="2" pageNumber="42">Figs 1–11</figureCitation>
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),
|
||
<taxonomicName id="4C514D38FFADFFF7C8C8B3EA3164FE57" box="[1157,1295,440,464]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFADFFF7C8C8B3EA3164FE57" box="[1157,1295,440,464]" italics="true" pageId="2" pageNumber="42">G. arenarius</emphasis>
|
||
</taxonomicName>
|
||
differs from both
|
||
<taxonomicName id="4C514D38FFADFFF7CC9FB38F35EDFE73" box="[210,390,476,500]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFADFFF7CC9FB38F35EDFE73" box="[210,390,476,500]" italics="true" pageId="2" pageNumber="42">G. carpophagus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C514D38FFADFFF7CD8FB38F3646FE73" box="[450,557,477,500]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="easoni">
|
||
<emphasis id="B925EAA9FFADFFF7CD8FB38F3646FE73" box="[450,557,477,500]" italics="true" pageId="2" pageNumber="42">G. easoni</emphasis>
|
||
</taxonomicName>
|
||
in lacking the transverse suture on the head and the so-called carpophagusstructures on the trunk, and having relatively stouter antennae and forcipular coxosternite (see also below, under Discussion). Additionally,
|
||
<taxonomicName id="4C514D38FFADFFF7CD8FB0773627FDBB" box="[450,588,549,572]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFADFFF7CD8FB0773627FDBB" box="[450,588,549,572]" italics="true" pageId="2" pageNumber="42">G. arenarius</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C514D38FFADFFF7CE93B07737E5FDBB" box="[734,910,548,572]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFADFFF7CE93B07737E5FDBB" box="[734,910,548,572]" italics="true" pageId="2" pageNumber="42">G. carpophagus</emphasis>
|
||
</taxonomicName>
|
||
in the distinctly smaller size at full growth, the usually blunt and more sclerotised tubercles lining the intermediate part of the labrum, and the presence of a minute denticle at the basis of the forcipular tarsungula.
|
||
<taxonomicName id="4C514D38FFADFFF7CE8AB03E37C4FD03" box="[711,943,620,644]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFADFFF7CE8AB03E37C4FD03" box="[711,943,620,644]" italics="true" pageId="2" pageNumber="42">Geophilus arenarius</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C514D38FFADFFF7C80AB03F30DBFD03" box="[1095,1200,621,644]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="easoni">
|
||
<emphasis id="B925EAA9FFADFFF7C80AB03F30DBFD03" box="[1095,1200,621,644]" italics="true" pageId="2" pageNumber="42">G. easoni</emphasis>
|
||
</taxonomicName>
|
||
in the fewer bristles lining the lateral parts of the labrum, in the greyish colour of the trunk, and the higher number of legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2E6633FFADFFF7CCDAB463316BF945" httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="2" pageNumber="42" targetBox="[167,1431,734,1563]" targetPageId="2">
|
||
<paragraph id="8BEE36BBFFADFFF7CCDAB463316BF945" blockId="2.[151,1437,1585,1730]" pageId="2" pageNumber="42">
|
||
<emphasis id="B925EAA9FFADFFF7CCDAB4633551F9C1" bold="true" box="[151,314,1585,1608]" pageId="2" pageNumber="42">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C514D38FFADFFF7CD72B4633664F9C0" box="[319,527,1585,1607]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFADFFF7CD72B4633664F9C0" box="[319,527,1585,1607]" italics="true" pageId="2" pageNumber="42">Geophilus arenarius</emphasis>
|
||
</taxonomicName>
|
||
; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1–5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1—head, without right antenna, setae omitted, dorsal view; 2—anterior part of head, without antennae, mandibles and maxillae, ventral view; 3—maxillary complex, left half, ventral view; 4—forcipular segment, setae omitted, ventral view; 5—ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6—ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34B6530FFADFFF0CC88B4BC35C7FE00" lastPageId="5" lastPageNumber="45" pageId="2" pageNumber="42" type="description">
|
||
<paragraph id="8BEE36BBFFADFFF7CC88B4BC3174F881" blockId="2.[151,1436,1774,2015]" box="[197,1311,1774,1799]" pageId="2" pageNumber="42">
|
||
<emphasis id="B925EAA9FFADFFF7CC88B4BC3645F880" bold="true" box="[197,558,1774,1799]" pageId="2" pageNumber="42">Description of an adult female</emphasis>
|
||
(one of the
|
||
<typeStatus id="54EA8819FFADFFF7CEF9B4BD3772F880" box="[692,793,1775,1799]" pageId="2" pageNumber="42" type="syntype">syntypes</typeStatus>
|
||
,
|
||
<quantity id="4CA99B5EFFADFFF7CF68B4BC371FF881" box="[805,884,1774,1798]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="42" unit="mm" value="34.0">34 mm</quantity>
|
||
long, with 57 pairs of legs;
|
||
<figureCitation id="136A2A3EFFADFFF7C8E0B4BC317AF881" box="[1197,1297,1774,1798]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="2" pageNumber="42">Figs 1–5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFADFFF7CC88B541370FF8C8" blockId="2.[151,1436,1774,2015]" pageId="2" pageNumber="42">
|
||
<emphasis id="B925EAA9FFADFFF7CC88B54135EFF8AC" box="[197,388,1811,1835]" italics="true" pageId="2" pageNumber="42">General features</emphasis>
|
||
. Body only slightly narrowing forward, more attenuated towards the posterior tip. Colour (preserved in 70% ethanol) almost uniform, pale brownish grey.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFADFFF6CC88B50936FCFE9B" blockId="2.[151,1436,1774,2015]" lastBlockId="3.[151,1437,151,284]" lastPageId="3" lastPageNumber="43" pageId="2" pageNumber="42">
|
||
<emphasis id="B925EAA9FFADFFF7CC88B50935FBF8F4" box="[197,400,1883,1907]" italics="true" pageId="2" pageNumber="42">Cephalic capsule.</emphasis>
|
||
Cephalic plate (
|
||
<figureCitation id="136A2A3EFFADFFF7CE03B50836FAF8F5" box="[590,657,1882,1906]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="2" pageNumber="42">Fig. 1</figureCitation>
|
||
) sub-trapezoid, as long as wide; anterior margin slightly angulated, lateral margins evidently convex and convergent forward, posterior margin slightly concave; no transverse suture. Clypeus (
|
||
<figureCitation id="136A2A3EFFADFFF7CD48B5F03527F83D" box="[261,332,1954,1978]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="2" pageNumber="42">Fig. 2</figureCitation>
|
||
) uniformly areolate, without finely areolate clypeal areas and without plagulae; lateral margins complete; 10 pairs of setae, including a paramedian pair close to the anterior margin, most setae arranged in a transverse row in the anterior part of the clypeus, and two paramedian pairs posterior to the latter. Labrum (
|
||
<figureCitation id="136A2A3EFFACFFF6C90BB2C531E5FF28" box="[1350,1422,151,175]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="3" pageNumber="43">Fig. 2</figureCitation>
|
||
): intermediate part continuous with the clypeus, bearing 4 sclerotised stout tubercles; each lateral part separated from the clypeus by a complete margin, bearing ca. 8 bristles branching into fine short filaments close to the tip. Pleurites uniformly areolate, with 3 anterior setae.
|
||
</paragraph>
|
||
<caption id="DF2E6633FFACFFF6CCDAB4F336BFF973" httpUri="https://zenodo.org/record/207323/files/figure.png" pageId="3" pageNumber="43" targetBox="[151,1436,337,1675]" targetPageId="3">
|
||
<paragraph id="8BEE36BBFFACFFF6CCDAB4F336BFF973" blockId="3.[151,1436,1697,1780]" pageId="3" pageNumber="43">
|
||
<emphasis id="B925EAA9FFACFFF6CCDAB4F33550F930" bold="true" box="[151,315,1697,1720]" pageId="3" pageNumber="43">FIGURES 7–9.</emphasis>
|
||
<taxonomicName id="4C514D38FFACFFF6CD0EB4F3367FF930" box="[323,532,1697,1719]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="3" pageNumber="43" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFACFFF6CD0EB4F3367FF930" box="[323,532,1697,1719]" italics="true" pageId="3" pageNumber="43">Geophilus arenarius</emphasis>
|
||
</taxonomicName>
|
||
; photos taken from a syntype 3, 27 mm long, with 55 pairs of legs. 7—anterior part of body, dorsal view; 8—head and forcipular segment, ventral view; 9—part of trunk at ca. 20% of the series of leg-bearing segments (segments 9–12 out of a total of 55), ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEE36BBFFACFFF1CC88B57334A4FF53" blockId="3.[151,1436,1825,2029]" lastBlockId="4.[151,1437,151,2012]" lastPageId="4" lastPageNumber="44" pageId="3" pageNumber="43">
|
||
<emphasis id="B925EAA9FFACFFF6CC88B5733544F8BE" box="[197,303,1825,1849]" italics="true" pageId="3" pageNumber="43">Antennae</emphasis>
|
||
(
|
||
<figureCitation id="136A2A3EFFACFFF6CD73B57335EAF8BE" box="[318,385,1825,1849]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="3" pageNumber="43">Fig. 1</figureCitation>
|
||
). Each antenna ca. 2.2 times as long as the head; intermediate articles about as long as wide; article XIV ca. 1.6 times as long as wide. Setae of various size on the most basal articles, gradually denser and shorter from the basal articles to the distal ones. Apical sensilla ca. 14 μm long, spear-like, slender, slightly narrowing towards the tip. Club-like sensilla only on article XIV, grouped on the distal parts of both the internal and external sides. Three longitudinal rows of propioceptive spine-like sensilla at the bases of the articles; rows lacking on articles VI, X and XIV, where only a single, dorsal sensillum is present. Groups of 1–6 sensilla, similar to the apical ones but shorter, on both the ventro-internal and the dorso-external sides of the distal part of articles II, V, IX and XIII.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B2B23602FF70" blockId="4.[151,1437,151,2012]" box="[197,617,223,248]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B2B2355BFF7F" box="[197,304,224,248]" italics="true" pageId="4" pageNumber="44">Mandible</emphasis>
|
||
. A single pectinate lamella.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B3563718FE73" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B35635F3FE9B" box="[197,408,260,284]" italics="true" pageId="4" pageNumber="44">Maxillary complex</emphasis>
|
||
(
|
||
<figureCitation id="136A2A3EFFABFFF1CDE5B3563585FE9B" box="[424,494,260,284]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="4" pageNumber="44">Fig. 3</figureCitation>
|
||
). First maxillae: coxosternite entire, with short lappets; coxal projections sub-triangular, longer than wide; telopodites longer than the coxal projections, composed of two articles, with elongate lappets; coxal projections and the distal article of the telopodites bearing setae on the basal part, spine-like sensilla close to the tip, and minute scales on the tip. Second maxillae: coxosternite entire, uniformly areolate, the anterior margin widely concave, with setae close to the anterior margin; metameric pores featuring as transverse slits; telopodites composed of three articles, gradually narrowing towards the tip; claw simple, sub-conic, gradually tapering, slightly bent, with 1–2 short sensilla on the ventral side.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B0523700FCB0" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B05235CFFD9F" box="[197,420,512,536]" italics="true" pageId="4" pageNumber="44">Forcipular segment</emphasis>
|
||
(
|
||
<figureCitation id="136A2A3EFFABFFF1CDFBB3AD3595FD90" box="[438,510,511,535]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="4" pageNumber="44">Fig. 4</figureCitation>
|
||
). Forcipules, when closed, just reaching the anterior margin of the head. Tergite subtrapezoid, ca. 4.4 times as wide as long, lateral margins evidently converging forward; no pretergite exposed. Pleurites without a distinct dorsal ridge. Coxosternite: two paramedian groups of 4–5 small setae on the dorsal side, close to the anterior margin; no denticles; ventrally exposed part ca. 1.76 times as wide as long; coxopleural sutures entirely ventral, strongly converging backward; chitin-lines incomplete, diverging lateral from the condyles. Trochanteroprefemur ca. 1.1 times as wide as long, the external side ca. 2.0 times as long as the internal side. Tarsungulum abruptly narrowing near the base, the distal part curved and tapering uniformly, the internal margin entire. A tiny denticle at the basis of tarsungulum; no denticles on the other forcipular articles. Poison calyx relatively long, asymmetric, lodged in the distal part of the trochanteroprefemur.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B11634ACFBDF" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B11635D4FCDB" box="[197,447,836,860]" italics="true" pageId="4" pageNumber="44">Leg-bearing segments</emphasis>
|
||
. Tergite 1 wider than metatergite 2, lateral margins slightly converging backward; no distinct pretergite 1. Metasternites without
|
||
<taxonomicName id="4C514D38FFABFFF1CE31B1353761FCF8" box="[636,778,871,895]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="carpophagus">carpophagus</taxonomicName>
|
||
pits on the anterior margin, only a small projection on the posterior margin of those of the anterior segments (resembling the peg usually facing the pit in a typical carpophagus-structure;
|
||
<figureCitation id="136A2A3EFFABFFF1CD75B1FD35E3FC40" box="[312,392,943,967]" captionStart="FIGURES 10 – 11" captionStartId="5.[151,264,1524,1547]" captionTargetBox="[158,1429,455,1503]" captionTargetId="figure@5.[151,1436,447,1503]" captionTargetPageId="5" captionText="FIGURES 10 – 11. Comparison between G. carpophagus, G. e a s o n i, and G. arenarius; line-drawings from adult specimens, setae omitted. 10 — forcipular segment, ventral view; 11 — a trunk segment at about 20 % of the sequence of leg-bearing segments, ventral view." httpUri="https://zenodo.org/record/207324/files/figure.png" pageId="4" pageNumber="44">Fig. 11</figureCitation>
|
||
). Ventral pore-fields from segment 1 to the penultimate: a transverse posterior band, which is entire on the most anterior segments, medially interrupted on the remaining segments; anterior segments with additional small pores-fields close to the anterior corners of the metasternite and on procoxae and metacoxae. Legs 1 smaller than the other legs. Claws with two accessory spines, the anterior one distinctly longer than the posterior one.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B63637DAFAFF" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B6363671FBFB" box="[197,538,1124,1148]" italics="true" pageId="4" pageNumber="44">Ultimate leg-bearing segment</emphasis>
|
||
(
|
||
<figureCitation id="136A2A3EFFABFFF1CE61B636361FFBFB" box="[556,628,1124,1148]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="4" pageNumber="44">Fig. 5</figureCitation>
|
||
). Pleuropretergite entire, without sutures. Metatergite ca. 1.8 times as wide as long, lateral margins convex and converging backward, posterior margin truncate. Metasternite sub-trapezoid, ca. 2.0 times as wide as long, lateral margins convex, posterior margin almost straight; setae uniformly scattered. Coxopleura moderately swollen, reaching forward the posterior margin of the presternite; setae uniformly scattered. Coxal organs opening through distinct pores, all on the ventral side along the lateral margins of the metasternite, 10 on one side and 9 on the other side. Ultimate telopodites ca. 1.1 times as long as, and as wide as, the penultimate; setae uniformly scattered on both dorsal and ventral sides; claws similar to those of precedent legs, only slightly smaller than those, the accessory spines distinctly shorter.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B7D63743FA47" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B7D635CEFA1B" box="[197,421,1412,1436]" italics="true" pageId="4" pageNumber="44">Postpedal segments</emphasis>
|
||
(
|
||
<figureCitation id="136A2A3EFFABFFF1CDFAB7D63595FA1B" box="[439,510,1412,1436]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="4" pageNumber="44">Fig. 5</figureCitation>
|
||
). First genital pleurosternite entire, without distinct pleurites. Gonopodal lamina distinctly bilobed. A pair of distinct anal organs and pores.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B799308DFA63" blockId="4.[151,1437,151,2012]" box="[197,1254,1483,1509]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B799367FFA63" bold="true" box="[197,532,1483,1508]" pageId="4" pageNumber="44">Differences in an adult male</emphasis>
|
||
(one of the
|
||
<typeStatus id="54EA8819FFABFFF1CED6B79F376BFA62" box="[667,768,1485,1509]" pageId="4" pageNumber="44" type="syntype">syntypes</typeStatus>
|
||
,
|
||
<quantity id="4CA99B5EFFABFFF1CF41B79E3731FA63" box="[780,858,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="44" unit="mm" value="27.0">27 mm</quantity>
|
||
long, with 55 pairs of legs;
|
||
<figureCitation id="136A2A3EFFABFFF1C8D9B79E30B3FA63" box="[1172,1240,1484,1508]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="4" pageNumber="44">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B7A237FAF9F3" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B7A23678F98F" box="[197,531,1520,1544]" italics="true" pageId="4" pageNumber="44">Ultimate leg-bearing segment</emphasis>
|
||
(
|
||
<figureCitation id="136A2A3EFFABFFF1CE6EB7A2360CF98F" box="[547,615,1520,1544]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1585,1608]" captionTargetBox="[167,1431,734,1563]" captionTargetId="figure@2.[151,1436,733,1564]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Geophilus arenarius; line-drawings from a syntype Ƥ, 34 mm long, with 57 pairs of legs (Figs 1 – 5), and a syntype 3, 27 mm long, with 55 pairs of legs (Fig. 6). 1 — head, without right antenna, setae omitted, dorsal view; 2 — anterior part of head, without antennae, mandibles and maxillae, ventral view; 3 — maxillary complex, left half, ventral view; 4 — forcipular segment, setae omitted, ventral view; 5 — ultimate leg-bearing segment and postpedal segments in the female, setae omitted, ventral view; 6 — ultimate leg-bearing segment and postpedal segments in the male, setae omitted, ventral view." httpUri="https://zenodo.org/record/207322/files/figure.png" pageId="4" pageNumber="44">Fig. 6</figureCitation>
|
||
). Distinctly denser, shorter setae on the posterior part of the metasternite, the postero-mesal parts of coxopleura, and the ventral side of all articles of the telopodites. Ultimate telopodites moderately swollen, ca. 1.3 times as wide as the penultimate. First genital sternite separated from pleurites by distinct sutures. Gonopods bi-articulate, with conical penis in between.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF1CC88B42D30F2F8AF" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="44">
|
||
<emphasis id="B925EAA9FFABFFF1CC88B42D366DF91F" bold="true" box="[197,518,1663,1688]" pageId="4" pageNumber="44">Geographical distribution.</emphasis>
|
||
<taxonomicName id="4C514D38FFABFFF1CE5CB4D336F4F91F" box="[529,671,1665,1688]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFABFFF1CE5CB4D336F4F91F" box="[529,671,1665,1688]" italics="true" pageId="4" pageNumber="44">G. arenarius</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54EA8819FFABFFF1CF85B4D33793F91E" box="[968,1016,1665,1689]" pageId="4" pageNumber="44">type</typeStatus>
|
||
locality, near Annaba,
|
||
<collectingCountry id="F346762BFFABFFF1C946B4D23108F91F" box="[1291,1379,1664,1688]" name="Algeria" pageId="4" pageNumber="44">Algeria</collectingCountry>
|
||
. Our direct examination of other specimens of the
|
||
<taxonomicName id="4C514D38FFABFFF1CEE7B4F63757F93B" box="[682,828,1700,1724]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFABFFF1CEE7B4F63757F93B" box="[682,828,1700,1724]" italics="true" pageId="4" pageNumber="44">carpophagus</emphasis>
|
||
</taxonomicName>
|
||
species-complex collected throughout north-western Africa (from
|
||
<collectingCountry id="F346762BFFABFFF1CD7EB49B35F3F967" box="[307,408,1737,1760]" name="Morocco" pageId="4" pageNumber="44">Morocco</collectingCountry>
|
||
to
|
||
<collectingCountry id="F346762BFFABFFF1CD8EB49A3672F967" box="[451,537,1736,1760]" name="Tunisia" pageId="4" pageNumber="44">Tunisia</collectingCountry>
|
||
, including coastal and mountainous sites; listed under Material and methods) failed to detect other specimens of
|
||
<emphasis id="B925EAA9FFABFFF1CE79B4BF36D5F883" box="[564,702,1773,1796]" italics="true" pageId="4" pageNumber="44">G. a re n a r i u s</emphasis>
|
||
. Instead, these specimens confirm that north-western Africa is inhabited by at least two species in the
|
||
<taxonomicName id="4C514D38FFABFFF1CE00B54236B4F8AF" box="[589,735,1808,1832]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFABFFF1CE00B54236B4F8AF" box="[589,735,1808,1832]" italics="true" pageId="4" pageNumber="44">carpophagus</emphasis>
|
||
</taxonomicName>
|
||
complex, one of which is
|
||
<taxonomicName id="4C514D38FFABFFF1C845B54330F9F8AF" box="[1032,1170,1809,1832]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFABFFF1C845B54330F9F8AF" box="[1032,1170,1809,1832]" italics="true" pageId="4" pageNumber="44">G. arenarius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEE36BBFFABFFF0CC88B56635C7FE00" blockId="4.[151,1437,151,2012]" lastBlockId="5.[151,1436,151,391]" lastPageId="5" lastPageNumber="45" pageId="4" pageNumber="44">
|
||
Records of
|
||
<taxonomicName id="4C514D38FFABFFF1CD04B56735BEF8CB" box="[329,469,1845,1868]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFABFFF1CD04B56735BEF8CB" box="[329,469,1845,1868]" italics="true" pageId="4" pageNumber="44">G. arenarius</emphasis>
|
||
</taxonomicName>
|
||
from different localities in
|
||
<collectingCountry id="F346762BFFABFFF1CF5DB566370DF8CB" box="[784,870,1844,1868]" name="Croatia" pageId="4" pageNumber="44">Croatia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F346762BFFABFFF1CF3FB56737BFF8CA" box="[882,980,1845,1869]" name="Hungary" pageId="4" pageNumber="44">Hungary</collectingCountry>
|
||
and
|
||
<collectingCountry id="F346762BFFABFFF1C840B5663019F8CB" box="[1037,1138,1844,1868]" name="Romania" pageId="4" pageNumber="44">Romania</collectingCountry>
|
||
have been listed synthetically by
|
||
<bibRefCitation id="EFC04B4AFFABFFF1CCB5B50A35A2F8F7" author="Tomosvary" box="[248,457,1880,1904]" pageId="4" pageNumber="44" refString="Tomosvary, E. (1880) Beitrag zur Kenntnis der Myriopoden Ungarns. I. Die Chilopoden. Zoologischer Anzeiger, 3, 617 - 619." type="journal article" year="1880">Tömösváry (1880)</bibRefCitation>
|
||
and in more detail by
|
||
<bibRefCitation id="EFC04B4AFFABFFF1CE8AB50A370EF8F7" author="Daday" box="[711,869,1880,1904]" pageId="4" pageNumber="44" refString="Daday, E. (1889) Myriopoda regni Hungariae. Kiralyi Magyar Termeszettudomanyi Tarsulat, Budapest, 104 pp." type="book" year="1889">Daday (1889)</bibRefCitation>
|
||
. These have been repeatedly mentioned acritically by subsequent authors, and an additional record from
|
||
<collectingCountry id="F346762BFFABFFF1CF44B52F3700F812" box="[777,875,1917,1941]" name="Hungary" pageId="4" pageNumber="44">Hungary</collectingCountry>
|
||
has been given by
|
||
<bibRefCitation id="EFC04B4AFFABFFF1C81CB52E3081F813" author="Szabo" box="[1105,1258,1916,1940]" pageId="4" pageNumber="44" refString="Szabo, M. (1932) Die Myriopoden der Halbinsel Tihany. Arbeiten des Ungarischen Biologischen Forschungsinstitutes in Tihany, 5, 44 - 50." type="journal article" year="1932">Szabó (1932)</bibRefCitation>
|
||
under the misspelled name
|
||
<emphasis id="B925EAA9FFABFFF1CD7BB5F335E2F83F" box="[310,393,1953,1976]" italics="true" pageId="4" pageNumber="44">arenius</emphasis>
|
||
. However, all these old records have not been confirmed by modern authors (
|
||
<bibRefCitation id="EFC04B4AFFABFFF1C95DB5F231FCF83F" author="Matic" box="[1296,1431,1952,1976]" pageId="4" pageNumber="44" refString="Matic, Z. (1972) Clasa Chilopoda, Subclasa Epimorpha. Fauna Republicii Socialiste Romania 6. Academiei Republicii Socialiste Romania, Bucuresti, 220 pp." type="book" year="1972">Matic 1972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC04B4AFFABFFF1CCDAB5973563F85B" author="Kos" box="[151,264,1988,2012]" pageId="4" pageNumber="44" refString="Kos, I. (1992) A review of the taxonomy, geographical distribution and ecology of the centipedes of Yugoslavia (Myriapoda Chilopoda). Bericht des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, suppl. 10, 353 - 360." type="journal article" year="1992">Kos 1992</bibRefCitation>
|
||
; L. Danyi, pers. comm.), or by our preliminary examination of
|
||
<taxonomicName id="4C514D38FFABFFF1CFAAB5963031F85B" box="[999,1114,1988,2012]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="4" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925EAA9FFABFFF1CFAAB5963031F85B" box="[999,1114,1988,2012]" italics="true" pageId="4" pageNumber="44">Geophilus</emphasis>
|
||
</taxonomicName>
|
||
specimens from throughout Europe.
|
||
<bibRefCitation id="EFC04B4AFFAAFFF0CCB8B2C535ECFF28" author="Matic" box="[245,391,151,175]" pageId="5" pageNumber="45" refString="Matic, Z. (1972) Clasa Chilopoda, Subclasa Epimorpha. Fauna Republicii Socialiste Romania 6. Academiei Republicii Socialiste Romania, Bucuresti, 220 pp." type="book" year="1972">Matic (1972)</bibRefCitation>
|
||
suspected explicitly that the records from
|
||
<collectingCountry id="F346762BFFAAFFF0CF2DB2C537AEFF28" box="[864,965,151,175]" name="Romania" pageId="5" pageNumber="45">Romania</collectingCountry>
|
||
should actually be referred to
|
||
<taxonomicName id="4C514D38FFAAFFF0C95AB2CA34A7FF53" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFAAFFF0C95AB2CA34A7FF53" italics="true" pageId="5" pageNumber="45">G. carpophagus</emphasis>
|
||
</taxonomicName>
|
||
. Taking into account the information given by
|
||
<bibRefCitation id="EFC04B4AFFAAFFF0CEA1B2EE37E0FF53" author="Daday" box="[748,907,188,212]" pageId="5" pageNumber="45" refString="Daday, E. (1889) Myriopoda regni Hungariae. Kiralyi Magyar Termeszettudomanyi Tarsulat, Budapest, 104 pp." type="book" year="1889">Daday (1889)</bibRefCitation>
|
||
, the specimens examined by him depart obviously from the
|
||
<typeStatus id="54EA8819FFAAFFF0CD09B2B235CDFF7F" box="[324,422,224,248]" pageId="5" pageNumber="45" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C514D38FFAAFFF0CD80B2B23633FF7F" box="[461,600,224,248]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="arenarius">
|
||
<emphasis id="B925EAA9FFAAFFF0CD80B2B23633FF7F" box="[461,600,224,248]" italics="true" pageId="5" pageNumber="45">G. arenarius</emphasis>
|
||
</taxonomicName>
|
||
in major morphological features (complete chitin-lines, higher number of trunk segments, distinct
|
||
<taxonomicName id="4C514D38FFAAFFF0CDE4B356365CFE9B" box="[425,567,260,284]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="carpophagus">carpophagus</taxonomicName>
|
||
structure), so that they are obviously misidentified specimens of another species, possibly not in the
|
||
<taxonomicName id="4C514D38FFAAFFF0CDE7B37A3657FEC7" box="[426,572,296,320]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFAAFFF0CDE7B37A3657FEC7" box="[426,572,296,320]" italics="true" pageId="5" pageNumber="45">carpophagus</emphasis>
|
||
</taxonomicName>
|
||
species-complex, more probably
|
||
<emphasis id="B925EAA9FFAAFFF0CFF6B37A3028FEC7" box="[955,1091,296,320]" italics="true" pageId="5" pageNumber="45">G. e l e c t r i c u s</emphasis>
|
||
Linnaeus. Even the record by
|
||
<bibRefCitation id="EFC04B4AFFAAFFF0CCDAB31E3546FEE3" author="Szabo" box="[151,301,332,356]" pageId="5" pageNumber="45" refString="Szabo, M. (1932) Die Myriopoden der Halbinsel Tihany. Arbeiten des Ungarischen Biologischen Forschungsinstitutes in Tihany, 5, 44 - 50." type="journal article" year="1932">Szabó (1932)</bibRefCitation>
|
||
is most probably to be referred to
|
||
<taxonomicName id="4C514D38FFAAFFF0CEFBB31F370CFEE3" box="[694,871,332,356]" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="carpophagus">
|
||
<emphasis id="B925EAA9FFAAFFF0CEFBB31F370CFEE3" box="[694,871,332,356]" italics="true" pageId="5" pageNumber="45">G. carpophagus</emphasis>
|
||
</taxonomicName>
|
||
, based on investigations on surrounding localities (L. Danyi, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |