232 lines
36 KiB
XML
232 lines
36 KiB
XML
<document id="DF0E52B90B2E2E117DE8F56ECAD4BBD9" ID-CLB-Dataset="9477" ID-DOI="10.11646/zootaxa.4885.4.8" ID-GBIF-Dataset="8cd9d74c-2aac-43b8-be04-c649f9300b1d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296987" ID-ZooBank="16AC9A8F-D326-42DC-91A2-029E2CF1A0ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606685483353" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03EC87C09F4BF30A20D8FF6DFC44FE50" docLanguage="en" docName="zootaxa.4885.4.8.pdf" docOrigin="Zootaxa 4885 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discheramocephalus vasilii in Darby 2013" docType="treatment" docVersion="8" lastPageNumber="594" masterDocId="FFD5FFB89F49F309204FFFB2FFDEFF81" masterDocTitle="Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae)" masterLastPageNumber="595" masterPageNumber="591" pageNumber="593" updateTime="1698846858831" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="142ECB968A20D4D02D947FD5D6802069">Notes on Malagasy Discheramocephalus Johnson (Coleoptera: Ptiliidae)</mods:title>
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<mods:namePart id="43E81FF7D22AF663EC42A34AD10FF056">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="C80190B4044B6CB72DF8BD97B9D853EB">Zootaxa</mods:title>
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<mods:date id="95C9E4ADC3E6568D6A7D3A97CAD6B614">2020</mods:date>
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<treatment id="03EC87C09F4BF30A20D8FF6DFC44FE50" ID-DOI="http://doi.org/10.5281/zenodo.4324251" ID-GBIF-Taxon="170599100" ID-Zenodo-Dep="4324251" LSID="urn:lsid:plazi:treatment:03EC87C09F4BF30A20D8FF6DFC44FE50" httpUri="http://treatment.plazi.org/id/03EC87C09F4BF30A20D8FF6DFC44FE50" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593">
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<subSubSection id="C35F655D9F4BF30B20D8FF6DFDE2FF78" box="[151,572,223,250]" pageId="2" pageNumber="593" type="nomenclature">
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<paragraph id="8BFA36D69F4BF30B20D8FF6DFDE2FF78" blockId="2.[151,572,223,286]" box="[151,572,223,250]" pageId="2" pageNumber="593">
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<heading id="D0B281BA9F4BF30B20D8FF6DFDE2FF78" bold="true" box="[151,572,223,250]" fontSize="11" level="1" pageId="2" pageNumber="593" reason="1">
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<taxonomicName id="4C454D559F4BF30B20D8FF6DFDE2FF78" ID-CoL="36SCJ" authority="Darby" authorityName="in Darby" authorityYear="2013" box="[151,572,223,250]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="vasilii">
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<emphasis id="B931EAC49F4BF30B20D8FF6DFDE2FF78" bold="true" box="[151,572,223,250]" pageId="2" pageNumber="593">
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<emphasis id="B931EAC49F4BF30B20D8FF6DFE3BFF7B" bold="true" box="[151,485,223,250]" italics="true" pageId="2" pageNumber="593">Discheramocephalus vasilii</emphasis>
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Darby
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C35F655D9F4BF30B20D8FEB1FEA9FE9C" box="[151,375,259,286]" pageId="2" pageNumber="593" type="description">
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<paragraph id="8BFA36D69F4BF30B20D8FEB1FEA9FE9C" blockId="2.[151,572,223,286]" box="[151,375,259,286]" pageId="2" pageNumber="593">
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(
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<figureCitation id="137E2A539F4BF30B20EFFEB1FEDEFE9C" box="[160,256,259,285]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">Fig. 1–3</figureCitation>
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,
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<figureCitation id="137E2A539F4BF30B2143FEB6FEC2FE9F" box="[268,284,260,286]" captionStart="FIGURES 7–9" captionStartId="2.[151,264,1967,1991]" captionTargetBox="[166,1421,1562,1926]" captionTargetId="figure-566@2.[151,1436,1528,1943]" captionTargetPageId="2" captionText="FIGURES 7–9. Abdominal internal ‘cavities’ of segment VIII. Discheramocephalus vasilii Darby, male (7); D. bisulcatus Darby, male (8) and female (9)." figureDoi="http://doi.org/10.5281/zenodo.4296991" httpUri="https://zenodo.org/record/4296991/files/figure.png" pageId="2" pageNumber="593">7</figureCitation>
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,
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<figureCitation id="137E2A539F4BF30B2167FEB1FEB1FE9C" box="[296,367,259,285]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="2" pageNumber="593">10–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F655D9F4BF30B20D8FEFAFC91FEE1" box="[151,847,327,352]" pageId="2" pageNumber="593" type="reference_group">
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<paragraph id="8BFA36D69F4BF30B20D8FEFAFC91FEE1" blockId="2.[151,847,327,352]" box="[151,847,327,352]" pageId="2" pageNumber="593">
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<treatmentCitationGroup id="AB5511F89F4BF30B20D8FEFAFC91FEE1" box="[151,847,327,352]" pageId="2" pageNumber="593">
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<treatmentCitation id="0AE410C79F4BF30B20D8FEFAFDB4FEE1" ID-DOI="http://doi.org/10.5281/zenodo.6154589" ID-Zenodo-Dep="6154589" LSID="urn:lsid:zoobank.org:act:53653D9C-FDD2-4A00-8FC8-FFB143CD0544" author="Darby, M." box="[151,618,328,352]" httpUri="http://treatment.plazi.org/id/187B87854057091A3BE6FA14FD22F4D5" page="241" pageId="2" pageNumber="593" year="2013">
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<taxonomicName id="4C454D559F4BF30B20D8FEFAFDB4FEE1" ID-CoL="36SCJ" authority="Darby, 2013: 241" authorityName="in Darby" authorityPageNumber="241" authorityYear="2013" box="[151,618,328,352]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="vasilii">
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<emphasis id="B931EAC49F4BF30B20D8FEFAFE6FFEE1" box="[151,433,328,352]" italics="true" pageId="2" pageNumber="593">Discheramocephalus vasilii</emphasis>
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<bibRefCitation id="EFD44B279F4BF30B21F7FEFAFDB4FEE1" author="Darby, M." box="[440,618,328,352]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby, 2013: 241</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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,
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<figureCitation id="137E2A539F4BF30B223AFEFAFD6FFEE1" box="[629,689,328,352]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">Fig. 2</figureCitation>
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, 20, 22–25, 28.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F655D9F4BF30B20D8FE3FFA8DFB8D" pageId="2" pageNumber="593" type="materials_examined">
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<paragraph id="8BFA36D69F4BF30B20D8FE3FFCB3FE4D" blockId="2.[151,1437,397,1504]" pageId="2" pageNumber="593">
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<emphasis id="B931EAC49F4BF30B20D8FE3FFE80FE26" bold="true" box="[151,350,397,423]" pageId="2" pageNumber="593">Material studied</emphasis>
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:
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<materialsCitation id="3B2D3C8B9F4BF30B2122FE3CFCB4FE4D" ID-GBIF-Occurrence="2985804301" collectingDate="2019-12-04" collectionCode="MNHW" collectorName="P. Jaloszynski" country="Madagascar" elevation="950" latitude="-21.26" location="Ranomafana National Park" longLatPrecision="758" longitude="47.43" pageId="2" pageNumber="593" specimenCount="1">
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1 ♁,
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||
<collectingCountry id="F35276469F4BF30B21E9FE3CFDF1FE29" box="[422,559,398,424]" name="Madagascar" pageId="2" pageNumber="593">Madagascar</collectingCountry>
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,
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<location id="8E9A600D9F4BF30B2274FE3CFCB0FE29" LSID="urn:lsid:plazi:treatment:03EC87C09F4BF30A20D8FF6DFC44FE50:8E9A600D9F4BF30B2274FE3CFCB0FE29" box="[571,878,398,424]" country="Madagascar" latitude="-21.26" longLatPrecision="758" longitude="47.43" name="Ranomafana National Park" pageId="2" pageNumber="593">Ranomafana National Park</location>
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,
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<geoCoordinate id="EE7150119F4BF30B2335FE3CFC15FE29" box="[890,971,398,424]" degrees="21.26" direction="south" orientation="latitude" pageId="2" pageNumber="593" precision="555" value="-21.26">21.26S</geoCoordinate>
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,
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<geoCoordinate id="EE7150119F4BF30B2398FE3CFBF4FE29" box="[983,1066,398,424]" degrees="47.43" direction="east" orientation="longitude" pageId="2" pageNumber="593" precision="555" value="47.43">47.43E</geoCoordinate>
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,
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<quantity id="4CBD9B339F4BF30B2479FE3CFB1BFE29" box="[1078,1221,398,424]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="2" pageNumber="593" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
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<elevation id="0068D1E59F4BF30B2479FE3CFB1BFE29" box="[1078,1221,398,424]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="2" pageNumber="593" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</elevation>
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</quantity>
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,
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<date id="FFFB10169F4BF30B249EFE3CFA8CFE29" box="[1233,1362,398,424]" pageId="2" pageNumber="593" value="2019-12-04">
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<collectingDate id="EFBFE9FE9F4BF30B249EFE3CFA8CFE29" box="[1233,1362,398,424]" pageId="2" pageNumber="593" value="2019-12-04">12.04.2019</collectingDate>
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</date>
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, sifted from leaf litter in secondary forest, leg.
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<collectorName id="26B053009F4BF30B221AFE00FD2EFE4D" box="[597,752,434,460]" pageId="2" pageNumber="593">P. Jałoszyński</collectorName>
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(
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<collectionCode id="ED54AE139F4BF30B22B0FE00FCBFFE4D" box="[767,865,434,460]" pageId="2" pageNumber="593">MNHW</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BFA36D69F4BF30B2088FE64FD4CFD6D" blockId="2.[151,1437,397,1504]" pageId="2" pageNumber="593">
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<emphasis id="B931EAC49F4BF30B2088FE64FE69FE6E" bold="true" box="[199,439,469,496]" pageId="2" pageNumber="593">Emended diagnosis.</emphasis>
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Head with arcuate, anteriorly convex frontal groove accompanied at each side by elongate, oblique impression and with a pair of large shallow impressions each near mesal margin of supraantennal region; pronotum with six pairs of longitudinal sulci, of which only the lateral pair reaches anterior pronotal margin, each of four median sulci strongly broadened behind middle, pronotum with transverse, oval antebasal impression between the two lateralmost sulci; all antennomeres less than 3 times as long as broad; ‘cavities’ of abdominal segment VIII in male conspicuously large, each about as broad as 1/4 width of segment; posterior margin of sternite IX in male with broad median area devoid of denticles; aedeagus slender, in dorsal view parallel-sided between base and distal 1/4, in lateral view weakly sinuate.
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</paragraph>
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<paragraph id="8BFA36D69F4BF30B2088FD47FE82FC21" blockId="2.[151,1437,397,1504]" pageId="2" pageNumber="593">
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<emphasis id="B931EAC49F4BF30B2088FD47FDCDFC8E" bold="true" box="[199,531,757,783]" pageId="2" pageNumber="593">Supplementary description.</emphasis>
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Head in male with paired impressions, these internally with smooth surface, not filled with setae (
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<figureCitation id="137E2A539F4BF30B2112FCA8FE61FCB5" box="[349,447,794,820]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">Fig. 1–2</figureCitation>
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); antennomeres 3–9 elongate but each less than 3 times as long as broad (
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<figureCitation id="137E2A539F4BF30B2546FCA8FA93FCB5" box="[1289,1357,794,820]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">Fig. 3</figureCitation>
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); male abdominal segment VIII (
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<figureCitation id="137E2A539F4BF30B21F8FC8CFE24FCD9" box="[439,506,830,856]" captionStart="FIGURES 7–9" captionStartId="2.[151,264,1967,1991]" captionTargetBox="[166,1421,1562,1926]" captionTargetId="figure-566@2.[151,1436,1528,1943]" captionTargetPageId="2" captionText="FIGURES 7–9. Abdominal internal ‘cavities’ of segment VIII. Discheramocephalus vasilii Darby, male (7); D. bisulcatus Darby, male (8) and female (9)." figureDoi="http://doi.org/10.5281/zenodo.4296991" httpUri="https://zenodo.org/record/4296991/files/figure.png" pageId="2" pageNumber="593">Fig. 7</figureCitation>
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) with a pair of large internal ‘cavities’, each nearly as broad as 1/4 of width of segment; posterior margin of sternite IX in male (
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<figureCitation id="137E2A539F4BF30B22EFFCD0FD2CFCFD" box="[672,754,866,892]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="2" pageNumber="593">Fig. 10</figureCitation>
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) with 4–5 sharp denticles on each side, median area arcuate, lacking denticles.
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</paragraph>
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<paragraph id="8BFA36D69F4BF30B2088FC18FA8DFB8D" blockId="2.[151,1437,397,1504]" pageId="2" pageNumber="593">
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Aedeagus (
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<figureCitation id="137E2A539F4BF30B210BFC18FE1AFC45" box="[324,452,938,964]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="2" pageNumber="593">Fig. 11– 12</figureCitation>
|
||
) slender and conspicuously large, ~1/3 as long as body, in dorsal view (
|
||
<figureCitation id="137E2A539F4BF30B24AEFC18FAF1FC45" box="[1249,1327,938,964]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="2" pageNumber="593">Fig. 11</figureCitation>
|
||
) parallelsided from base to distal 1/4, in lateral view (
|
||
<figureCitation id="137E2A539F4BF30B22C2FC7CFD00FC69" box="[653,734,974,1000]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1643,1667]" captionTargetBox="[291,1296,481,1621]" captionTargetId="figure-161@3.[284,1304,474,1628]" captionTargetPageId="3" captionText="FIGURES 10–18. Discheramocephalus vasilii Darby, male (10–12), and D. bisulcatus Darby, male (13, 15–17) and female (14, 18). Posterior margin of abdominal sternite IX (10, 13–14); aedeagus in dorsal (11, 15) and lateral (12, 17) views; apex of aedeagus in dorsal view (16); spermatheca (18)." figureDoi="http://doi.org/10.5281/zenodo.4296993" httpUri="https://zenodo.org/record/4296993/files/figure.png" pageId="2" pageNumber="593">Fig. 12</figureCitation>
|
||
) weakly sinuate; endophallic structures indistinct, with numerous transverse folds of internal sac barely discernible under light microscope (better so with phase contrast).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F655D9F4BF30A2088FBA4FC44FE50" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593" type="discussion">
|
||
<paragraph id="8BFA36D69F4BF30B2088FBA4FBF3FA61" blockId="2.[151,1437,397,1504]" pageId="2" pageNumber="593">
|
||
<emphasis id="B931EAC49F4BF30B2088FBA4FEE5FBB1" bold="true" box="[199,315,1046,1072]" pageId="2" pageNumber="593">Remarks.</emphasis>
|
||
Details of the aedeagus and abdominal structures were not described previously for this species. As a part of diagnosis, “large, shallow, setae bearing pits on the head” were mentioned by
|
||
<bibRefCitation id="EFD44B279F4BF30B24C1FB88FAF7FBD5" author="Darby, M." box="[1166,1321,1082,1108]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby (2013)</bibRefCitation>
|
||
. His SEM micrographs of the pronotum and head (
|
||
<bibRefCitation id="EFD44B279F4BF30B2218FBECFD3FFBF9" author="Darby, M." box="[599,737,1118,1144]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby 2013</bibRefCitation>
|
||
: Fig. 20) agree with the characters seen in the specimen examined during the present study. The antenna (
|
||
<bibRefCitation id="EFD44B279F4BF30B22C4FB30FCC6FB1D" author="Darby, M." box="[651,792,1154,1180]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby 2013</bibRefCitation>
|
||
: Fig. 28) generally agrees with that illustrated here (
|
||
<figureCitation id="137E2A539F4BF30B2520FB30FF78FB41" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">Fig. 3</figureCitation>
|
||
), although with slight differences in the shape of some flagellomeres. Similar differences between the antennae of
|
||
<taxonomicName id="4C454D559F4BF30B20D8FB79FEF2FB65" authorityName="Darby" authorityYear="2013" box="[151,300,1226,1252]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="bisulcatus">
|
||
<emphasis id="B931EAC49F4BF30B20D8FB79FEF2FB65" box="[151,300,1226,1252]" italics="true" pageId="2" pageNumber="593">D. bisulcatus</emphasis>
|
||
</taxonomicName>
|
||
figured by
|
||
<bibRefCitation id="EFD44B279F4BF30B21E0FB78FD9CFB65" author="Darby, M." box="[431,578,1226,1252]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby (2013</bibRefCitation>
|
||
: Fig. 27) and that in
|
||
<figureCitation id="137E2A539F4BF30B2362FB78FCB0FB65" box="[813,878,1226,1252]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">
|
||
Fig.
|
||
<quantity id="4CBD9B339F4BF30B232FFB78FCB0FB65" box="[864,878,1226,1252]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="593" unit="in" value="6.0">6</quantity>
|
||
</figureCitation>
|
||
in the present paper can be seen, and they can be attributed to different study methods (light microscopy of a specimen in liquid medium and hand drawing, vs. SEM of dried specimens). A close-up of the head of
|
||
<taxonomicName id="4C454D559F4BF30B22FCFAA1FC09FAAD" authority="in Darby (2013" authorityName="in Darby" authorityYear="2013" box="[691,983,1298,1324]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="vasilii">
|
||
<emphasis id="B931EAC49F4BF30B22FCFAA1FCC5FAAD" box="[691,795,1298,1324]" italics="true" pageId="2" pageNumber="593">D. vasilii</emphasis>
|
||
in
|
||
<bibRefCitation id="EFD44B279F4BF30B230CFAA0FC09FAAD" author="Darby, M." box="[835,983,1298,1324]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby (2013</bibRefCitation>
|
||
</taxonomicName>
|
||
: Fig. 23–24) shows paired impressions, similar to those found during the present study, except that there is another, small median impression in front of the transverse arcuate sulcus. The impressions do not seem to bear any setae, but appear as filled with dirt. The same paired impressions in the specimen examined and illustrated in the present study (
|
||
<figureCitation id="137E2A539F4BF30B246DFACCFBBAFA19" box="[1058,1124,1406,1432]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1887,1911]" captionTargetBox="[151,1436,381,1864]" captionTargetId="figure-82@1.[151,1436,381,1864]" captionTargetPageId="1" captionText="FIGURES 1–6. Discheramocephalus vasilii Darby, male (1–3), and D. bisulcatus Darby, male (4–6). Dorsal habitus (1, 4); head, pronotum and elytral base in dorsal view (2, 5); antenna in dorsal view (3, 6)." figureDoi="http://doi.org/10.5281/zenodo.4296989" httpUri="https://zenodo.org/record/4296989/files/figure.png" pageId="2" pageNumber="593">Fig. 2</figureCitation>
|
||
) are smooth and empty, and the median impression is missing. The presence and absence of the latter may reflect variability within species, or sexual dimorphism (the specimen illustrated by
|
||
<bibRefCitation id="EFD44B279F4BF30B22FEFA74FC97FA61" author="Darby, M." box="[689,841,1478,1504]" pageId="2" pageNumber="593" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby (2013)</bibRefCitation>
|
||
is of unknown sex).
|
||
</paragraph>
|
||
<caption id="DF3A665E9F4BF30B20D8F81DFE09F86A" ID-DOI="http://doi.org/10.5281/zenodo.4296991" ID-Zenodo-Dep="4296991" httpUri="https://zenodo.org/record/4296991/files/figure.png" pageId="2" pageNumber="593" startId="2.[151,264,1967,1991]" targetBox="[166,1421,1562,1926]" targetPageId="2">
|
||
<paragraph id="8BFA36D69F4BF30B20D8F81DFE09F86A" blockId="2.[151,1436,1967,2027]" pageId="2" pageNumber="593">
|
||
<emphasis id="B931EAC49F4BF30B20D8F81DFEE0F846" bold="true" box="[151,318,1967,1991]" pageId="2" pageNumber="593">FIGURES 7–9.</emphasis>
|
||
Abdominal internal ‘cavities’ of segment VIII.
|
||
<taxonomicName id="4C454D559F4BF30B2378F81DFB43F846" authority="Darby" authorityName="in Darby" authorityYear="2013" box="[823,1181,1967,1991]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="vasilii">
|
||
<emphasis id="B931EAC49F4BF30B2378F81DFB8AF846" box="[823,1108,1967,1991]" italics="true" pageId="2" pageNumber="593">Discheramocephalus vasilii</emphasis>
|
||
Darby
|
||
</taxonomicName>
|
||
, male (7);
|
||
<taxonomicName id="4C454D559F4BF30B255CF802FF09F86A" authority="Darby" authorityName="Darby" authorityYear="2013" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="bisulcatus">
|
||
<emphasis id="B931EAC49F4BF30B255CF802FA42F846" box="[1299,1436,1967,1991]" italics="true" pageId="2" pageNumber="593">D. bisulcatus</emphasis>
|
||
Darby
|
||
</taxonomicName>
|
||
, male (8) and female (9).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFA36D69F4AF30A2088FF25FC44FE50" blockId="3.[151,1437,151,465]" pageId="3" pageNumber="594">
|
||
<taxonomicName id="4C454D559F4AF30A2088FF25FE20FF30" authorityName="in Darby" authorityYear="2013" box="[199,510,151,177]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="vasilii">
|
||
<emphasis id="B931EAC49F4AF30A2088FF25FE20FF30" box="[199,510,151,177]" italics="true" pageId="3" pageNumber="594">Discheramocephalus vasilii</emphasis>
|
||
</taxonomicName>
|
||
, as noted by
|
||
<bibRefCitation id="EFD44B279F4AF30A22DEFF25FCF5FF30" author="Darby, M." box="[657,811,151,177]" pageId="3" pageNumber="594" pagination="239 - 266" refId="ref2429" refString="Darby, M. (2013) Studies of Madagascan Ptiliidae (Coleoptera) 1: the tribe Discheramocephalini including eighteen new species. Zootaxa, 3718 (3), 239 - 266. https: // doi. org / 10.11646 / zootaxa. 3718.3.2" type="journal article" year="2013">Darby (2013)</bibRefCitation>
|
||
, shares the same number of pronotal sulci with
|
||
<emphasis id="B931EAC49F4AF30A2573FF25FF3CFF54" italics="true" pageId="3" pageNumber="594">D. elisa- bethae</emphasis>
|
||
<bibRefCitation id="EFD44B279F4AF30A20A6FF09FE1CFF54" author="Grebennikov, V. V." box="[233,450,187,213]" pageId="3" pageNumber="594" pagination="313 - 328" refId="ref2516" refString="Grebennikov, V. V. (2008) How small you can go: Factors limiting body miniaturization in winged insects with a review of the pantropical genus Discheramocephalus and description of six new species of the smallest beetles (Pterygota: Coleoptera: Ptiliidae). European Journal of Entomology, 105, 313 - 328. https: // doi. org / 10.14411 / eje. 2008.039" type="journal article" year="2008">Grebennikov, 2008</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="4C454D559F4AF30A21B3FF09FC8FFF54" authority="Grebennikov, 2008" authorityName="Grebennikov" authorityYear="2008" box="[508,849,187,213]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="mikaeli">
|
||
<emphasis id="B931EAC49F4AF30A21B3FF09FDAEFF54" box="[508,624,187,213]" italics="true" pageId="3" pageNumber="594">D. mikaeli</emphasis>
|
||
<bibRefCitation id="EFD44B279F4AF30A2238FF09FC8FFF54" author="Grebennikov, V. V." box="[631,849,187,213]" pageId="3" pageNumber="594" pagination="313 - 328" refId="ref2516" refString="Grebennikov, V. V. (2008) How small you can go: Factors limiting body miniaturization in winged insects with a review of the pantropical genus Discheramocephalus and description of six new species of the smallest beetles (Pterygota: Coleoptera: Ptiliidae). European Journal of Entomology, 105, 313 - 328. https: // doi. org / 10.14411 / eje. 2008.039" type="journal article" year="2008">Grebennikov, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C454D559F4AF30A2314FF09FB10FF54" box="[859,1230,187,213]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="elisabethae">
|
||
<emphasis id="B931EAC49F4AF30A2314FF09FB10FF54" box="[859,1230,187,213]" italics="true" pageId="3" pageNumber="594">Discheramocephalus elisabethae</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<collectingCountry id="F35276469F4AF30A2494FF08FA88FF54" box="[1243,1366,186,213]" name="Cameroon" pageId="3" pageNumber="594">Cameroon</collectingCountry>
|
||
] lacks the arcuate transverse frontal groove, its postocular transverse groove bears submedian longitudinal carinulae flanking a narrow smooth area, and the four median pronotal sulci are indistinctly broadening posterad. Each of these characters is different in
|
||
<taxonomicName id="4C454D559F4AF30A21FBFE95FDC5FEC0" authorityName="in Darby" authorityYear="2013" box="[436,539,295,321]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="vasilii">
|
||
<emphasis id="B931EAC49F4AF30A21FBFE95FDC5FEC0" box="[436,539,295,321]" italics="true" pageId="3" pageNumber="594">D. vasilii</emphasis>
|
||
</taxonomicName>
|
||
, which has a transverse frontal groove on the head, the smooth median region of the postocular groove not demarcated by longitudinal carinulae, and the pronotal sulci strongly, almost abruptly broadening posterad.
|
||
<taxonomicName id="4C454D559F4AF30A21C8FEDDFD15FE08" authorityName="Grebennikov" authorityYear="2008" box="[391,715,367,393]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="mikaeli">
|
||
<emphasis id="B931EAC49F4AF30A21C8FEDDFD15FE08" box="[391,715,367,393]" italics="true" pageId="3" pageNumber="594">Discheramocephalus mikaeli</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<collectingCountry id="F35276469F4AF30A2297FEDCFC9DFE08" box="[728,835,366,393]" name="Tanzania" pageId="3" pageNumber="594">Tanzania</collectingCountry>
|
||
] differs from
|
||
<taxonomicName id="4C454D559F4AF30A2396FEDDFBE0FE08" authorityName="in Darby" authorityYear="2013" box="[985,1086,367,393]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="vasilii">
|
||
<emphasis id="B931EAC49F4AF30A2396FEDDFBE0FE08" box="[985,1086,367,393]" italics="true" pageId="3" pageNumber="594">D. vasilii</emphasis>
|
||
</taxonomicName>
|
||
in the same cephalic characters as
|
||
<taxonomicName id="4C454D559F4AF30A20FAFE21FE89FE2C" box="[181,343,403,429]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="elisabethae">
|
||
<emphasis id="B931EAC49F4AF30A20FAFE21FE89FE2C" box="[181,343,403,429]" italics="true" pageId="3" pageNumber="594">D. elisabethae</emphasis>
|
||
</taxonomicName>
|
||
, and additionally in the median pair of pronotal sulci much broader and reaching anterior pronotal margin, vs. narrow and not reaching the anterior margin in
|
||
<taxonomicName id="4C454D559F4AF30A2362FE05FC4DFE50" authorityName="in Darby" authorityYear="2013" box="[813,915,439,465]" class="Insecta" family="Ptiliidae" genus="Discheramocephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="vasilii">
|
||
<emphasis id="B931EAC49F4AF30A2362FE05FC4DFE50" box="[813,915,439,465]" italics="true" pageId="3" pageNumber="594">D. vasilii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |