treatments-xml/data/03/F8/3B/03F83B49FFC1FFAFFF0FFEECFC9CFE0F.xml

<document id="0097CA7CC6765B12612B3AB683218D34" ID-CLB-Dataset="8387" ID-DOI="10.11646/zootaxa.4920.2.9" ID-GBIF-Dataset="03094c25-9057-4f2e-8f1a-296aadfca17d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4475313" ID-ZooBank="901B8453-CC3C-4C59-894C-7052186B46EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611821655991" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Maenosono, Tadafumi &amp; Naruse, Tohru" docDate="2021" docId="03F83B49FFC1FFAFFF0FFEECFC9CFE0F" docLanguage="en" docName="zootaxa.4920.2.9.pdf" docOrigin="Zootaxa 4920 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salmoneus singularis Komai &amp; Maenosono &amp; Naruse 2021, n. sp." docType="treatment" docUuid="22FD59C2-669E-45EA-9B56-E4FC55F2C0A5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="294" masterDocId="FFC14331FFC0FFA8FF98FFCBFF9DFF86" masterDocTitle="A new species of alpheid shrimp tentatively assigned to Salmoneus Holthuis, 1955 (Decapoda: Caridea) from the Ryukyu Islands, Japan" masterLastPageNumber="296" masterPageNumber="287" pageNumber="288" updateTime="1698925736767" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A9735A0598A8DE82FCD40D7B7209E11D">A new species of alpheid shrimp tentatively assigned to Salmoneus Holthuis, 1955 (Decapoda: Caridea) from the Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="47DC3A36A11FA05B60CEB3BFDA19BB77">Komai, Tomoyuki</mods:namePart>
<mods:affiliation id="92386A68B21AA31CE75BAEBEDA446BE0">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba 260 - 8682, Japan.</mods:affiliation>
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<mods:namePart id="18AEA3BFB4EDE25DF3103B33390AD2B8">Maenosono, Tadafumi</mods:namePart>
<mods:affiliation id="13FC6FB6508A61317DBB27C9A1D8708D">Kankyosha, 1 - 4 - 5 - 102 Kyozuka, Urasoe, Okinawa 901 - 2111, Japan.</mods:affiliation>
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<mods:namePart id="EA203060EFDCC40BDFAE28CDA4C66F5E">Naruse, Tohru</mods:namePart>
<mods:affiliation id="919FD4463B297C42E7C6A0B6590AE135">Tropical Biosphere Research Center, Iriomote Station, University of the Ryukyus, 870 Uehara, Okinawa 907 - 1541, Japan. naruse @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0001 - 9212 - 5246</mods:affiliation>
<mods:nameIdentifier id="608E0EBD509DAFB09EC28EB4A5A6518B" type="email">naruse@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
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<heading id="D0A63D33FFC1FFA9FF0FFEECFE7AFEC7" bold="true" box="[151,487,295,322]" fontSize="11" level="1" pageId="1" pageNumber="288" reason="1">
<emphasis id="B925564DFFC1FFA9FF0FFEECFE7AFEC7" bold="true" box="[151,487,295,322]" pageId="1" pageNumber="288">
<taxonomicName id="4C51F1DCFFC1FFA9FF0FFEECFE00FEC4" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[151,413,295,322]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="288" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC1FFA9FF0FFEECFE00FEC4" bold="true" box="[151,413,295,322]" italics="true" pageId="1" pageNumber="288">Salmoneus singularis</emphasis>
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<taxonomicNameLabel id="A216EB36FFC1FFA9FE3CFEECFE7AFEC7" box="[420,487,295,321]" pageId="1" pageNumber="288" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BEE8A5FFFC1FFA9FF0FFE87FD4AFEE3" blockId="1.[151,727,295,394]" box="[151,727,330,358]" pageId="1" pageNumber="288">[New Japanese name: Kawari-nokogiri-teppou-ebi]</paragraph>
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<paragraph id="8BEE8A5FFFC1FFA9FF0FFEA4FE8FFE0C" blockId="1.[151,727,295,394]" box="[151,274,367,394]" pageId="1" pageNumber="288">
(
<figureCitation id="136A96DAFFC1FFA9FF07FEA4FE94FE0C" box="[159,265,367,394]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1031,1055]" captionStartId-1="3.[151,250,1490,1514]" captionStartId-2="4.[151,250,1904,1928]" captionStartId-3="6.[151,250,1935,1959]" captionStartId-4="7.[151,250,1073,1097]" captionTargetBox-0="[159,1427,189,998]" captionTargetBox-1="[160,1426,189,1455]" captionTargetBox-2="[193,1371,202,1880]" captionTargetBox-3="[167,1423,609,1897]" captionTargetBox-4="[155,1428,424,1041]" captionTargetId-0="figure-21@2.[151,1436,181,1007]" captionTargetId-1="figure-17@3.[151,1436,181,1466]" captionTargetId-2="figure-21@4.[177,1411,178,1880]" captionTargetId-3="figure-243@6.[156,1431,595,1911]" captionTargetId-4="figure-134@7.[151,1436,417,1050]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, habitus in lateral view." captionText-1="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." captionText-2="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." captionText-3="FIGURE 4. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, right pereopod 1 (major cheliped), lateral view; B, same, coxa to carpus, lateral view in different angle; C, same, chela, extensor view; D, same, carpus and chela, flexor view; E, same, carpus and chela, mesial view." captionText-4="FIGURE 5. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, basal part of left maxilliped 3 and precoxa with arthrobranch, ventral view; B, distal part of ultimate article of left maxilliped 3, lateral view; C–E, distal part of propodus and dactylus of left pereopods 3–5, mesial view." figureDoi-0="http://doi.org/10.5281/zenodo.4475315" figureDoi-1="http://doi.org/10.5281/zenodo.4475317" figureDoi-2="http://doi.org/10.5281/zenodo.4475319" figureDoi-3="http://doi.org/10.5281/zenodo.4475321" figureDoi-4="http://doi.org/10.5281/zenodo.4475323" httpUri-0="https://zenodo.org/record/4475315/files/figure.png" httpUri-1="https://zenodo.org/record/4475317/files/figure.png" httpUri-2="https://zenodo.org/record/4475319/files/figure.png" httpUri-3="https://zenodo.org/record/4475321/files/figure.png" httpUri-4="https://zenodo.org/record/4475323/files/figure.png" pageId="1" pageNumber="288">Figs. 1–5</figureCitation>
)
</paragraph>
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<subSubSection id="C34BD9D4FFC1FFA9FF0FFE7BFD9DFDDD" pageId="1" pageNumber="288" type="materials_examined">
<paragraph id="8BEE8A5FFFC1FFA9FF0FFE7BFBDEFD95" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
<emphasis id="B925564DFFC1FFA9FF0FFE7BFEE1FE4C" bold="true" box="[151,380,432,458]" pageId="1" pageNumber="288">Material examined</emphasis>
. 
<materialsCitation id="3B398002FFC1FFA9FE15FE7BFBDEFD95" ID-GBIF-Occurrence="3032940301" collectingDate="2014-05-14" collectionCode="T" county="southern Ryukyu Islands" latitude="24.3646" location="Uchipanari Islet" longLatPrecision="7" longitude="123.726" municipality="Yaeyama Group" pageId="1" pageNumber="288" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EA34FDFFC1FFA9FE15FE7BFE67FE4C" box="[397,506,432,458]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
: CBM-ZC 16164, non-ovigerous specimen (cl 
<quantity id="4CA927BAFFC1FFA9FB81FE7BFBEBFE4D" box="[1049,1142,432,459]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="288" unit="mm" value="5.4">5.4 mm</quantity>
), NE point of 
<location id="8E8EDC84FFC1FFA9FAB8FE7BFF57FE68" LSID="urn:lsid:plazi:treatment:03F83B49FFC1FFAFFF0FFEECFC9CFE0F:8E8EDC84FFC1FFA9FAB8FE7BFF57FE68" county="southern Ryukyu Islands" latitude="24.3646" longLatPrecision="7" longitude="123.726" municipality="Yaeyama Group" name="Uchipanari Islet" pageId="1" pageNumber="288">Uchipanari Islet</location>
, 
<location id="8E8EDC84FFC1FFA9FF4CFE1FFE1FFE68" LSID="urn:lsid:plazi:treatment:03F83B49FFC1FFAFFF0FFEECFC9CFE0F:8E8EDC84FFC1FFA9FF4CFE1FFE1FFE68" box="[212,386,468,494]" county="southern Ryukyu Islands" latitude="24.3646" longLatPrecision="7" longitude="123.726" municipality="Yaeyama Group" name="Iriomote Island" pageId="1" pageNumber="288">Iriomote Island</location>
, 
<collectingMunicipality id="6B8A1025FFC1FFA9FE13FE1EFDDFFE69" box="[395,578,469,495]" pageId="1" pageNumber="288">Yaeyama Group</collectingMunicipality>
, 
<collectingCounty id="628FF2D3FFC1FFA9FDD5FE1FFCFEFE68" box="[589,867,468,494]" pageId="1" pageNumber="288">southern Ryukyu Islands</collectingCounty>
, 
<geoCoordinate id="EE65EC98FFC1FFA9FCF5FE1FFC76FE68" box="[877,1003,468,494]" degrees="24.3646" direction="north" orientation="latitude" pageId="1" pageNumber="288" precision="5" value="24.3646">24.3646°N</geoCoordinate>
, 
<geoCoordinate id="EE65EC98FFC1FFA9FC6DFE1FFBE0FE68" box="[1013,1149,468,494]" degrees="123.7260" direction="east" orientation="longitude" pageId="1" pageNumber="288" precision="5" value="123.726">123.7260°E</geoCoordinate>
, lower intertidal, seagrass bed, extracted from soft sediments, yabby pump, coll. 
<collectionCode id="ED40129AFFC1FFA9FD60FE32FC97FD95" box="[760,778,505,531]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="288">T</collectionCode>
. Maenosono, 
<date id="FFEFAC9FFFC1FFA9FC3EFE32FBA2FD95" box="[934,1087,505,531]" pageId="1" pageNumber="288" value="2014-05-14">
<collectingDate id="EFAB5577FFC1FFA9FC3EFE32FBA2FD95" box="[934,1087,505,531]" pageId="1" pageNumber="288" value="2014-05-14">14 May 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE8A5FFFC1FFA9FF5FFDD6FD9DFDDD" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
<typeStatus id="54EA34FDFFC1FFA9FF5FFDD6FEA5FDB1" box="[199,312,541,567]" pageId="1" pageNumber="288" type="paratype">Paratypes</typeStatus>
. Same data as holotype: RUMF-ZC-5923, 1 non-ovigerous specimen (cl 
<quantity id="4CA927BAFFC1FFA9FBF6FDD6FB55FDB1" box="[1134,1224,541,567]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="288" unit="mm" value="3.7">3.7 mm</quantity>
); CBM-ZC 16165, 1 ovigerous female (cl 
<quantity id="4CA927BAFFC1FFA9FE01FD8BFE6EFDDD" box="[409,499,576,603]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="288" unit="mm" value="5.8">5.8 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C34BD9D4FFC1FFA9FF5FFDAFFBC0FBF0" pageId="1" pageNumber="288" type="diagnosis">
<paragraph id="8BEE8A5FFFC1FFA9FF5FFDAFFBC0FBF0" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
<emphasis id="B925564DFFC1FFA9FF5FFDAFFEA7FDF8" bold="true" box="[199,314,612,638]" pageId="1" pageNumber="288">Diagnosis</emphasis>
. Medium-sized species of 
<taxonomicName id="4C51F1DCFFC1FFA9FDF4FDAFFD78FDF8" authorityName="Holthuis" authorityYear="1955" box="[620,741,612,638]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC1FFA9FDF4FDAFFD78FDF8" box="[620,741,612,638]" italics="true" pageId="1" pageNumber="288">Salmoneus</emphasis>
</taxonomicName>
(cl of ovigerous specimen 
<quantity id="4CA927BAFFC1FFA9FB84FDAFFBEAFDF9" box="[1052,1143,612,639]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="288" unit="mm" value="5.8">5.8 mm</quantity>
), with moderately slender body. Rostrum 0.3–0.4 times as long as carapace, reaching or overreaching distal end of article 2 of antennular peduncle, narrowly triangular in dorsal view; mid-dorsal carina obsolete; ventral margin with 1 minute subterminal spine. Carapace with strong postorbital spine on each side; orbital spine reduced to small, bluntly triangular projection. Telson with deep, V-shaped, posteromedian notch. Eyes mostly covered by carapace in dorsal view, with only anterolateral part of cornea visible; eyestalk with minute distomesial projection; cornea moderately developed. Antennule with stylocerite strong, reaching mid-length of peduncular article 2. Antenna with carpocerite short, reaching mid-length of scaphocerite. Maxilliped 3 with lateral plate strongly produced dorsally, subacutely pointed. Chelipeds greatly unequal and dissimilar. Major cheliped with ischium unarmed; merus slender, elongate, mesioventrally flattened; palm subcylindrical, not particularly inflated, without grooves; fingers slightly shorter than palm, occlusal margins with 15–16 blunt triangular teeth over entire length. Minor cheliped much smaller and shorter than major cheliped; ischium with 1 spiniform seta; carpus subequal in length to merus; chela shorter than carpus. Pereopod 2 slender; ischium with 2 spiniform setae. Pereopods 3–5 slender; ischium of pereopods 3 and 4 each with 3 spiniform setae; dactylus very slender, exceeding half-length of propodus in pereopods 3 and 4. Second pleopod with appendix masculina elongate, almost reaching distal margin of endopod.
</paragraph>
</subSubSection>
<subSubSection id="C34BD9D4FFC1FFADFF5FFB4BFD93FC45" lastPageId="5" lastPageNumber="292" pageId="1" pageNumber="288" type="description">
<paragraph id="8BEE8A5FFFC1FFA9FF5FFB4BFC38FB1D" blockId="1.[151,1437,432,2043]" box="[199,933,1152,1179]" pageId="1" pageNumber="288">
<emphasis id="B925564DFFC1FFA9FF5FFB4BFECCFB1C" bold="true" box="[199,337,1152,1178]" pageId="1" pageNumber="288">Description</emphasis>
. 
<emphasis id="B925564DFFC1FFA9FEF8FB4BFE53FB1C" box="[352,462,1152,1178]" italics="true" pageId="1" pageNumber="288">
<typeStatus id="54EA34FDFFC1FFA9FEF8FB4BFE57FB1C" box="[352,458,1152,1178]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
.
</emphasis>
Body moderately stout for genus (
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).
</paragraph>
<paragraph id="8BEE8A5FFFC1FFA9FF5FFB6EFDD7FA3C" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
Rostrum (
<figureCitation id="136A96DAFFC1FFA9FEA0FB6FFE2CFB39" box="[312,433,1188,1215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="1" pageNumber="288">Fig. 2A, B</figureCitation>
) relatively long, 0.3 times as long as carapace, overreaching distal margin of article 2 of antennular peduncle, dorsoventrally flattened, narrowly triangular in dorsal view, distinctly longer than wide at base, nearly horizontal in lateral view, without distinct mid-dorsal carina; dorsal margin slightly sloping toward tip; lateral margins faintly concave in dorsal view; tip acute; ventral surface sharply carinate medially, with minute subapical spine. Carapace (
<figureCitation id="136A96DAFFC1FFA9FEF8FAFFFE2CFAC9" box="[352,433,1332,1359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1031,1055]" captionTargetBox="[159,1427,189,998]" captionTargetId="figure-21@2.[151,1436,181,1007]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, habitus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475315" httpUri="https://zenodo.org/record/4475315/files/figure.png" pageId="1" pageNumber="288">Figs. 1</figureCitation>
, 
<figureCitation id="136A96DAFFC1FFA9FE5CFAFEFD96FAC9" box="[452,523,1333,1359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="1" pageNumber="288">2A, B</figureCitation>
) slightly inflated dorsally, surface glabrous, armed with 1 pair of dorsolateral spines in postorbital position; dorsal mid-line without tubercle or spine; cardiac notch moderately deep; orbital spine reduced to small, bluntly triangular projection; anterolateral margin below orbital teeth broadly sinuous; pterygostomial angle broadly rounded.
</paragraph>
<paragraph id="8BEE8A5FFFC1FFA9FF5FFA0FFF78F95C" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
Pleon (
<figureCitation id="136A96DAFFC1FFA9FE83FA0FFEFDFA59" box="[283,352,1476,1503]" captionStart="FIGURE 1" captionStartId="2.[151,250,1031,1055]" captionTargetBox="[159,1427,189,998]" captionTargetId="figure-21@2.[151,1436,181,1007]" captionTargetPageId="2" captionText="FIGURE 1. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, habitus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475315" httpUri="https://zenodo.org/record/4475315/files/figure.png" pageId="1" pageNumber="288">Fig. 1</figureCitation>
) with anterior 3 pleura rounded posteroventrally; pleuron 4 pointed at posteroventral angle, not acute; pleuron 5 with minute posteroventral spine. Pleomere 6 1.2 times as long as high, without articulated posteroventral plate, with faint suture; preanal plate rounded; posterolateral process short, terminating acutely. Telson (
<figureCitation id="136A96DAFFC1FFA9FF68F9FBFED9F9CD" box="[240,324,1584,1611]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="1" pageNumber="288">Fig. 2C</figureCitation>
) moderately slender, tapering distally, 2.6 times as long as maximal (proximal) width; dorsal surface with 2 pairs of small spiniform setae both inserted at some distance from lateral margin, first pair at about telson mid-length, second pair at about 0.7 of telson length; posterior margin with deep V-shaped notch, remaining lateral portions straight, each bearing 2 unequal spiniform setae, mesial much stouter and about 1.3 times longer than lateral.
</paragraph>
<paragraph id="8BEE8A5FFFC1FFA9FF5FF92FFE88F8C0" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
Eyestalks (
<figureCitation id="136A96DAFFC1FFA9FEDFF92FFE6DF979" box="[327,496,1764,1791]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="1" pageNumber="288">Fig. 2A, B, D</figureCitation>
) mostly concealed by anterior part of carapace, with minute dorsomesial distal tubercle; anterolateral part of cornea visible in dorsal view; cornea located at anterolateral part of eyestalk, darkly pigmented.
</paragraph>
<paragraph id="8BEE8A5FFFC1FFA9FF5FF89BFCA3F87C" blockId="1.[151,1437,432,2043]" pageId="1" pageNumber="288">
Antennular peduncle (
<figureCitation id="136A96DAFFC1FFA9FE5BF89BFDA0F8ED" box="[451,573,1872,1899]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="1" pageNumber="288">Fig. 2A, B</figureCitation>
) stout. Article 1 with small, anteriorly directed tooth on ventromesial ridge; stylocerite slender, with acute tip reaching beyond mid-length of article 2, its dorsal margin sharply carinate and broadly convex in lateral view. Article 2 subquadrate, 1.1 times as long as wide, slightly overreaching distal end of antennal scaphocerite. Lateral flagellum (
<figureCitation id="136A96DAFFC1FFA9FD0DF877FD6DF851" box="[661,752,1980,2007]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="1" pageNumber="288">Fig. 2A</figureCitation>
) biramous, with fused portion very short; secondary ramus composed of 4 units with several groups of long aesthetascs.
</paragraph>
<caption id="DF2EDAD7FFC2FFAAFF0FFBCCFF53FBC5" ID-DOI="http://doi.org/10.5281/zenodo.4475315" ID-Zenodo-Dep="4475315" httpUri="https://zenodo.org/record/4475315/files/figure.png" pageId="2" pageNumber="289" startId="2.[151,250,1031,1055]" targetBox="[159,1427,189,998]" targetPageId="2">
<paragraph id="8BEE8A5FFFC2FFAAFF0FFBCCFF53FBC5" blockId="2.[151,1436,1031,1091]" pageId="2" pageNumber="289">
<emphasis id="B925564DFFC2FFAAFF0FFBCCFE88FB99" bold="true" box="[151,277,1031,1055]" pageId="2" pageNumber="289">FIGURE 1.</emphasis>
<taxonomicName id="4C51F1DCFFC2FFAAFE85FBCCFE6AFB99" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[285,503,1031,1055]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="289" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC2FFAAFE85FBCCFE6AFB99" box="[285,503,1031,1055]" italics="true" pageId="2" pageNumber="289">Salmoneus singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC2FFAAFD98FBCCFDA7FB99" bold="true" box="[512,570,1031,1055]" pageId="2" pageNumber="289">
<taxonomicNameLabel id="A216EB36FFC2FFAAFD98FBCCFDA7FB99" box="[512,570,1031,1055]" pageId="2" pageNumber="289" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, habitus in lateral view.
</paragraph>
</caption>
<paragraph id="8BEE8A5FFFC2FFAAFF5FFBB9FC0DFB71" blockId="2.[151,1437,1137,2028]" pageId="2" pageNumber="289">
Antenna (
<figureCitation id="136A96DAFFC2FFAAFEAFFBBAFE4FFB0A" box="[311,466,1137,1164]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="2" pageNumber="289">Fig. 2A, B, E</figureCitation>
) with basicerite bearing small, ventrolateral distal spine; dorsomesial margin broadly bilobed distally, mesial lobe overhanging base of scaphocerite. Scaphocerite subovate with nearly straight lateral margin and gently convex mesial margin; distal blade broadly rounded, reaching slightly beyond small distolateral spine. Carpocerite short, stout, reaching mid-length of scaphocerite.
</paragraph>
<paragraph id="8BEE8A5FFFC2FFAAFF5FFACAFB13FA9D" blockId="2.[151,1437,1137,2028]" box="[199,1166,1281,1308]" pageId="2" pageNumber="289">Epistomial sclerites each with short protuberance. Upper lip with crested median lobe.</paragraph>
<paragraph id="8BEE8A5FFFC2FFAAFF5FFAEEFC52FAC6" blockId="2.[151,1437,1137,2028]" box="[199,975,1317,1344]" pageId="2" pageNumber="289">Mouthparts typical for genus in external observation (not illustrated).</paragraph>
<paragraph id="8BEE8A5FFFC2FFAAFF5FFA82FB3CFA56" blockId="2.[151,1437,1137,2028]" pageId="2" pageNumber="289">
Maxilliped 3 (
<figureCitation id="136A96DAFFC2FFAAFEFEFA82FE54FAE5" box="[358,457,1353,1379]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[193,1371,202,1880]" captionTargetId="figure-21@4.[177,1411,178,1880]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="2" pageNumber="289">Figs. 3A</figureCitation>
, 
<figureCitation id="136A96DAFFC2FFAAFE4DFA82FD85FAE2" box="[469,536,1353,1380]" captionStart="FIGURE 5" captionStartId="7.[151,250,1073,1097]" captionTargetBox="[155,1428,424,1041]" captionTargetId="figure-134@7.[151,1436,417,1050]" captionTargetPageId="7" captionText="FIGURE 5. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, basal part of left maxilliped 3 and precoxa with arthrobranch, ventral view; B, distal part of ultimate article of left maxilliped 3, lateral view; C–E, distal part of propodus and dactylus of left pereopods 3–5, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4475323" httpUri="https://zenodo.org/record/4475323/files/figure.png" pageId="2" pageNumber="289">5A, B</figureCitation>
) moderately slender, reaching distal margin of antennal scaphocerite. Coxa with lateral plate rounded-subquadrate. Antepenultimate article sinuously curved, proximal half flattened dorsoventrally. Penultimate article slightly widened distally. Ultimate article tapering distally from mid-length to acute tip, with 1 prominent subterminal seta; mesial face with several transverse tracts of stiff serrulate setae.
</paragraph>
<paragraph id="8BEE8A5FFFC2FFAAFF5FFA12FC82F991" blockId="2.[151,1437,1137,2028]" pageId="2" pageNumber="289">Pereopods 1 (chelipeds) strongly unequal in size and asymmetrical in shape; major (right) cheliped exceeding twice length of minor (right) cheliped, much more robust.</paragraph>
<paragraph id="8BEE8A5FFFC2FFAAFF5FF9EAFB56F8DD" blockId="2.[151,1437,1137,2028]" pageId="2" pageNumber="289">
Major cheliped (
<figureCitation id="136A96DAFFC2FFAAFEE6F9EAFE22F9BD" box="[382,447,1569,1595]" captionStart="FIGURE 4" captionStartId="6.[151,250,1935,1959]" captionTargetBox="[167,1423,609,1897]" captionTargetId="figure-243@6.[156,1431,595,1911]" captionTargetPageId="6" captionText="FIGURE 4. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, right pereopod 1 (major cheliped), lateral view; B, same, coxa to carpus, lateral view in different angle; C, same, chela, extensor view; D, same, carpus and chela, flexor view; E, same, carpus and chela, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4475321" httpUri="https://zenodo.org/record/4475321/files/figure.png" pageId="2" pageNumber="289">Fig. 4</figureCitation>
A–E) elongate, relatively slender, in full extension overreaching distal margin of antennal scaphocerite by length of chela and carpus. Ischium nearly straight, without spiniform seta; ventrolateral margin bluntly carinate. Merus fairly depressed dorsoventrally, slightly arcuate, slightly widened distally, about twice length of ischium; ventral surface shallowly concave to accommodate flexor surface of chela. Carpus short, widened distally, cup-shaped; distomesial margin produced into blunt projection. Chela 0.75 times as long as carapace, subcylindrical, smooth, without grooves, mesial margin gently sinuous; palm 3 times as long as wide; fingers not particularly elongate, 0.9 times as long as palm, strongly compressed; fingertips strongly curved, crossing, acuminate; occlusal margin of dactylus with 15 narrowly spaced, small, subtriangular, distally subacute or rounded teeth along its entire length; occlusal margin of pollex with 16 teeth similar to those of dactylus.
</paragraph>
<paragraph id="8BEE8A5FFFC2FFAAFF5FF8AEFAECF86D" blockId="2.[151,1437,1137,2028]" pageId="2" pageNumber="289">
Minor cheliped (
<figureCitation id="136A96DAFFC2FFAAFE1AF8AEFE65F8F9" box="[386,504,1893,1919]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[193,1371,202,1880]" captionTargetId="figure-21@4.[177,1411,178,1880]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="2" pageNumber="289">Fig. 3B, C</figureCitation>
) in full extension reaching mid-length of scaphocerite by tip of chela. Ischium with 1 spiniform seta at mid-length of ventrolateral surface. Merus 1.3 times as long as ischium. Carpus subequal in length to merus, widened distally, 5 times as long as distal width. Chela 0.8 times as long as carpus; palm slightly depressed; fingers subequal to palm in length, not crossing distally, cutting edges unarmed, forming thin blade.
</paragraph>
<caption id="DF2EDAD7FFC3FFABFF0FFA19FE62F9D1" ID-DOI="http://doi.org/10.5281/zenodo.4475317" ID-Zenodo-Dep="4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="3" pageNumber="290" startId="3.[151,250,1490,1514]" targetBox="[160,1426,189,1455]" targetPageId="3">
<paragraph id="8BEE8A5FFFC3FFABFF0FFA19FE62F9D1" blockId="3.[151,1436,1490,1623]" pageId="3" pageNumber="290">
<emphasis id="B925564DFFC3FFABFF0FFA19FE89FA6C" bold="true" box="[151,276,1490,1514]" pageId="3" pageNumber="290">FIGURE 2.</emphasis>
<taxonomicName id="4C51F1DCFFC3FFABFE83FA18FE68FA6D" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[283,501,1491,1515]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC3FFABFE83FA18FE68FA6D" box="[283,501,1491,1515]" italics="true" pageId="3" pageNumber="290">Salmoneus singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC3FFABFE64FA18FDABFA6D" bold="true" box="[508,566,1491,1515]" pageId="3" pageNumber="290">
<taxonomicNameLabel id="A216EB36FFC3FFABFE64FA18FDABFA6D" box="[508,566,1491,1515]" pageId="3" pageNumber="290" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane).
</paragraph>
</caption>
<paragraph id="8BEE8A5FFFC3FFABFF5FF94EFD78F8A9" blockId="3.[151,1437,1669,2020]" pageId="3" pageNumber="290">
Pereopod 2 (
<figureCitation id="136A96DAFFC3FFABFECBF94EFE34F919" box="[339,425,1669,1695]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[193,1371,202,1880]" captionTargetId="figure-21@4.[177,1411,178,1880]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="3" pageNumber="290">Fig. 3D</figureCitation>
) slender, moderately long, distinctly longer than minor cheliped, not reaching distal margin of scaphocerite. Basis unarmed. Ischium with 2 spiniform setae on lateral surface ventrally. Merus slightly longer than ischium. Carpus about 0.7 times as long as ischium and merus combined, subdivided into 5 articles, ratio of carpal subarticles from proximal to distal equal to: 3.1: 0.7: 0.5: 0.5: 1.0. Chela 1.8 times as long as distal-most carpal subarticle; dactylus 1.7 times as long as palm.
</paragraph>
<paragraph id="8BEE8A5FFFC3FFADFF5FF8F2FEB8FE0E" blockId="3.[151,1437,1669,2020]" lastBlockId="5.[151,1437,151,2036]" lastPageId="5" lastPageNumber="292" pageId="3" pageNumber="290">
Pereopods 3–5 relatively slender, moderately long. Pereopod 3 (
<figureCitation id="136A96DAFFC3FFABFC00F8F2FB88F8D5" box="[920,1045,1849,1875]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[193,1371,202,1880]" captionTargetId="figure-21@4.[177,1411,178,1880]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="3" pageNumber="290">Figs. 3E, F</figureCitation>
, 
<figureCitation id="136A96DAFFC3FFABFBB8F8F2FBDFF8D2" box="[1056,1090,1849,1876]" captionStart="FIGURE 5" captionStartId="7.[151,250,1073,1097]" captionTargetBox="[155,1428,424,1041]" captionTargetId="figure-134@7.[151,1436,417,1050]" captionTargetPageId="7" captionText="FIGURE 5. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, basal part of left maxilliped 3 and precoxa with arthrobranch, ventral view; B, distal part of ultimate article of left maxilliped 3, lateral view; C–E, distal part of propodus and dactylus of left pereopods 3–5, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4475323" httpUri="https://zenodo.org/record/4475323/files/figure.png" pageId="3" pageNumber="290">5C</figureCitation>
) overreaching distal margin of scaphocerite by length of dactylus and 0.2 of propodus; ischium with 3 spiniform setae on ventrolateral surface; merus about 14 times as long as wide, unarmed; carpus slenderer than merus, slightly shorter, unarmed, subequal in length to propodus; propodus with 3 widely spaced, minute, slender, spiniform setae on flexor margin and 2 longer spiniform seta at flexor distal margin; dactylus about 0.5 times as long as propodus, very slender (11.3 times as long as wide), simple, gently curving, with tuft of minute setae at distal 0.3 of dactylar length (near demarcation of unguis); unguis elongate, occupying 0.3 length of dactylus. Pereopod 4 (
<figureCitation id="136A96DAFFC5FFADFB95FF70FBE9FF53" box="[1037,1140,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[193,1371,202,1880]" captionTargetId="figure-21@4.[177,1411,178,1880]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="5" pageNumber="292">Figs. 3G</figureCitation>
, 
<figureCitation id="136A96DAFFC5FFADFB1CFF71FB3AFF53" box="[1156,1191,186,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1073,1097]" captionTargetBox="[155,1428,424,1041]" captionTargetId="figure-134@7.[151,1436,417,1050]" captionTargetPageId="7" captionText="FIGURE 5. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, basal part of left maxilliped 3 and precoxa with arthrobranch, ventral view; B, distal part of ultimate article of left maxilliped 3, lateral view; C–E, distal part of propodus and dactylus of left pereopods 3–5, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4475323" httpUri="https://zenodo.org/record/4475323/files/figure.png" pageId="5" pageNumber="292">5D</figureCitation>
) similar to pereopod 3, overreaching scaphocerite by length of dactylus; ischium with 3 spiniform setae. Pereopod 5 (
<figureCitation id="136A96DAFFC5FFADFB66FF14FAF9FF7F" box="[1278,1380,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[193,1371,202,1880]" captionTargetId="figure-21@4.[177,1411,178,1880]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="5" pageNumber="292">Figs. 3H</figureCitation>
, 
<figureCitation id="136A96DAFFC5FFADFAECFF15FA09FF7F" box="[1396,1428,222,249]" captionStart="FIGURE 5" captionStartId="7.[151,250,1073,1097]" captionTargetBox="[155,1428,424,1041]" captionTargetId="figure-134@7.[151,1436,417,1050]" captionTargetPageId="7" captionText="FIGURE 5. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, basal part of left maxilliped 3 and precoxa with arthrobranch, ventral view; B, distal part of ultimate article of left maxilliped 3, lateral view; C–E, distal part of propodus and dactylus of left pereopods 3–5, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4475323" httpUri="https://zenodo.org/record/4475323/files/figure.png" pageId="5" pageNumber="292">5E</figureCitation>
) distinctly longer than pereopods 3 and 4 (in particular, length of carpus-propodus combined 1.3 times of that of pereopods 3 and 4); propodus with grooming apparatus consisting of setal brush on distal half of flexor margin and 2 slender spiniform seta on flexor distal margin; dactylus similar to those of pereopods 3 and 4, 0.4 times as long as propodus.
</paragraph>
<caption id="DF2EDAD7FFC4FFACFF0FF8BBFE2CF872" ID-DOI="http://doi.org/10.5281/zenodo.4475319" ID-Zenodo-Dep="4475319" httpUri="https://zenodo.org/record/4475319/files/figure.png" pageId="4" pageNumber="291" startId="4.[151,250,1904,1928]" targetBox="[193,1371,202,1880]" targetPageId="4">
<paragraph id="8BEE8A5FFFC4FFACFF0FF8BBFE2CF872" blockId="4.[151,1437,1904,2036]" pageId="4" pageNumber="291">
<emphasis id="B925564DFFC4FFACFF0FF8BBFE85F80E" bold="true" box="[151,280,1904,1928]" pageId="4" pageNumber="291">FIGURE 3.</emphasis>
<taxonomicName id="4C51F1DCFFC4FFACFEBBF8BBFD9CF80E" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[291,513,1904,1928]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="291" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC4FFACFEBBF8BBFD9CF80E" box="[291,513,1904,1928]" italics="true" pageId="4" pageNumber="291">Salmoneus singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC4FFACFD95F8BBFDD6F80E" bold="true" box="[525,587,1904,1928]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="A216EB36FFC4FFACFD95F8BBFDD6F80E" box="[525,587,1904,1928]" pageId="4" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164, left thoracic appendages. A, maxilliped 3, lateral view; B, pereopod 1 (minor cheliped), lateral view; C, same, chela, extensor view; D, pereopod 2, lateral view; E, pereopod 3, lateral view; F, same, propodus and dactylus, lateral view; G, pereopod 4, lateral view; H, pereopod 5, lateral view.
</paragraph>
</caption>
<paragraph id="8BEE8A5FFFC5FFADFF5FFE5AFB0FFE49" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
Pleopod 2 (
<figureCitation id="136A96DAFFC5FFADFEDEFE5AFE04FE2A" box="[326,409,401,428]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="5" pageNumber="292">Fig. 2F</figureCitation>
) with appendix masculina elongate, distinctly longer than appendix interna, almost reaching distal margin of endopod, with some short setae distally; appendix interna curved mesially.
</paragraph>
<paragraph id="8BEE8A5FFFC5FFADFF5FFE12FDF2FDD9" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
Uropod (
<figureCitation id="136A96DAFFC5FFADFEB7FE12FE15FE75" box="[303,392,473,499]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="5" pageNumber="292">Fig. 2G</figureCitation>
) with lateral lobe of protopod terminating in minute spine. Endopod and exopod narrowly ovoid, former slightly shorter than latter. Exopod with straight lateral margin terminating posteriorly in minute blunt spine, latter reaching slightly beyond base of small distolateral spiniform seta; diaeresis sinuous, without noticeable tooth mesial to distolateral spiniform seta.
</paragraph>
<paragraph id="8BEE8A5FFFC5FFADFF5FFDA3FA04FD6B" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
Gill-exopod formula typical for genus, consisting of 1 arthrobranch above base of maxilliped 3 (well developed, multi-lamellate; 
<figureCitation id="136A96DAFFC5FFADFECCFD47FE2DFD20" box="[340,432,651,678]" captionStart="FIGURE 5" captionStartId="7.[151,250,1073,1097]" captionTargetBox="[155,1428,424,1041]" captionTargetId="figure-134@7.[151,1436,417,1050]" captionTargetPageId="7" captionText="FIGURE 5. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, basal part of left maxilliped 3 and precoxa with arthrobranch, ventral view; B, distal part of ultimate article of left maxilliped 3, lateral view; C–E, distal part of propodus and dactylus of left pereopods 3–5, mesial view." figureDoi="http://doi.org/10.5281/zenodo.4475323" httpUri="https://zenodo.org/record/4475323/files/figure.png" pageId="5" pageNumber="292">Fig. 5A</figureCitation>
), 1 pleurobranch on each thoracomeres 4–8 (pereopods 1–5), epipods on maxilliped 1 through pereopod 4 (those on maxilliped 3 to pereopod 4 strap-like with terminal hook), flagellum-like exopod on maxilliped 1–3 and setobranch on each pereopod 1–5, corresponding to epipod on maxilliped 3 and pereopod 1–4.
</paragraph>
<paragraph id="8BEE8A5FFFC5FFADFF5FFD33FF4AFC26" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFD33FEA4FC97" box="[199,313,760,785]" italics="true" pageId="5" pageNumber="292">
<typeStatus id="54EA34FDFFC5FFADFF5FFD33FEA4FC97" box="[199,313,760,785]" pageId="5" pageNumber="292" type="paratype">Paratypes</typeStatus>
</emphasis>
. Two 
<typeStatus id="54EA34FDFFC5FFADFEE7FD3CFE42FC97" box="[383,479,759,785]" pageId="5" pageNumber="292" type="paratype">paratype</typeStatus>
specimens, including one ovigerous and one non-ovigerous specimens, are generally similar to 
<typeStatus id="54EA34FDFFC5FFADFE93FCD0FEECFCB3" box="[267,369,795,821]" pageId="5" pageNumber="292" type="holotype">holotype</typeStatus>
. Major cheliped clearly in process of regeneration in ovigerous specimen and missing in nonovigerous specimen. Rostrum reaching or overreaching distal margin of article 2 of antennular peduncle. Minor cheliped ischium armed with either 1 or 2 spiniform setae. Pereopods 3 and 4 ischia each armed with 3 spiniform setae.
</paragraph>
<paragraph id="8BEE8A5FFFC5FFADFF5FFC62FD93FC45" blockId="5.[151,1437,151,2036]" box="[199,526,937,964]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFC62FEF8FC45" bold="true" box="[199,357,937,963]" pageId="5" pageNumber="292">Colour in life</emphasis>
. Not recorded.
</paragraph>
</subSubSection>
<subSubSection id="C34BD9D4FFC5FFADFF5FFC06FB90FB8A" pageId="5" pageNumber="292" type="etymology">
<paragraph id="8BEE8A5FFFC5FFADFF5FFC06FB90FB8A" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFC06FED5FC61" bold="true" box="[199,328,973,999]" pageId="5" pageNumber="292">Etymology</emphasis>
. The Latin adjective 
<taxonomicName id="4C51F1DCFFC5FFADFDADFC06FD38FC61" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[565,677,973,999]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="B925564DFFC5FFADFDADFC06FD38FC61" box="[565,677,973,999]" italics="true" pageId="5" pageNumber="292">singularis</emphasis>
</taxonomicName>
refers to the uniqueness of the new species within 
<taxonomicName id="4C51F1DCFFC5FFADFB44FC06FAC8FC61" authorityName="Holthuis" authorityYear="1955" box="[1244,1365,973,999]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFB44FC06FAC8FC61" box="[1244,1365,973,999]" italics="true" pageId="5" pageNumber="292">Salmoneus</emphasis>
</taxonomicName>
, more precisely in possessing a pair of postorbital spines on the carapace (see below).
</paragraph>
</subSubSection>
<subSubSection id="C34BD9D4FFC5FFADFF5FFBDEFE99FBD4" pageId="5" pageNumber="292" type="materials_examined">
<paragraph id="8BEE8A5FFFC5FFADFF5FFBDEFE99FBD4" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFBDEFEF9FBA8" bold="true" box="[199,356,1044,1071]" pageId="5" pageNumber="292">Type locality</emphasis>
. Uchipanari Islet, Iriomote Island, Yaeyama Islands, southern Ryukyu Islands, 
<collectingCountry id="F346CACFFFC5FFADFA91FBDEFAD1FBA9" box="[1289,1356,1045,1071]" name="Japan" pageId="5" pageNumber="292">Japan</collectingCountry>
; lower intertidal.
</paragraph>
</subSubSection>
<subSubSection id="C34BD9D4FFC5FFADFF5FFB97FB58FB1C" pageId="5" pageNumber="292" type="distribution">
<paragraph id="8BEE8A5FFFC5FFADFF5FFB97FB38FBF1" blockId="5.[151,1437,151,2036]" box="[199,1189,1116,1143]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFB97FEC4FBF0" bold="true" box="[199,345,1116,1142]" pageId="5" pageNumber="292">Distribution</emphasis>
. Presently known only from the 
<typeStatus id="54EA34FDFFC5FFADFD51FB96FD64FBF1" box="[713,761,1117,1143]" pageId="5" pageNumber="292">type</typeStatus>
locality in the Ryukyu Islands, 
<collectingCountry id="F346CACFFFC5FFADFBC5FB96FB3DFBF1" box="[1117,1184,1117,1143]" name="Japan" pageId="5" pageNumber="292">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BEE8A5FFFC5FFADFF5FFB4BFB58FB1C" blockId="5.[151,1437,151,2036]" box="[199,1221,1152,1179]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFB4BFEBEFB1C" bold="true" box="[199,291,1152,1178]" pageId="5" pageNumber="292">Habitat</emphasis>
. Extracted from soft sediments on a seagrass bed, suggesting an infaunal life style.
</paragraph>
</subSubSection>
<subSubSection id="C34BD9D4FFC5FFAFFF5FFB6FFC9CFE0F" lastPageId="7" lastPageNumber="294" pageId="5" pageNumber="292" type="discussion">
<paragraph id="8BEE8A5FFFC5FFADFF5FFB6FFE02F9A2" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
<emphasis id="B925564DFFC5FFADFF5FFB6FFEA9FB38" bold="true" box="[199,308,1188,1214]" pageId="5" pageNumber="292">Remarks</emphasis>
. The possession of a pair of postorbital spines on the carapace is clearly the most diagnostic feature of 
<taxonomicName id="4C51F1DCFFC5FFADFF2DFB0CFE3BFB67" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[181,422,1223,1249]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC5FFADFF2DFB0CFE3BFB67" box="[181,422,1223,1249]" italics="true" pageId="5" pageNumber="292">Salmoneus singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC5FFADFE34FB03FE76FB64" bold="true" box="[428,491,1224,1250]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A216EB36FFC5FFADFE34FB03FE76FB64" box="[428,491,1224,1250]" pageId="5" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, separating it from all other species in the genus. Considering the combination of all remaining morphological features, the new species shows substantial affinity to several species referred to the 
<taxonomicName id="4C51F1DCFFC5FFADFA1FFB27FE0FFAAF" authority="Miya, 1972" authorityName="Miya" authorityYear="1972" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B925564DFFC5FFADFA1FFB27FE9AFAAF" italics="true" pageId="5" pageNumber="292">S. gracilipes</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFE88FAC4FE0FFAAF" author="Miya, Y." box="[272,402,1295,1321]" pageId="5" pageNumber="292" pagination="23 - 101" refId="ref6037" refString="Miya, Y. (1972) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3 (1), 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
</taxonomicName>
species group, i.e., 
<taxonomicName id="4C51F1DCFFC5FFADFDEEFAC4FBBBFAAF" authority="Anker &amp; Marin, 2006" authorityName="Anker &amp; Marin" authorityYear="2006" box="[630,1062,1295,1321]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="alpheophilus">
<emphasis id="B925564DFFC5FFADFDEEFAC4FCB9FAAF" box="[630,804,1295,1321]" italics="true" pageId="5" pageNumber="292">S. alpheophilus</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFCB3FAC4FBBBFAAF" author="Anker, A. &amp; Marin, I. N." box="[811,1062,1295,1321]" pageId="5" pageNumber="292" pagination="295 - 319" refId="ref4994" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea: Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54 (2), 295 - 319." type="journal article" year="2006">Anker &amp; Marin, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFBABFAC4FF4FFACB" authority="Felder &amp; Manning, 1986" authorityName="Felder &amp; Manning" authorityYear="1986" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="cavicolus">
<emphasis id="B925564DFFC5FFADFBABFAC4FB27FAAF" box="[1075,1210,1295,1321]" italics="true" pageId="5" pageNumber="292">S. cavicolus</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFB5AFAC4FF4FFACB" author="Felder, D. L. &amp; Manning, R. B." pageId="5" pageNumber="292" pagination="497 - 508" refId="ref5527" refString="Felder, D. L. &amp; Manning, R. B. (1986) A new genus and two new species of alpheid shrimps (Decapoda: Caridea) from South Florida. Journal of Crustacean Biology, 6 (3), 497 - 508. https: // doi. org / 10.1163 / 193724086 X 00343" type="journal article" year="1986">Felder &amp; Manning, 1986</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFF44FAF8FDBEFACB" authority="De Grave, 2004" authorityName="De Grave" authorityYear="2004" box="[220,547,1331,1357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="colinorum">
<emphasis id="B925564DFFC5FFADFF44FAF8FEF7FACB" box="[220,362,1331,1357]" italics="true" pageId="5" pageNumber="292">S. colinorum</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFEE8FAF8FDBEFACB" author="De Grave, S." box="[368,547,1331,1357]" pageId="5" pageNumber="292" pagination="41 - 48" refId="ref5250" refString="De Grave, S. (2004) A new species of Salmoneus (Crustacea: Decapoda: Alpheidae) from Palau. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie, 74, 41 - 48." type="journal article" year="2004">De Grave, 2004</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFDB5FAF8FC4DFACB" authority="Anker &amp; Marin, 2006" authorityName="Anker &amp; Marin" authorityYear="2006" box="[557,976,1331,1357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="falcidactylus">
<emphasis id="B925564DFFC5FFADFDB5FAF8FD4AFACB" box="[557,727,1331,1357]" italics="true" pageId="5" pageNumber="292">S. falcidactylus</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFD44FAF8FC4DFACB" author="Anker, A. &amp; Marin, I. N." box="[732,976,1331,1357]" pageId="5" pageNumber="292" pagination="295 - 319" refId="ref4994" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea: Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54 (2), 295 - 319." type="journal article" year="2006">Anker &amp; Marin, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFC42FAF8FBF8FACB" authorityName="Miya" authorityYear="1972" box="[986,1125,1331,1357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B925564DFFC5FFADFC42FAF8FBF8FACB" box="[986,1125,1331,1357]" italics="true" pageId="5" pageNumber="292">S. gracilipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFBEAFAF8FF4FFAF6" authority="Anker &amp; Marin, 2006" authorityName="Anker &amp; Marin" authorityYear="2006" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B925564DFFC5FFADFBEAFAF8FB79FACB" box="[1138,1252,1331,1357]" italics="true" pageId="5" pageNumber="292">S. pusillus</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFB71FAF8FF4FFAF6" author="Anker, A. &amp; Marin, I. N." pageId="5" pageNumber="292" pagination="295 - 319" refId="ref4994" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea: Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54 (2), 295 - 319." type="journal article" year="2006">Anker &amp; Marin, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFF47FA9CFD4FFAF6" authority="Ashrafi, Duris &amp; Naderloo, 2020" authorityName="Ashrafi, Duris &amp; Naderloo" authorityYear="2020" box="[223,722,1366,1395]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="rashedi">
<emphasis id="B925564DFFC5FFADFF47FA9CFECCFAF6" box="[223,337,1366,1393]" italics="true" pageId="5" pageNumber="292">S. rashedi</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFEC0FA9DFD4FFAF6" author="Ashrafi, H. &amp; Duris, Z. &amp; Naderloo, R." box="[344,722,1366,1395]" pageId="5" pageNumber="292" pagination="503 - 514" refId="ref5045" refString="Ashrafi, H., Duris, Z. &amp; Naderloo, R. (2020) A new species and a new record of alpheid shrimps of the genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) from Iran. Zootaxa, 4838 (4), 503 - 514. https: // doi. org / 10.11646 / zootaxa. 4838.4.4" type="journal article" year="2020">Ashrafi, Ďuriš &amp; Naderloo, 2020</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C51F1DCFFC5FFADFC95FA9CFB29FAF6" authority="Banner &amp; Banner, 1966" authorityName="Banner &amp; Banner" authorityYear="1966" box="[781,1204,1366,1393]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="tafaongae">
<emphasis id="B925564DFFC5FFADFC95FA9CFC06FAF6" box="[781,923,1366,1393]" italics="true" pageId="5" pageNumber="292">S. tafaongae</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFC3CFA9CFB29FAF6" author="Banner, A. H. &amp; Banner, D. M." box="[932,1204,1366,1393]" pageId="5" pageNumber="292" pagination="145 - 188" refId="ref5166" refString="Banner, A. H. &amp; Banner, D. M. (1966) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part X. Collections from Fiji, Tonga and Samoa. Pacific Science, 20, 145 - 188." type="journal article" year="1966">Banner &amp; Banner, 1966</bibRefCitation>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC0F7AEFFC5FFADFB72FA9DFF4EFA12" author="Anker, A. &amp; Marin, I. N." pageId="5" pageNumber="292" pagination="295 - 319" refId="ref4994" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea: Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54 (2), 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC5FFADFF46FAB1FE24FA12" author="Ashrafi, H. &amp; Duris, Z. &amp; Naderloo, R." box="[222,441,1402,1428]" pageId="5" pageNumber="292" pagination="503 - 514" refId="ref5045" refString="Ashrafi, H., Duris, Z. &amp; Naderloo, R. (2020) A new species and a new record of alpheid shrimps of the genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) from Iran. Zootaxa, 4838 (4), 503 - 514. https: // doi. org / 10.11646 / zootaxa. 4838.4.4" type="journal article" year="2020">
Ashrafi 
<emphasis id="B925564DFFC5FFADFEA3FAB0FEE8FA12" box="[315,373,1402,1428]" italics="true" pageId="5" pageNumber="292">et al.</emphasis>
2020
</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC5FFADFE5EFAB0FD21FA12" author="De Grave, S. &amp; Park, J. - H. &amp; Anker, A." box="[454,700,1402,1429]" pageId="5" pageNumber="292" pagination="173 - 186" refId="ref5346" refString="De Grave, S., Park, J. - H. &amp; Anker, A. (2020) A redescription of Salmoneus tafaongae Banner &amp; Banner, 1966 (Malacostraca: Decapoda: Alpheidae). Crustacean Research, 49, 173 - 186. https: // doi. org / 10.18353 / crustacea. 49.0 _ 173" type="journal article" year="2020">
De Grave 
<emphasis id="B925564DFFC5FFADFDA4FAB0FDEBFA12" box="[572,630,1402,1428]" italics="true" pageId="5" pageNumber="292">et al.</emphasis>
2020
</bibRefCitation>
). Shared characters are the relatively long, slender rostrum, with a small subdistal spine on the ventral margin; the dorsally partly concealed eyes (with only the anterior portion of the córneas being visible in dorsal view); the slender dactyli of the pereopods 3–5; and the strongly asymmetrical chelipeds, with the major cheliped fingers armed with a row of teeth over the entire length of the occlusal margins (
<bibRefCitation id="EFC0F7AEFFC5FFADFF07F9C2FE0EF9A2" author="Anker, A. &amp; Marin, I. N." box="[159,403,1545,1572]" pageId="5" pageNumber="292" pagination="295 - 319" refId="ref4994" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea: Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54 (2), 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
).
</paragraph>
<paragraph id="8BEE8A5FFFC5FFADFF5FF9E6FCF1F82A" blockId="5.[151,1437,151,2036]" pageId="5" pageNumber="292">
The presence of postorbital spines on the carapace makes the generic placement of the present new species somewhat problematic. The presence of these spines would seem to suggest a possible relationship of 
<taxonomicName id="4C51F1DCFFC5FFADFA88F99AFA01F9EC" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[1296,1436,1616,1643]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC5FFADFA88F99AFA01F9EC" box="[1296,1436,1616,1643]" italics="true" pageId="5" pageNumber="292">S. singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC5FFADFF0FF9BFFF4BF908" bold="true" box="[151,214,1652,1678]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A216EB36FFC5FFADFF0FF9BFFF4BF908" box="[151,214,1652,1678]" pageId="5" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
with some members of 
<taxonomicName id="4C51F1DCFFC5FFADFE7CF9BFFD0FF908" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[484,658,1652,1678]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFE7CF9BFFD0FF908" box="[484,658,1652,1678]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
, such as 
<taxonomicName id="4C51F1DCFFC5FFADFD60F9BEFA05F908" authority="Alvarez, Iliffe, Gonzalez &amp; Villalobos, 2012" authorityName="Alvarez, Iliffe, Gonzalez &amp; Villalobos" authorityYear="2012" box="[760,1432,1652,1678]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="akumalensis">
<emphasis id="B925564DFFC5FFADFD60F9BEFC03F908" box="[760,926,1652,1678]" italics="true" pageId="5" pageNumber="292">T. akumalensis</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFC3CF9BFFA05F908" author="Alvarez, F. &amp; Iliffe, T. M. &amp; Gonzalez, B. &amp; Villalobos, J. L." box="[932,1432,1652,1678]" pageId="5" pageNumber="292" pagination="61 - 68" refId="ref4180" refString="Alvarez, F., Iliffe, T. M., Gonzalez, B. &amp; Villalobos, J. L. (2012) Triacanthoneus akumalensis, a new species of alpheid shrimp (Crustacea: Caridea: Alpheidae) from an anchialine cave in Quintana Roo, Mexico. Zootaxa, 3154 (1), 61 - 68. https: // doi. org / 10.11646 / zootaxa. 3154.1.5" type="journal article" year="2012">Alvarez, Iliffe, Gonzalez &amp; Villalobos, 2012</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFF0FF952FE5EF934" authority="Anker, 2010 b" authorityName="Anker" authorityYear="2010" box="[151,451,1688,1714]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="alacranes">
<emphasis id="B925564DFFC5FFADFF0FF952FEBCF934" box="[151,289,1688,1714]" italics="true" pageId="5" pageNumber="292">T. alacranes</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFEBFF953FE5EF934" author="Anker, A." box="[295,451,1688,1714]" pageId="5" pageNumber="292" pagination="47 - 63" refId="ref4377" refString="Anker, A. (2010 b) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker, 2010b</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFE57F952FD4BF934" authority="Anker, 2020 a" authorityName="Anker" authorityYear="2020" box="[463,726,1688,1714]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="blanca">
<emphasis id="B925564DFFC5FFADFE57F952FDABF934" box="[463,566,1688,1714]" italics="true" pageId="5" pageNumber="292">T. blanca</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFDA4F953FD4BF934" author="Anker, A." box="[572,726,1688,1714]" pageId="5" pageNumber="292" pagination="450 - 468" refId="ref4609" refString="Anker, A. (2020 a) Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010, with description of a second eastern Pacific species (Malacostraca: Decapoda). Zootaxa, 4772 (3), 450 - 468. https: // doi. org / 10.11646 / zootaxa. 4772.3.2" type="journal article" year="2020">Anker, 2020a</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC5FFADFD7AF952FB9FF934" authority="Anker, 2010 b" authorityName="Anker" authorityYear="2010" box="[738,1026,1688,1714]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B925564DFFC5FFADFD7AF952FCFEF934" box="[738,867,1688,1714]" italics="true" pageId="5" pageNumber="292">T. pacificus</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFCF1F953FB9FF934" author="Anker, A." box="[873,1026,1688,1714]" pageId="5" pageNumber="292" pagination="47 - 63" refId="ref4377" refString="Anker, A. (2010 b) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker, 2010b</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C51F1DCFFC5FFADFBA1F952FAB8F934" authority="Anker, 2010 b" authorityName="Anker" authorityYear="2010" box="[1081,1317,1688,1714]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="toro">
<emphasis id="B925564DFFC5FFADFBA1F952FB1EF934" box="[1081,1155,1689,1714]" italics="true" pageId="5" pageNumber="292">T. toro</emphasis>
<bibRefCitation id="EFC0F7AEFFC5FFADFB10F953FAB8F934" author="Anker, A." box="[1160,1317,1688,1714]" pageId="5" pageNumber="292" pagination="47 - 63" refId="ref4377" refString="Anker, A. (2010 b) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker, 2010b</bibRefCitation>
</taxonomicName>
. However, the position of these spines is somewhat different between 
<taxonomicName id="4C51F1DCFFC5FFADFCACF977FC5CF950" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[820,961,1724,1750]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC5FFADFCACF977FC5CF950" box="[820,961,1724,1750]" italics="true" pageId="5" pageNumber="292">S. singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC5FFADFC51F977FB95F950" bold="true" box="[969,1032,1724,1750]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A216EB36FFC5FFADFC51F977FB95F950" box="[969,1032,1724,1750]" pageId="5" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and the afore-mentioned species of 
<taxonomicName id="4C51F1DCFFC5FFADFF0FF914FED8F97F" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[151,325,1759,1785]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFF0FF914FED8F97F" box="[151,325,1759,1785]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
. In five species of 
<taxonomicName id="4C51F1DCFFC5FFADFD87F914FD50F97F" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[543,717,1759,1785]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFD87F914FD50F97F" box="[543,717,1759,1785]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
, they are in “hepatic” or “post-hepatic” position (
<bibRefCitation id="EFC0F7AEFFC5FFADFA98F914FA05F97F" author="Anker, A." box="[1280,1432,1759,1785]" pageId="5" pageNumber="292" pagination="47 - 63" refId="ref4377" refString="Anker, A. (2010 b) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010b</bibRefCitation>
, 
<bibRefCitation id="EFC0F7AEFFC5FFADFF0FF8C8FF7DF89B" author="Anker, A." box="[151,224,1795,1821]" pageId="5" pageNumber="292" pagination="450 - 468" refId="ref4609" refString="Anker, A. (2020 a) Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010, with description of a second eastern Pacific species (Malacostraca: Decapoda). Zootaxa, 4772 (3), 450 - 468. https: // doi. org / 10.11646 / zootaxa. 4772.3.2" type="journal article" year="2020">2020a</bibRefCitation>
), although they are located slightly more dorsally to the hepatic spines of other carideans, e.g., crangonids and palaemonids (cf. 
<bibRefCitation id="EFC0F7AEFFC5FFADFE16F8ECFDAEF8C7" author="Holthuis, L. B." box="[398,563,1831,1857]" pageId="5" pageNumber="292" refId="ref5717" refString="Holthuis, L. B. (1993) The Recent Genera of the Caridean and Stenopodidean Shrimps (Crustacea, Decapoda) with an Appendix on the Order Amphionidacea. Nationaal Natuurhistorisch Museum, Leiden, 328 pp. https: // doi. org / 10.5962 / bhl. title. 152891" type="book" year="1993">Holthuis 1993</bibRefCitation>
). In addition, the carapacial spines are in general better developed and more posteriorly located in 
<taxonomicName id="4C51F1DCFFC5FFADFE12F880FDA5F8E3" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[394,568,1867,1893]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFE12F880FDA5F8E3" box="[394,568,1867,1893]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
than in 
<taxonomicName id="4C51F1DCFFC5FFADFD0AF880FC80F8E3" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[658,797,1867,1893]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC5FFADFD0AF880FC80F8E3" box="[658,797,1867,1893]" italics="true" pageId="5" pageNumber="292">S. singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC5FFADFCBCF880FCFFF8E3" bold="true" box="[804,866,1867,1893]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A216EB36FFC5FFADFCBCF880FCFFF8E3" box="[804,866,1867,1893]" pageId="5" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The most parsimonious hypothesis may be that the dorsolateral spines are homologous between the new species and the five species of 
<taxonomicName id="4C51F1DCFFC5FFADFBC3F8A5FA94F80E" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[1115,1289,1902,1928]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFBC3F8A5FA94F80E" box="[1115,1289,1902,1928]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
, but it is not easy to fully assess the homology without phylogenetic analysis.
</paragraph>
<paragraph id="8BEE8A5FFFC5FFAEFF5FF87DFBD6FDBD" blockId="5.[151,1437,151,2036]" lastBlockId="6.[151,1437,151,572]" lastPageId="6" lastPageNumber="293" pageId="5" pageNumber="292">
The new species differs from all members of 
<taxonomicName id="4C51F1DCFFC5FFADFD5BF87DFCECF856" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[707,881,1974,2000]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFD5BF87DFCECF856" box="[707,881,1974,2000]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
in the lack of a mid-dorsal spine on the carapace, which is one of the main diagnostic characters of 
<taxonomicName id="4C51F1DCFFC5FFADFD23F812FCF4F875" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[699,873,2009,2035]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC5FFADFD23F812FCF4F875" box="[699,873,2009,2035]" italics="true" pageId="5" pageNumber="292">Triacanthoneus</emphasis>
</taxonomicName>
, although its position is somewhat variable among the species (
<bibRefCitation id="EFC0F7AEFFC6FFAEFEBCFF5CFE23FF37" author="Anker, A." box="[292,446,151,177]" pageId="6" pageNumber="293" pagination="47 - 63" refId="ref4377" refString="Anker, A. (2010 b) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010b</bibRefCitation>
, 
<bibRefCitation id="EFC0F7AEFFC6FFAEFE53FF5CFD8EFF37" author="Anker, A." box="[459,531,151,177]" pageId="6" pageNumber="293" pagination="450 - 468" refId="ref4609" refString="Anker, A. (2020 a) Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010, with description of a second eastern Pacific species (Malacostraca: Decapoda). Zootaxa, 4772 (3), 450 - 468. https: // doi. org / 10.11646 / zootaxa. 4772.3.2" type="journal article" year="2020">2020a</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC6FFAEFD87FF5CFD62FF37" author="Alvarez, F. &amp; Iliffe, T. M. &amp; Gonzalez, B. &amp; Villalobos, J. L." box="[543,767,151,177]" pageId="6" pageNumber="293" pagination="61 - 68" refId="ref4180" refString="Alvarez, F., Iliffe, T. M., Gonzalez, B. &amp; Villalobos, J. L. (2012) Triacanthoneus akumalensis, a new species of alpheid shrimp (Crustacea: Caridea: Alpheidae) from an anchialine cave in Quintana Roo, Mexico. Zootaxa, 3154 (1), 61 - 68. https: // doi. org / 10.11646 / zootaxa. 3154.1.5" type="journal article" year="2012">
Alvarez 
<emphasis id="B925564DFFC6FFAEFD19FF5CFD26FF37" box="[641,699,151,177]" italics="true" pageId="6" pageNumber="293">et al.</emphasis>
2012
</bibRefCitation>
, 
<bibRefCitation id="EFC0F7AEFFC6FFAEFC94FF5CFCD4FF37" author="Alvarez, F. &amp; Iliffe, T. M. &amp; Villalobos, J. L." box="[780,841,151,177]" pageId="6" pageNumber="293" pagination="88 - 94" refId="ref4255" refString="Alvarez, F., Iliffe, T. M. &amp; Villalobos, J. L. (2014) A new species of the alpheid shrimp genus Triacanthoneus Anker, 2010 (Crustacea: Alpheidae) from Belize. Zootaxa, 3768 (1), 88 - 94. https: // doi. org / 10.11646 / zootaxa. 3768.1.6" type="journal article" year="2014">2014</bibRefCitation>
). Other similarities between 
<taxonomicName id="4C51F1DCFFC6FFAEFB0AFF5CFABDFF37" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[1170,1312,151,177]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC6FFAEFB0AFF5CFABDFF37" box="[1170,1312,151,177]" italics="true" pageId="6" pageNumber="293">S. singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC6FFAEFAB1FF5CFAF7FF37" bold="true" box="[1321,1386,151,177]" pageId="6" pageNumber="293">
<taxonomicNameLabel id="A216EB36FFC6FFAEFAB1FF5CFAF7FF37" box="[1321,1386,151,177]" pageId="6" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C51F1DCFFC6FFAEFF0FFF71FED8FF52" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[151,325,186,212]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFF0FFF71FED8FF52" box="[151,325,186,212]" italics="true" pageId="6" pageNumber="293">Triacanthoneus</emphasis>
</taxonomicName>
are the presence of a subdistal spine on the ventral margin of the rostrum and the flexed ischio-meral articulation of the pereopods 3–5 (
<bibRefCitation id="EFC0F7AEFFC6FFAEFD87FF15FD24FF7E" author="Anker, A." box="[543,697,222,248]" pageId="6" pageNumber="293" pagination="450 - 468" refId="ref4609" refString="Anker, A. (2020 a) Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010, with description of a second eastern Pacific species (Malacostraca: Decapoda). Zootaxa, 4772 (3), 450 - 468. https: // doi. org / 10.11646 / zootaxa. 4772.3.2" type="journal article" year="2020">Anker 2020a</bibRefCitation>
). These features, however, are also present in some other species of 
<taxonomicName id="4C51F1DCFFC6FFAEFF2EFEC9FEB2FE9A" authorityName="Holthuis" authorityYear="1955" box="[182,303,258,284]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFF2EFEC9FEB2FE9A" box="[182,303,258,284]" italics="true" pageId="6" pageNumber="293">Salmoneus</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation id="EFC0F7AEFFC6FFAEFEEEFEC9FDF4FE9A" author="Anker, A. &amp; Marin, I. N." box="[374,617,258,284]" pageId="6" pageNumber="293" pagination="295 - 319" refId="ref4994" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea: Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. Raffles Bulletin of Zoology, 54 (2), 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC6FFAEFDEDFEC9FCDBFE9B" author="Komai, T. &amp; Yamada, Y. &amp; Yunokawa, Y." box="[629,838,258,285]" pageId="6" pageNumber="293" pagination="413 - 420" refId="ref5969" refString="Komai, T., Yamada, Y. &amp; Yunokawa, Y. (2015) A new cave-dwelling species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the Ryukyu Islands, southern Japan. Zootaxa, 4028 (3), 413 - 420. https: // doi. org / 10.11646 / zootaxa. 4028.3.6" type="journal article" year="2015">
Komai 
<emphasis id="B925564DFFC6FFAEFD50FEC8FC9CFE9A" box="[712,769,258,284]" italics="true" pageId="6" pageNumber="293">et al.</emphasis>
2015
</bibRefCitation>
). On the other side, in the general shape of the major cheliped, especially the cup-shaped, non-elongate carpus, 
<taxonomicName id="4C51F1DCFFC6FFAEFCB4FEEDFC27FEC6" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[812,954,294,320]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC6FFAEFCB4FEEDFC27FEC6" box="[812,954,294,320]" italics="true" pageId="6" pageNumber="293">S. singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC6FFAEFC5AFEEDFB9EFEC6" bold="true" box="[962,1027,294,320]" pageId="6" pageNumber="293">
<taxonomicNameLabel id="A216EB36FFC6FFAEFC5AFEEDFB9EFEC6" box="[962,1027,294,320]" pageId="6" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is closer to almost all other species currently placed in 
<taxonomicName id="4C51F1DCFFC6FFAEFEF7FE82FE75FEE5" authorityName="Holthuis" authorityYear="1955" box="[367,488,329,355]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFEF7FE82FE75FEE5" box="[367,488,329,355]" italics="true" pageId="6" pageNumber="293">Salmoneus</emphasis>
</taxonomicName>
, except for the cavernicolous 
<taxonomicName id="4C51F1DCFFC6FFAEFCAFFE81FBB6FEE5" authority="Fransen, 1991" authorityName="Fransen" authorityYear="1991" box="[823,1067,329,356]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="sketi">
<emphasis id="B925564DFFC6FFAEFCAFFE81FC1BFEE5" box="[823,902,329,356]" italics="true" pageId="6" pageNumber="293">S. sketi</emphasis>
<bibRefCitation id="EFC0F7AEFFC6FFAEFC15FE81FBB6FEE5" author="Fransen, C. H. J. M." box="[909,1067,329,356]" pageId="6" pageNumber="293" pagination="171 - 179" refId="ref5588" refString="Fransen, C. H. J. M. (1991) Salmoneus sketi, a new species of alpheid shrimp (Crustacea Decapoda: Caridea) from a submarine cave in the Adriatic. Zoologische Mededelingen, 65, 171 - 179." type="journal article" year="1991">Fransen, 1991</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C51F1DCFFC6FFAEFBF8FE81FEEDFE0E" authority="Komai, Yamada &amp; Yunokawa, 2015" authorityName="Komai, Yamada &amp; Yunokawa" authorityYear="2015" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="antricola">
<emphasis id="B925564DFFC6FFAEFBF8FE81FB7EFEE5" box="[1120,1251,329,356]" italics="true" pageId="6" pageNumber="293">S. antricola</emphasis>
<bibRefCitation id="EFC0F7AEFFC6FFAEFB71FE82FEEDFE0E" author="Komai, T. &amp; Yamada, Y. &amp; Yunokawa, Y." pageId="6" pageNumber="293" pagination="413 - 420" refId="ref5969" refString="Komai, T., Yamada, Y. &amp; Yunokawa, Y. (2015) A new cave-dwelling species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the Ryukyu Islands, southern Japan. Zootaxa, 4028 (3), 413 - 420. https: // doi. org / 10.11646 / zootaxa. 4028.3.6" type="journal article" year="2015">Komai, Yamada &amp; Yunokawa, 2015</bibRefCitation>
</taxonomicName>
, in which the carpus is vase-shaped or long and cylindrical, as in 
<taxonomicName id="4C51F1DCFFC6FFAEFBD8FEA6FB73FE01" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[1088,1262,365,391]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFBD8FEA6FB73FE01" box="[1088,1262,365,391]" italics="true" pageId="6" pageNumber="293">Triacanthoneus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC0F7AEFFC6FFAEFB64FEA6FA0AFE01" author="Fransen, C. H. J. M." box="[1276,1431,365,391]" pageId="6" pageNumber="293" pagination="171 - 179" refId="ref5588" refString="Fransen, C. H. J. M. (1991) Salmoneus sketi, a new species of alpheid shrimp (Crustacea Decapoda: Caridea) from a submarine cave in the Adriatic. Zoologische Mededelingen, 65, 171 - 179." type="journal article" year="1991">Fransen 1991</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC6FFAEFF0FFE5AFEF1FE2A" author="Komai, T. &amp; Yamada, Y. &amp; Yunokawa, Y." box="[151,364,401,428]" pageId="6" pageNumber="293" pagination="413 - 420" refId="ref5969" refString="Komai, T., Yamada, Y. &amp; Yunokawa, Y. (2015) A new cave-dwelling species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the Ryukyu Islands, southern Japan. Zootaxa, 4028 (3), 413 - 420. https: // doi. org / 10.11646 / zootaxa. 4028.3.6" type="journal article" year="2015">
Komai 
<emphasis id="B925564DFFC6FFAEFF74FE59FEBBFE2D" box="[236,294,401,427]" italics="true" pageId="6" pageNumber="293">et al.</emphasis>
2015
</bibRefCitation>
). It is important to note here that the development of the notch on the posterior margin of the telson is variable within both genera (
<bibRefCitation id="EFC0F7AEFFC6FFAEFDDEFE7EFD7DFE49" author="Anker, A." box="[582,736,437,463]" pageId="6" pageNumber="293" pagination="47 - 63" refId="ref4377" refString="Anker, A. (2010 b) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010b</bibRefCitation>
, 
<bibRefCitation id="EFC0F7AEFFC6FFAEFD75FE7EFCA9FE49" author="Anker, A." box="[749,820,437,463]" pageId="6" pageNumber="293" pagination="450 - 468" refId="ref4609" refString="Anker, A. (2020 a) Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010, with description of a second eastern Pacific species (Malacostraca: Decapoda). Zootaxa, 4772 (3), 450 - 468. https: // doi. org / 10.11646 / zootaxa. 4772.3.2" type="journal article" year="2020">2020a</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC6FFAEFCD8FE7EFB82FE49" author="Alvarez, F. &amp; Iliffe, T. M. &amp; Gonzalez, B. &amp; Villalobos, J. L." box="[832,1055,437,463]" pageId="6" pageNumber="293" pagination="61 - 68" refId="ref4180" refString="Alvarez, F., Iliffe, T. M., Gonzalez, B. &amp; Villalobos, J. L. (2012) Triacanthoneus akumalensis, a new species of alpheid shrimp (Crustacea: Caridea: Alpheidae) from an anchialine cave in Quintana Roo, Mexico. Zootaxa, 3154 (1), 61 - 68. https: // doi. org / 10.11646 / zootaxa. 3154.1.5" type="journal article" year="2012">
Alvarez 
<emphasis id="B925564DFFC6FFAEFC39FE7DFC46FE49" box="[929,987,437,463]" italics="true" pageId="6" pageNumber="293">et al.</emphasis>
2012
</bibRefCitation>
, 
<bibRefCitation id="EFC0F7AEFFC6FFAEFBB3FE7EFBF5FE49" author="Alvarez, F. &amp; Iliffe, T. M. &amp; Villalobos, J. L." box="[1067,1128,437,463]" pageId="6" pageNumber="293" pagination="88 - 94" refId="ref4255" refString="Alvarez, F., Iliffe, T. M. &amp; Villalobos, J. L. (2014) A new species of the alpheid shrimp genus Triacanthoneus Anker, 2010 (Crustacea: Alpheidae) from Belize. Zootaxa, 3768 (1), 88 - 94. https: // doi. org / 10.11646 / zootaxa. 3768.1.6" type="journal article" year="2014">2014</bibRefCitation>
); however, most species of 
<taxonomicName id="4C51F1DCFFC6FFAEFF0FFE12FED8FE75" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[151,325,473,499]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFF0FFE12FED8FE75" box="[151,325,473,499]" italics="true" pageId="6" pageNumber="293">Triacanthoneus</emphasis>
</taxonomicName>
do not have one, whereas most species of 
<taxonomicName id="4C51F1DCFFC6FFAEFC87FE12FC05FE75" authorityName="Holthuis" authorityYear="1955" box="[799,920,473,499]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFC87FE12FC05FE75" box="[799,920,473,499]" italics="true" pageId="6" pageNumber="293">Salmoneus</emphasis>
</taxonomicName>
present a fairly deep notch, as is the case of 
<taxonomicName id="4C51F1DCFFC6FFAEFA1FFE12FE9AFD91" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC6FFAEFA1FFE12FE9AFD91" italics="true" pageId="6" pageNumber="293">S. singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC6FFAEFE95FE36FED6FD91" bold="true" box="[269,331,509,535]" pageId="6" pageNumber="293">
<taxonomicNameLabel id="A216EB36FFC6FFAEFE95FE36FED6FD91" box="[269,331,509,535]" pageId="6" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136A96DAFFC6FFAEFEC2FE36FE2CFD91" box="[346,433,509,535]" captionStart="FIGURE 2" captionStartId="3.[151,250,1490,1514]" captionTargetBox="[160,1426,189,1455]" captionTargetId="figure-17@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus singularis n. sp., holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, left eye, anterolateral view; E, left antenna, ventral view (setae omitted); F, left pleopod 2, dorsal (anterior) view (setae omitted); G, left uropod, dorsal view (perpendicular to horizontal plane)." figureDoi="http://doi.org/10.5281/zenodo.4475317" httpUri="https://zenodo.org/record/4475317/files/figure.png" pageId="6" pageNumber="293">Fig. 2C</figureCitation>
). Based on the presently available evidence, it seems most reasonable to assign the present new species to 
<taxonomicName id="4C51F1DCFFC6FFAEFED9FDEAFE27FDBD" authorityName="Holthuis" authorityYear="1955" box="[321,442,545,571]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFED9FDEAFE27FDBD" box="[321,442,545,571]" italics="true" pageId="6" pageNumber="293">Salmoneus</emphasis>
</taxonomicName>
and not 
<taxonomicName id="4C51F1DCFFC6FFAEFD84FDEAFD57FDBD" baseAuthorityName="Anker" baseAuthorityYear="2020" box="[540,714,545,571]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC6FFAEFD84FDEAFD57FDBD" box="[540,714,545,571]" italics="true" pageId="6" pageNumber="293">Triacanthoneus</emphasis>
</taxonomicName>
as rediagnosed by 
<bibRefCitation id="EFC0F7AEFFC6FFAEFC07FDEAFBDAFDBD" author="Anker, A." box="[927,1095,545,571]" pageId="6" pageNumber="293" pagination="450 - 468" refId="ref4609" refString="Anker, A. (2020 a) Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010, with description of a second eastern Pacific species (Malacostraca: Decapoda). Zootaxa, 4772 (3), 450 - 468. https: // doi. org / 10.11646 / zootaxa. 4772.3.2" type="journal article" year="2020">Anker (2020a)</bibRefCitation>
.
</paragraph>
<caption id="DF2EDAD7FFC6FFAEFF0FF844FCFDF869" ID-DOI="http://doi.org/10.5281/zenodo.4475321" ID-Zenodo-Dep="4475321" httpUri="https://zenodo.org/record/4475321/files/figure.png" pageId="6" pageNumber="293" startId="6.[151,250,1935,1959]" targetBox="[167,1423,609,1897]" targetPageId="6">
<paragraph id="8BEE8A5FFFC6FFAEFF0FF844FCFDF869" blockId="6.[151,1437,1935,2031]" pageId="6" pageNumber="293">
<emphasis id="B925564DFFC6FFAEFF0FF844FE8EF821" bold="true" box="[151,275,1935,1959]" pageId="6" pageNumber="293">FIGURE 4.</emphasis>
<taxonomicName id="4C51F1DCFFC6FFAEFE81F844FE6FF821" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[281,498,1935,1959]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC6FFAEFE81F844FE6FF821" box="[281,498,1935,1959]" italics="true" pageId="6" pageNumber="293">Salmoneus singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC6FFAEFE61F844FDAFF821" bold="true" box="[505,562,1935,1959]" pageId="6" pageNumber="293">
<taxonomicNameLabel id="A216EB36FFC6FFAEFE61F844FDAFF821" box="[505,562,1935,1959]" pageId="6" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, non-ovigerous specimen (cl 5.4 mm), CBM-ZC 16164. A, right pereopod 1 (major cheliped), lateral view; B, same, coxa to carpus, lateral view in different angle; C, same, chela, extensor view; D, same, carpus and chela, flexor view; E, same, carpus and chela, mesial view.
</paragraph>
</caption>
<paragraph id="8BEE8A5FFFC7FFAFFF5FFF5CFC9CFE0F" blockId="7.[151,1437,151,393]" pageId="7" pageNumber="294">
<taxonomicName id="4C51F1DCFFC7FFAFFF5FFF5CFE24FF37" authority="Komai &amp; Maenosono &amp; Naruse, 2021" authorityName="Komai &amp; Maenosono &amp; Naruse" authorityYear="2021" box="[199,441,151,177]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="B925564DFFC7FFAFFF5FFF5CFE24FF37" box="[199,441,151,177]" italics="true" pageId="7" pageNumber="294">Salmoneus singularis</emphasis>
</taxonomicName>
<emphasis id="B925564DFFC7FFAFFE59FF5CFD9CFF37" bold="true" box="[449,513,151,177]" pageId="7" pageNumber="294">
<taxonomicNameLabel id="A216EB36FFC7FFAFFE59FF5CFD9CFF37" box="[449,513,151,177]" pageId="7" pageNumber="294" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
represents the eighth species of 
<taxonomicName id="4C51F1DCFFC7FFAFFCEDFF5CFC73FF37" authorityName="Holthuis" authorityYear="1955" box="[885,1006,151,177]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="B925564DFFC7FFAFFCEDFF5CFC73FF37" box="[885,1006,151,177]" italics="true" pageId="7" pageNumber="294">Salmoneus</emphasis>
</taxonomicName>
to be recorded from Japanese waters, the other seven species being 
<taxonomicName id="4C51F1DCFFC7FFAFFE75FF70FDEFFF53" authorityName="Komai, Yamada &amp; Yunokawa" authorityYear="2015" box="[493,626,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="antricola">
<emphasis id="B925564DFFC7FFAFFE75FF70FDEFFF53" box="[493,626,187,213]" italics="true" pageId="7" pageNumber="294">S. antricola</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC7FFAFFD19FF70FC77FF53" authority="Miyake &amp; Miya, 1966" authorityName="Miyake &amp; Miya" authorityYear="1966" box="[641,1002,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B925564DFFC7FFAFFD19FF70FD42FF53" box="[641,735,187,213]" italics="true" pageId="7" pageNumber="294">S. babai</emphasis>
<bibRefCitation id="EFC0F7AEFFC7FFAFFD7FFF70FC77FF53" author="Miyake, S. &amp; Miya, Y." box="[743,1002,187,213]" pageId="7" pageNumber="294" pagination="133 - 141" refId="ref6087" refString="Miyake, S. &amp; Miya, Y. (1966) On a new species and a new record of alpheid shrimps from Japan. Journal of the Faculty of Agriculture, Kyushu University, 14 (1), 133 - 141." type="journal article" year="1966">Miyake &amp; Miya, 1966</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC7FFAFFC6EFF70FB66FF53" authority="Komai, 2009" authorityName="Komai" authorityYear="2009" box="[1014,1275,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="B925564DFFC7FFAFFC6EFF70FBC1FF53" box="[1014,1116,187,213]" italics="true" pageId="7" pageNumber="294">S. brucei</emphasis>
<bibRefCitation id="EFC0F7AEFFC7FFAFFBFCFF70FB66FF53" author="Komai, T." box="[1124,1275,187,213]" pageId="7" pageNumber="294" pagination="869 - 880" refId="ref5771" refString="Komai, T. (2009) A new species of the alpheid shrimp genus Salmoneus (Decapoda, Caridea) from the Ryukyu Islands, Japan, associated with a callianassid ghost shrimp (Decapoda, Thalassinidea). Crustaceana, 82 (7), 869 - 880. https: // doi. org / 10.1163 / 156854009 X 427405" type="journal article" year="2009">Komai, 2009</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC7FFAFFA90FF70FA08FF53" authorityName="Miya" authorityYear="1972" box="[1288,1429,187,213]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B925564DFFC7FFAFFA90FF70FA08FF53" box="[1288,1429,187,213]" italics="true" pageId="7" pageNumber="294">S. gracilipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC7FFAFFF0FFF14FD8CFF7F" authority="Komai &amp; Anker, 2012" authorityName="Komai &amp; Anker" authorityYear="2012" box="[151,529,223,249]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="B925564DFFC7FFAFFF0FFF14FE9AFF7E" box="[151,263,223,249]" italics="true" pageId="7" pageNumber="294">S. pinguis</emphasis>
<bibRefCitation id="EFC0F7AEFFC7FFAFFE88FF14FD8CFF7F" author="Komai, T. &amp; Anker, A." box="[272,529,223,249]" pageId="7" pageNumber="294" pagination="60 - 68" refId="ref5834" refString="Komai, T. &amp; Anker, A. (2012) A distinctive new species of the alpheid genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the northwestern Pacific. Zootaxa, 3344 (1), 60 - 68. https: // doi. org / 10.11646 / zootaxa. 3344.1.4" type="journal article" year="2012">Komai &amp; Anker, 2012</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C51F1DCFFC7FFAFFD85FF14FC08FF7C" authority="(Coutiere, 1896)" baseAuthorityName="Coutiere" baseAuthorityYear="1896" box="[541,917,223,250]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B925564DFFC7FFAFFD85FF14FD4FFF7F" box="[541,722,223,249]" italics="true" pageId="7" pageNumber="294">S. serratidigitus</emphasis>
(
<bibRefCitation id="EFC0F7AEFFC7FFAFFD7AFF14FC10FF7C" author="Coutiere, H." box="[738,909,223,250]" pageId="7" pageNumber="294" pagination="380 - 386" refId="ref5215" refString="Coutiere, H. (1896) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1896">Coutière, 1896</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C51F1DCFFC7FFAFFC57FF14FA9AFF7F" authority="Banner, 1959" authorityName="Banner" authorityYear="1959" box="[975,1287,222,249]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="tricristatus">
<emphasis id="B925564DFFC7FFAFFC57FF14FBFAFF7E" box="[975,1127,223,249]" italics="true" pageId="7" pageNumber="294">S. tricristatus</emphasis>
<bibRefCitation id="EFC0F7AEFFC7FFAFFBE8FF14FA9AFF7F" author="Banner, A. H." box="[1136,1287,222,249]" pageId="7" pageNumber="294" pagination="130 - 155" refId="ref5112" refString="Banner, A. H. (1959) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part IV: various small collections from the central Pacific area, including supplementary notes on alpheids from Hawaii. Pacific Science, 13 (2), 130 - 155." type="journal article" year="1959">Banner, 1959</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFC0F7AEFFC7FFAFFA8FFF14FA0AFF7F" author="Miya, Y." box="[1303,1431,223,249]" pageId="7" pageNumber="294" pagination="23 - 101" refId="ref6037" refString="Miya, Y. (1972) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3 (1), 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC7FFAFFF0FFEC8FEA4FE9B" author="Hayashi, K. I." box="[151,313,258,285]" pageId="7" pageNumber="294" pagination="520 - 524" refId="ref5633" refString="Hayashi, K. I. (1995) Prawns, shrimps and lobsters from Japan (85). Family Alpheidae-genera Automate, and Salmoneus. Aquabiology, 101, 520 - 524. [In Japanese]" type="journal article" year="1995">Hayashi 1995</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC7FFAFFED1FEC8FE46FE9B" author="Komai, T." box="[329,475,259,285]" pageId="7" pageNumber="294" pagination="869 - 880" refId="ref5771" refString="Komai, T. (2009) A new species of the alpheid shrimp genus Salmoneus (Decapoda, Caridea) from the Ryukyu Islands, Japan, associated with a callianassid ghost shrimp (Decapoda, Thalassinidea). Crustaceana, 82 (7), 869 - 880. https: // doi. org / 10.1163 / 156854009 X 427405" type="journal article" year="2009">Komai 2009</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC7FFAFFE73FEC8FD72FE9B" author="Komai, T. &amp; Anker, A." box="[491,751,259,285]" pageId="7" pageNumber="294" pagination="60 - 68" refId="ref5834" refString="Komai, T. &amp; Anker, A. (2012) A distinctive new species of the alpheid genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the northwestern Pacific. Zootaxa, 3344 (1), 60 - 68. https: // doi. org / 10.11646 / zootaxa. 3344.1.4" type="journal article" year="2012">Komai &amp; Anker 2012</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC7FFAFFD66FEC8FC44FE9B" author="Komai, T. &amp; Yamada, Y. &amp; Yunokawa, Y." box="[766,985,258,285]" pageId="7" pageNumber="294" pagination="413 - 420" refId="ref5969" refString="Komai, T., Yamada, Y. &amp; Yunokawa, Y. (2015) A new cave-dwelling species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the Ryukyu Islands, southern Japan. Zootaxa, 4028 (3), 413 - 420. https: // doi. org / 10.11646 / zootaxa. 4028.3.6" type="journal article" year="2015">
Komai 
<emphasis id="B925564DFFC7FFAFFCCDFEC8FC0CFE9B" box="[853,913,259,285]" italics="true" pageId="7" pageNumber="294">et al.</emphasis>
2015
</bibRefCitation>
). All species have been recorded from the subtropical Ryukyu Islands, with only 
<taxonomicName id="4C51F1DCFFC7FFAFFD1AFEECFC8CFEC7" authorityName="Miya" authorityYear="1972" box="[642,785,295,321]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B925564DFFC7FFAFFD1AFEECFC8CFEC7" box="[642,785,295,321]" italics="true" pageId="7" pageNumber="294">S. gracilipes</emphasis>
</taxonomicName>
also occurring in warm temperate areas of the Japanese mainland (
<bibRefCitation id="EFC0F7AEFFC7FFAFFE96FE80FE10FEE3" author="Miya, Y." box="[270,397,331,357]" pageId="7" pageNumber="294" pagination="23 - 101" refId="ref6037" refString="Miya, Y. (1972) The Alpheidae (Crustacea, Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3 (1), 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
; 
<bibRefCitation id="EFC0F7AEFFC7FFAFFE00FE80FDFFFEE3" author="Anker, A. &amp; Al-Kandari, M. &amp; De Grave, S." box="[408,610,331,357]" pageId="7" pageNumber="294" pagination="77 - 92" refId="ref4731" refString="Anker, A., Al-Kandari, M. &amp; De Grave, S. (2020) On two species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from Kuwait, one of them new to science (Malacostraca: Decapoda: Caridea). Zootaxa, 4780 (1), 77 - 92. https: // doi. org / 10.11646 / zootaxa. 4780.1.3" type="journal article" year="2020">
Anker 
<emphasis id="B925564DFFC7FFAFFE7DFE80FD8AFEE3" box="[485,535,331,357]" italics="true" pageId="7" pageNumber="294">et al</emphasis>
. 2020
</bibRefCitation>
). In addition, 
<bibRefCitation id="EFC0F7AEFFC7FFAFFD64FE80FC25FEE3" author="Yoshigou, H." box="[764,952,331,357]" pageId="7" pageNumber="294" pagination="221 - 273" refId="ref6132" refString="Yoshigou, H. (2009) A collection of alpheid shrimps (Crustacea; Decapoda; Alpheidae) from the mouth of rivers and anchialine caves in Japan. Miscellaneous Reports of Hiwa Museum for Natural History, 50, 221 - 273. [in Japanese]" type="journal article" year="2009">Yoshigou (2009)</bibRefCitation>
recorded an unidentified species similar to 
<taxonomicName id="4C51F1DCFFC7FFAFFF0FFEA4FEBBFE0F" authorityName="De Grave" authorityYear="2004" box="[151,294,367,393]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="colinorum">
<emphasis id="B925564DFFC7FFAFFF0FFEA4FEBBFE0F" box="[151,294,367,393]" italics="true" pageId="7" pageNumber="294">S. colinorum</emphasis>
</taxonomicName>
, but its identity remains to be determined.
</paragraph>
</subSubSection>
</treatment>
</document>