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<document id="E77089C5E12A33E52459D39B3164FF40" ID-DOI="10.11646/zootaxa.3683.4.5" ID-GBIF-Dataset="688f6814-7236-483e-9f65-45eefeba20f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="222164" ID-ZooBank="30BD249D-0F84-4FD5-8091-A0AC80262526" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460645156311" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="03F887A9FFB1FFA5FF7A66B9FB70FD96" docLanguage="en" docName="zt03683p426.pdf" docOrigin="Zootaxa 3683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cephennodes (Cephennodes) mtubatubanus Jałoszyński, 2013, sp. n." docType="treatment" docVersion="7" lastPageNumber="418" masterDocId="FFC1FFD1FFB7FFA2FFED662EFFA0FF87" masterDocTitle="The first record of the tribe Cephenniini in South Africa, with new species of Cephennomicrus Reitter and Cephennodes Reitter (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="426" masterPageNumber="411" pageNumber="417" updateTime="1698336797779" updateUser="plazi">
<mods:mods id="AB6A1DAA1DB9EB422DC04040ECE8423F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="934F5BA41DA1BD4F3BDE90083A0D9946">
<mods:title id="4D759FE4E2D08BE7A6A91A508560980A">The first record of the tribe Cephenniini in South Africa, with new species of Cephennomicrus Reitter and Cephennodes Reitter (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="25277FA601AD9C28D9744E081A817534" type="personal">
<mods:role id="429EC17EB05905D62668249B0D25CBF3">
<mods:roleTerm id="BE05CC49241C3DC941DE7F1EC2042325">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F925D6EB9C36E0BBC1CEA646C598F14B">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="0BDC1BF4D4BBF4E2D4FB0F6A6BF617BE">text</mods:typeOfResource>
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<mods:title id="62680D38A1F9859145A62D56EAEBE0E6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="01457FDDFAC1D920A12516BC77F64959">
<mods:date id="9B54BF9BD31E875CAF6AA9806709E7B8">2013</mods:date>
<mods:detail id="ED6344FC460555C4802F7C4115DE35A9" type="volume">
<mods:number id="9E1C98EA0B1E9C1AF0157698FE6A459D">3683</mods:number>
</mods:detail>
<mods:detail id="9A4CC3F97FF43FCB6A541B416BF670A6" type="issue">
<mods:number id="3118386AC375BCCAF7FADA48CD288B6B">4</mods:number>
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<mods:start id="7DAC7B52327AD7AD10A7F94BDA7D2284">411</mods:start>
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<mods:classification id="C606DA83F78061C18539900169419B56">journal article</mods:classification>
<mods:identifier id="21E8B5FD4E3036835959E16947D4847F" type="DOI">10.11646/zootaxa.3683.4.5</mods:identifier>
<mods:identifier id="3625E30044618AA847482E00A924E084" type="GBIF-Dataset">688f6814-7236-483e-9f65-45eefeba20f5</mods:identifier>
<mods:identifier id="9C088A08DF40C1E6139ABB1ECBBF0FDE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4D39022FC98373F5DD5DBDAB5D8FBFAE" type="Zenodo-Dep">222164</mods:identifier>
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<treatment id="03F887A9FFB1FFA5FF7A66B9FB70FD96" ID-DOI="http://doi.org/10.5281/zenodo.6155812" ID-GBIF-Taxon="119572019" ID-Zenodo-Dep="6155812" LSID="urn:lsid:plazi:treatment:03F887A9FFB1FFA5FF7A66B9FB70FD96" httpUri="http://treatment.plazi.org/id/03F887A9FFB1FFA5FF7A66B9FB70FD96" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417">
<subSubSection id="C34B6534FFB1FFA4FF7A66B9FEC6FF55" pageId="6" pageNumber="417" type="nomenclature">
<paragraph id="8BEE36BFFFB1FFA4FF7A66B9FD56FF36" blockId="6.[151,758,151,210]" box="[151,758,151,177]" pageId="6" pageNumber="417">
<heading id="D0A681D3FFB1FFA4FF7A66B9FD56FF36" bold="true" box="[151,758,151,177]" fontSize="11" level="1" pageId="6" pageNumber="417" reason="1">
<emphasis id="B925EAADFFB1FFA4FF7A66B9FD56FF36" bold="true" box="[151,758,151,177]" pageId="6" pageNumber="417">
<taxonomicName id="4C514D3CFFB1FFA4FF7A66B9FD0CFF36" box="[151,684,151,177]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="mtubatubanus" status="sp. nov." subGenus="Cephennodes">
<emphasis id="B925EAADFFB1FFA4FF7A66B9FE9AFF36" bold="true" box="[151,314,151,177]" italics="true" pageId="6" pageNumber="417">Cephennodes</emphasis>
(
<emphasis id="B925EAADFFB1FFA4FEA666B9FE4EFF36" bold="true" box="[331,494,151,177]" italics="true" pageId="6" pageNumber="417">Cephennodes</emphasis>
)
<emphasis id="B925EAADFFB1FFA4FE1366B9FD0CFF36" bold="true" box="[510,684,151,177]" italics="true" pageId="6" pageNumber="417">mtubatubanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21657D6FFB1FFA4FD5966B9FD56FF36" box="[692,758,151,177]" pageId="6" pageNumber="417" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF7A6694FEC6FF55" blockId="6.[151,758,151,210]" box="[151,358,186,210]" pageId="6" pageNumber="417">
(
<figureCitation id="136A2A3AFFB1FFA4FF726694FF4EFF55" box="[159,238,186,210]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1580,1603]" captionTargetBox="[158,1430,879,1549]" captionTargetId="figure@2.[151,1436,873,1559]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Dorsal habitus of holotype males. Cephennodes lucipetus sp. n. (1); Cephennodes zulunatalensis sp. n. (2); Cephennodes mtubatubanus sp. n. (3); Cephennodes sodwanaensis sp. n. (4)." httpUri="https://zenodo.org/record/222165/files/figure.png" pageId="6" pageNumber="417">Figs. 3</figureCitation>
,
<figureCitation id="136A2A3AFFB1FFA4FF176694FE95FF55" box="[250,309,186,210]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1499,1522]" captionTargetBox="[161,1430,196,1475]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Aedeagi of holotype males in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Cephennodes lucipetus sp. n. (5 – 6); Cephennodes zulunatalensis sp. n. (7 – 8); Cephennodes mtubatubanus sp. n. (9 – 10); Cephennodes sodwanaensis sp. n. (11 – 12). Abbreviations: ah, apical hook; aml, apex of median lobe; ap, apical projections. Arrows indicate structures important for species identification: a, general shape of median lobe and especially the shape of its left lateral margin in ventral view; b, shape of the apical region of median lobe in ventral view; c, position of parameral apices in relation to the apex of median lobe; d, general shape of apical projections in ventral view; e, shape of apical hook in dorsal and lateral views." httpUri="https://zenodo.org/record/222166/files/figure.png" pageId="6" pageNumber="417">9–10</figureCitation>
,
<figureCitation id="136A2A3AFFB1FFA4FEAD6694FEFDFF55" box="[320,349,186,210]" captionStart="FIGURES 28 – 29" captionStartId="13.[151,264,1645,1668]" captionTargetBox="[168,1436,459,1607]" captionTargetId="figure@13.[151,1436,445,1624]" captionTargetPageId="13" captionText="FIGURES 28 – 29. Distribution of continental Afrotropical Cephenniini (South Africa magnified in right panel). 28 ― genus Cephennodes (a ― C. leonensis Castellini; b ― C. atewaensis Jałoszyński; c ― C. nsukkaensis Jałoszyński; d ― C. bundibugyoensis Jałoszyński; e ― C. glabella Castellini; f ― C. ruandae (Cauchois); g ― C. indifferens Besuchet; h ― C. basilewskyi Besuchet; i ― C. leleupi Besuchet; j ― C. zanzibaricus (Schaufuss); k ― C. marginatus Besuchet; l ― C. zulunatalensis Jałoszyński; m ― C. sodwanaensis Jałoszyński; n ― C. lucipetus Jałoszyński; o ― C. mtubatubanus Jałoszyński); 29 ― genus Cephennomicrus (p ― Cm. zoiai Castellini; r ― Cm. perpusillus (Reitter); s ― Cm. ubomboensis Jałoszyński; t ― Cm. endroedyi Jałoszyński; u ― Cm. transkeianus Jałoszyński." httpUri="https://zenodo.org/record/222170/files/figure.png" pageId="6" pageNumber="417">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34B6534FFB1FFA4FF7A672DFEFEFE01" pageId="6" pageNumber="417" type="materials_examined">
<paragraph id="8BEE36BFFFB1FFA4FF7A672DFEFEFE01" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
<emphasis id="B925EAADFFB1FFA4FF7A672DFB2BFE9C" bold="true" box="[151,1163,258,283]" pageId="6" pageNumber="417">
<typeStatus id="54EA881DFFB1FFA4FF7A672DFF72FE9C" box="[151,210,259,283]" pageId="6" pageNumber="417">Type</typeStatus>
material.
<typeStatus id="54EA881DFFB1FFA4FEA1672DFE1DFE9C" box="[332,445,259,283]" pageId="6" pageNumber="417" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F346762FFFB1FFA4FE27672CFCC7FE9C" box="[458,871,258,283]" name="South Africa" pageId="6" pageNumber="417">REPUBLIC OF SOUTH AFRICA</collectingCountry>
(KwaZulu
<collectingCountry id="F346762FFFB1FFA4FC1C672DFB92FE9C" box="[1009,1074,259,283]" name="South Africa" pageId="6" pageNumber="417">Natal</collectingCountry>
Prov.):
</emphasis>
3, three labels: &quot;S.Afr.: Zululand / Mtubatuba /
<geoCoordinate id="EE655078FFB1FFA4FE456708FDA0FEB9" box="[424,512,294,318]" direction="south" orientation="latitude" pageId="6" pageNumber="417" precision="555" value="-28.29">28.29 S</geoCoordinate>
-
<geoCoordinate id="EE655078FFB1FFA4FDF66708FDD6FEB9" box="[539,630,294,318]" direction="east" orientation="longitude" pageId="6" pageNumber="417" precision="555" value="32.1">32.10 E</geoCoordinate>
&quot; [white, printed]; &quot;
<date id="FFEF107FFFB1FFA4FCBC6708FC1AFEB9" box="[849,954,294,318]" pageId="6" pageNumber="417" value="1974-04-07">7.4.1974</date>
; E-Y: 337 / sifted / leg. Endrödy-Younga&quot; [white, printed]; &quot;
<taxonomicName id="4C514D3CFFB1FFA4FE8A6765FD8EFEE5" box="[359,558,331,354]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAADFFB1FFA4FE8A6765FD8EFEE5" box="[359,558,331,354]" italics="true" pageId="6" pageNumber="417">CEPHENNODES</emphasis>
</taxonomicName>
(s. str.) /
<taxonomicName id="4C514D3CFFB1FFA4FD456765FCE6FEE4" box="[680,838,331,355]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB1FFA4FD456765FCE6FEE4" box="[680,838,331,355]" italics="true" pageId="6" pageNumber="417">mtubatubanus</emphasis>
</taxonomicName>
m. / det. P. Jałoszyński, '13 / HOLOTYPUS' [red, printed] (
<collectionCode id="ED40AE7AFFB1FFA4FF136740FEF2FE01" box="[254,338,366,390]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="417">TMSA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B6534FFB1FFA4FF2A67BCFC01FE75" pageId="6" pageNumber="417" type="diagnosis">
<paragraph id="8BEE36BFFFB1FFA4FF2A67BCFC01FE75" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
<emphasis id="B925EAADFFB1FFA4FF2A67BCFEE1FE2C" bold="true" box="[199,321,402,427]" pageId="6" pageNumber="417">Diagnosis.</emphasis>
Frons and vertex in male without modifications; median lobe of aedeagus with broad and moderately elongate subtriangular and distinctly asymmetrical apical part and narrow but blunt apex; the longest apical projection (apical hook) in lateral view evenly curved dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C34B6534FFB1FFA4FF2A67D0FE3EF9F5" pageId="6" pageNumber="417" type="description">
<paragraph id="8BEE36BFFFB1FFA4FF2A67D0FF6AFDD9" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
<emphasis id="B925EAADFFB1FFA4FF2A67D0FEF9FD90" bold="true" box="[199,345,510,535]" pageId="6" pageNumber="417">Description.</emphasis>
Body of male (
<figureCitation id="136A2A3AFFB1FFA4FDE867D0FDE7FD91" box="[517,583,510,534]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1580,1603]" captionTargetBox="[158,1430,879,1549]" captionTargetId="figure@2.[151,1436,873,1559]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Dorsal habitus of holotype males. Cephennodes lucipetus sp. n. (1); Cephennodes zulunatalensis sp. n. (2); Cephennodes mtubatubanus sp. n. (3); Cephennodes sodwanaensis sp. n. (4)." httpUri="https://zenodo.org/record/222165/files/figure.png" pageId="6" pageNumber="417">Fig. 3</figureCitation>
) moderately strongly convex, oval with feebly marked constriction between pronotum and elytra, dark brown with slightly lighter legs and palps, covered with yellowish vestiture. BL
<quantity id="4CA99B5AFFB1FFA4FA86640CFF66FDD9" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="417" unit="mm" value="1.1">1.10 mm</quantity>
.
</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF2A6444FC86FC41" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL
<quantity id="4CA99B5AFFB1FFA4FB676444FB50FD05" box="[1162,1264,618,642]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="6" pageNumber="417" unit="mm" value="0.18">0.18 mm</quantity>
, HW
<quantity id="4CA99B5AFFB1FFA4FADC6444FA37FD05" box="[1329,1431,618,642]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="417" unit="mm" value="0.28">0.28 mm</quantity>
; vertex and frons confluent, non-modified, in median part only slightly convex; supraantennal tubercles prominent; punctures on head dorsum very fine and inconspicuous; setae short and sparse, recumbent to suberect. Antennae moderately long, with slender proximal part and strongly broadened, distinct club composed of three terminal antennomeres, AnL 0.48, antennomere I 1.5x as long as broad; II only slightly narrower and shorter than I, nearly twice as long as broad; III distinctly narrower and much shorter than II, slightly broader than long; IV–VII subequal in length and width, each as broad as III but slightly longer, 1.1–1.2x as long as broad; VIII about as long as VII but distinctly broader, distinctly transverse; IX much broader and slightly longer than VIII, strongly transverse; X much broader and distinctly longer than IX, strongly transverse; XI about as long as IX–X together, 1.8x as long as broad, broader than X and with blunt apex.
</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF2A65FDFD97FB61" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
Pronotum semi-oval, broadest near middle; PL
<quantity id="4CA99B5AFFB1FFA4FD1C65FCFCFCFC6D" box="[753,860,978,1002]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="417" unit="mm" value="0.33">0.33 mm</quantity>
, PW
<quantity id="4CA99B5AFFB1FFA4FC4D65FCFBAAFC6D" box="[928,1034,978,1002]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="417" unit="mm" value="0.48">0.48 mm</quantity>
. Anterior margin strongly arcuate; lateral margins strongly convex in anterior third and only slightly rounded in posterior half, slightly convergent towards nearly right and sharply-marked posterior pronotal corners; posterior margin deeply bisinuate; lateral margins thickened and sharply demarcated from disc; lateral ante-basal foveae shallow but distinct, each located distinctly (but not much) closer to posterior than to lateral pronotal margin. Punctures on median part of pronotal disc inconspicuous, fine and spare, punctures becoming denser but not larger toward lateral margins of pronotum, punctures on thickened area along lateral margins small but dense and slightly coarse. Setae on pronotal disc moderately long and dense, suberect.
</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF2A62DCFB81FA1D" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
Elytra as convex as pronotum, oval, broadest near anterior third; EL
<quantity id="4CA99B5AFFB1FFA4FBE462DCFBD4FA8D" box="[1033,1140,1266,1290]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="417" unit="mm" value="0.6">0.60 mm</quantity>
, EW
<quantity id="4CA99B5AFFB1FFA4FB5362DCFA8AFA8D" box="[1214,1322,1266,1290]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="417" unit="mm" value="0.5">0.50 mm</quantity>
, EI 1.20; subhumeral lines well-developed and carinate, as long as 0.28x EL; basal elytral foveae located closer to lateral margin of mesoscutellum than to humerus; elytral apices non-modified, separately rounded. Punctures on elytra small and shallow but dense and more distinct than those on pronotal disc, those on anterior half of each elytron separated by spaces subequal to puncture diameters.
<collectingCountry id="F346762FFFB1FFA4FD0B63ACFCBEFA1D" box="[742,798,1410,1434]" name="India" pageId="6" pageNumber="417">Hind</collectingCountry>
wings well developed.
</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF2A6389FCC2FA39" blockId="6.[151,1436,258,2011]" box="[199,866,1446,1471]" pageId="6" pageNumber="417">Legs moderately long and slender; all tibiae nearly straight.</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF2A63E4FDE1F9C9" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
Aedeagus (
<figureCitation id="136A2A3AFFB1FFA4FEA563E4FE64FA65" box="[328,452,1482,1506]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1499,1522]" captionTargetBox="[161,1430,196,1475]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Aedeagi of holotype males in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Cephennodes lucipetus sp. n. (5 – 6); Cephennodes zulunatalensis sp. n. (7 – 8); Cephennodes mtubatubanus sp. n. (9 – 10); Cephennodes sodwanaensis sp. n. (11 – 12). Abbreviations: ah, apical hook; aml, apex of median lobe; ap, apical projections. Arrows indicate structures important for species identification: a, general shape of median lobe and especially the shape of its left lateral margin in ventral view; b, shape of the apical region of median lobe in ventral view; c, position of parameral apices in relation to the apex of median lobe; d, general shape of apical projections in ventral view; e, shape of apical hook in dorsal and lateral views." httpUri="https://zenodo.org/record/222166/files/figure.png" pageId="6" pageNumber="417">Figs. 9–10</figureCitation>
)
<emphasis id="B925EAADFFB1FFA4FE3A63E2FD8CFA64" box="[471,556,1484,1507]" italics="true" pageId="6" pageNumber="417">simonis</emphasis>
<typeStatus id="54EA881DFFB1FFA4FDDA63E5FDCAFA64" box="[567,618,1483,1507]" pageId="6" pageNumber="417">type</typeStatus>
, AeL
<quantity id="4CA99B5AFFB1FFA4FD5863E4FCBFFA65" box="[693,799,1482,1506]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="417" unit="mm" value="0.15">0.15 mm</quantity>
; median lobe with moderately broad subtriangular and blunt apex, apical projections broad, with robust and long apical hook evenly curved dorsally, in ventral view left lateral margin of dorsal projection just beneath apex slightly convex; parameres slender, their apices slightly extending above apex of median lobe.
</paragraph>
<paragraph id="8BEE36BFFFB1FFA4FF2A6074FE3EF9F5" blockId="6.[151,1436,258,2011]" box="[199,414,1626,1650]" pageId="6" pageNumber="417">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34B6534FFB1FFA4FF2A6050FCC8F911" box="[199,872,1662,1687]" pageId="6" pageNumber="417" type="distribution">
<paragraph id="8BEE36BFFFB1FFA4FF2A6050FCC8F911" blockId="6.[151,1436,258,2011]" box="[199,872,1662,1687]" pageId="6" pageNumber="417">
<emphasis id="B925EAADFFB1FFA4FF2A6050FEC0F910" bold="true" box="[199,352,1662,1687]" pageId="6" pageNumber="417">Distribution.</emphasis>
Eastern
<collectingCountry id="F346762FFFB1FFA4FE2F6050FE5BF911" box="[450,507,1662,1686]" name="South Africa" pageId="6" pageNumber="417">RSA</collectingCountry>
, KwaZulu-Natal Prov. (
<figureCitation id="136A2A3AFFB1FFA4FCEA6050FCFBF911" box="[775,859,1662,1686]" captionStart="FIGURES 28 – 29" captionStartId="13.[151,264,1645,1668]" captionTargetBox="[168,1436,459,1607]" captionTargetId="figure@13.[151,1436,445,1624]" captionTargetPageId="13" captionText="FIGURES 28 – 29. Distribution of continental Afrotropical Cephenniini (South Africa magnified in right panel). 28 ― genus Cephennodes (a ― C. leonensis Castellini; b ― C. atewaensis Jałoszyński; c ― C. nsukkaensis Jałoszyński; d ― C. bundibugyoensis Jałoszyński; e ― C. glabella Castellini; f ― C. ruandae (Cauchois); g ― C. indifferens Besuchet; h ― C. basilewskyi Besuchet; i ― C. leleupi Besuchet; j ― C. zanzibaricus (Schaufuss); k ― C. marginatus Besuchet; l ― C. zulunatalensis Jałoszyński; m ― C. sodwanaensis Jałoszyński; n ― C. lucipetus Jałoszyński; o ― C. mtubatubanus Jałoszyński); 29 ― genus Cephennomicrus (p ― Cm. zoiai Castellini; r ― Cm. perpusillus (Reitter); s ― Cm. ubomboensis Jałoszyński; t ― Cm. endroedyi Jałoszyński; u ― Cm. transkeianus Jałoszyński." httpUri="https://zenodo.org/record/222170/files/figure.png" pageId="6" pageNumber="417">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B6534FFB1FFA4FF2A608DFCACF93D" box="[199,780,1698,1723]" pageId="6" pageNumber="417" type="etymology">
<paragraph id="8BEE36BFFFB1FFA4FF2A608DFCACF93D" blockId="6.[151,1436,258,2011]" box="[199,780,1698,1723]" pageId="6" pageNumber="417">
<emphasis id="B925EAADFFB1FFA4FF2A608DFEEEF93C" bold="true" box="[199,334,1699,1723]" pageId="6" pageNumber="417">Etymology.</emphasis>
Locotypical, after the town Mtubatuba.
</paragraph>
</subSubSection>
<subSubSection id="C34B6534FFB1FFA5FF2A60E9FB70FD96" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417" type="discussion">
<paragraph id="8BEE36BFFFB1FFA4FF2A60E9FEEFF85C" blockId="6.[151,1436,258,2011]" pageId="6" pageNumber="417">
<emphasis id="B925EAADFFB1FFA4FF2A60E9FE9BF958" bold="true" box="[199,315,1735,1759]" pageId="6" pageNumber="417">Remarks.</emphasis>
<taxonomicName id="4C514D3CFFB1FFA4FEAE60E9FD20F958" box="[323,640,1735,1759]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB1FFA4FEAE60E9FD20F958" box="[323,640,1735,1759]" italics="true" pageId="6" pageNumber="417">Cephennodes mtubatubanus</emphasis>
</taxonomicName>
is highly similar to all its Afrotropical congeners, and additionally its aedeagus resembles that of
<taxonomicName id="4C514D3CFFB1FFA4FE0860C5FCE7F885" authority="Jaloszynski. The" authorityName="Jaloszynski. The" box="[485,839,1769,1795]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="papuanus" status="sp. nov.">
<emphasis id="B925EAADFFB1FFA4FE0860C5FDD9F884" box="[485,633,1771,1795]" italics="true" pageId="6" pageNumber="417">C. papuanus</emphasis>
Jałoszyński. The
</taxonomicName>
<taxonomicNameLabel id="A21657D6FFB1FFA4FCB860C5FC43F885" box="[853,995,1770,1794]" pageId="6" pageNumber="417" rank="species">new species</taxonomicNameLabel>
can be easily distinguished from
<taxonomicName id="4C514D3CFFB1FFA4FA6F60C5FEA4F8A0" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B925EAADFFB1FFA4FA6F60C5FEA4F8A0" italics="true" pageId="6" pageNumber="417">C. papuanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D3CFFB1FFA4FEFE6121FE61F8A1" box="[275,449,1806,1830]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="zanzibaricus">
<emphasis id="B925EAADFFB1FFA4FEFE6121FE61F8A1" box="[275,449,1806,1830]" italics="true" pageId="6" pageNumber="417">C. zanzibaricus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D3CFFB1FFA4FE3C6121FDD8F8A1" box="[465,632,1806,1830]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="nsukkaensis">
<emphasis id="B925EAADFFB1FFA4FE3C6121FDD8F8A1" box="[465,632,1806,1830]" italics="true" pageId="6" pageNumber="417">C. nsukkaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D3CFFB1FFA4FD6A6121FC84F8A1" box="[647,804,1807,1830]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="atewaensis">
<emphasis id="B925EAADFFB1FFA4FD6A6121FC84F8A1" box="[647,804,1807,1830]" italics="true" pageId="6" pageNumber="417">C. atewaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D3CFFB1FFA4FCB16121FB99F8A1" box="[860,1081,1806,1830]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="bundibugyoensis">
<emphasis id="B925EAADFFB1FFA4FCB16121FB99F8A1" box="[860,1081,1806,1830]" italics="true" pageId="6" pageNumber="417">C. bundibugyoensis</emphasis>
</taxonomicName>
on the basis of a non-modified vertex and frons (all these species have modifications on the head dorsum). From
<taxonomicName id="4C514D3CFFB1FFA4FBDC611DFB0FF8CC" box="[1073,1199,1843,1867]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="ruandae">
<emphasis id="B925EAADFFB1FFA4FBDC611DFB0FF8CC" box="[1073,1199,1843,1867]" italics="true" pageId="6" pageNumber="417">C. ruandae</emphasis>
</taxonomicName>
it differs in the body shape; in
<taxonomicName id="4C514D3CFFB1FFA4FEEF6179FE20F8E9" box="[258,384,1878,1902]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="ruandae">
<emphasis id="B925EAADFFB1FFA4FEEF6179FE20F8E9" box="[258,384,1878,1902]" italics="true" pageId="6" pageNumber="417">C. ruandae</emphasis>
</taxonomicName>
the pronotum is strikingly broader than elytra, while in
<taxonomicName id="4C514D3CFFB1FFA4FC1E6179FB11F8E9" box="[1011,1201,1878,1902]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB1FFA4FC1E6179FB11F8E9" box="[1011,1201,1878,1902]" italics="true" pageId="6" pageNumber="417">C. mtubatubanus</emphasis>
</taxonomicName>
elytra and pronotum have comparable width. Distinguishing
<taxonomicName id="4C514D3CFFB1FFA4FD816155FC91F814" box="[620,817,1915,1939]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB1FFA4FD816155FC91F814" box="[620,817,1915,1939]" italics="true" pageId="6" pageNumber="417">C. mtubatubanus</emphasis>
</taxonomicName>
from the remaining Afrotropical congeners requires examination of the aedeagus. The aedeagus of
<taxonomicName id="4C514D3CFFB1FFA4FD4761B1FCCBF831" box="[682,875,1950,1974]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB1FFA4FD4761B1FCCBF831" box="[682,875,1950,1974]" italics="true" pageId="6" pageNumber="417">C. mtubatubanus</emphasis>
</taxonomicName>
shows the following major differences compared to its congeners:
</paragraph>
<paragraph id="8BEE36BFFFB0FFA5FF7566B6FC56FF53" blockId="7.[152,1436,151,464]" pageId="7" pageNumber="418">
- in
<taxonomicName id="4C514D3CFFB0FFA5FF0A66B6FE2CFF37" box="[231,396,152,176]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="B925EAADFFB0FFA5FF0A66B6FE2CFF37" box="[231,396,152,176]" italics="true" pageId="7" pageNumber="418">C. basilewskyi</emphasis>
</taxonomicName>
the median lobe is shorter and has a differently shaped apex, the apex of apical hook in ventral view is directed to the left, while in
<taxonomicName id="4C514D3CFFB0FFA5FD406693FCCDFF53" box="[685,877,188,212]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB0FFA5FD406693FCCDFF53" box="[685,877,188,212]" italics="true" pageId="7" pageNumber="418">C. mtubatubanus</emphasis>
</taxonomicName>
to the right;
</paragraph>
<paragraph id="8BEE36BFFFB0FFA5FF7566CEFCADFE9B" blockId="7.[152,1436,151,464]" pageId="7" pageNumber="418">
- in
<taxonomicName id="4C514D3CFFB0FFA5FF0B66CEFEF5FF7F" box="[230,341,224,248]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="B925EAADFFB0FFA5FF0B66CEFEF5FF7F" box="[230,341,224,248]" italics="true" pageId="7" pageNumber="418">C. leleupi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D3CFFB0FFA5FE8866CEFE5FFF7F" box="[357,511,224,248]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="indifferens">
<emphasis id="B925EAADFFB0FFA5FE8866CEFE5FFF7F" box="[357,511,224,248]" italics="true" pageId="7" pageNumber="418">C. indifferens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D3CFFB0FFA5FDD466CEFD7BFF7F" box="[569,731,224,248]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B925EAADFFB0FFA5FDD466CEFD7BFF7F" box="[569,731,224,248]" italics="true" pageId="7" pageNumber="418">C. marginatus</emphasis>
</taxonomicName>
the apex of median lobe is broadly rounded, while that in
<taxonomicName id="4C514D3CFFB0FFA5FA6F66CEFEC4FE9B" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB0FFA5FA6F66CEFEC4FE9B" italics="true" pageId="7" pageNumber="418">C. mtubatubanus</emphasis>
</taxonomicName>
is much narrower and nearly pointed;
</paragraph>
<paragraph id="8BEE36BFFFB0FFA5FF756706FD7AFEE3" blockId="7.[152,1436,151,464]" pageId="7" pageNumber="418">
- in
<taxonomicName id="4C514D3CFFB0FFA5FF0E6706FECDFEC7" box="[227,365,296,320]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="leonensis">
<emphasis id="B925EAADFFB0FFA5FF0E6706FECDFEC7" box="[227,365,296,320]" italics="true" pageId="7" pageNumber="418">C. leonensis</emphasis>
</taxonomicName>
the apical part of median lobe is strongly projecting and forming nearly parallel-sided, long and slender rod (subtriangular in
<taxonomicName id="4C514D3CFFB0FFA5FDE76763FD68FEE3" box="[522,712,332,356]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB0FFA5FDE76763FD68FEE3" box="[522,712,332,356]" italics="true" pageId="7" pageNumber="418">C. mtubatubanus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BEE36BFFFB0FFA5FF75675EFDE2FE48" blockId="7.[152,1436,151,464]" pageId="7" pageNumber="418">
- in
<taxonomicName id="4C514D3CFFB0FFA5FF08675EFEC3FE0F" box="[229,355,368,392]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="glabella">
<emphasis id="B925EAADFFB0FFA5FF08675EFEC3FE0F" box="[229,355,368,392]" italics="true" pageId="7" pageNumber="418">C. glabella</emphasis>
</taxonomicName>
the apex of median lobe is very short and pointed (longer and blunt in
<taxonomicName id="4C514D3CFFB0FFA5FB74675EFAFAFE0F" box="[1177,1370,368,392]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB0FFA5FB74675EFAFAFE0F" box="[1177,1370,368,392]" italics="true" pageId="7" pageNumber="418">C. mtubatubanus</emphasis>
</taxonomicName>
), and apices of parameres are extending far above the apex of median lobe (only slightly extending above apex of median lobe in
<taxonomicName id="4C514D3CFFB0FFA5FE9F6796FD92FE57" box="[370,562,440,464]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="species" species="mtubatubanus">
<emphasis id="B925EAADFFB0FFA5FE9F6796FD92FE57" box="[370,562,440,464]" italics="true" pageId="7" pageNumber="418">C. mtubatubanus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BEE36BFFFB0FFA5FF2A67D7FB70FD96" blockId="7.[199,1232,505,529]" box="[199,1232,505,529]" pageId="7" pageNumber="418">
Differences between species of
<taxonomicName id="4C514D3CFFB0FFA5FDC767D7FD61FD96" box="[554,705,505,529]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="418" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAADFFB0FFA5FDC767D7FD61FD96" box="[554,705,505,529]" italics="true" pageId="7" pageNumber="418">Cephennodes</emphasis>
</taxonomicName>
newly described here are shown in
<figureCitation id="136A2A3AFFB0FFA5FBBE67D7FB6CFD96" box="[1107,1228,505,529]" captionStart="FIGURES 5 – 12" captionStartId="4.[151,264,1499,1522]" captionTargetBox="[161,1430,196,1475]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURES 5 – 12. Aedeagi of holotype males in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. Cephennodes lucipetus sp. n. (5 – 6); Cephennodes zulunatalensis sp. n. (7 – 8); Cephennodes mtubatubanus sp. n. (9 – 10); Cephennodes sodwanaensis sp. n. (11 – 12). Abbreviations: ah, apical hook; aml, apex of median lobe; ap, apical projections. Arrows indicate structures important for species identification: a, general shape of median lobe and especially the shape of its left lateral margin in ventral view; b, shape of the apical region of median lobe in ventral view; c, position of parameral apices in relation to the apex of median lobe; d, general shape of apical projections in ventral view; e, shape of apical hook in dorsal and lateral views." httpUri="https://zenodo.org/record/222166/files/figure.png" pageId="7" pageNumber="418">Figs. 5–12</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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