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<document id="77F23CD55BCDEDC9F1BA43EF5CCBA31F" ID-CLB-Dataset="46129" ID-DOI="10.11646/zootaxa.3784.1.2" ID-GBIF-Dataset="e2720913-7963-4303-bff2-e11872684c54" ID-ISSN="1175-5326" ID-Zenodo-Dep="230341" ID-ZooBank="1DF6FF6B-D761-46C4-B3CE-F94DA3DE156E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460402476791" checkinUser="plazi" docAuthor="Nguyen, Sang Ngoc, Yang, Jun-Xiao, Le, Thanh-Ngan Thi, Nguyen, Luan Thanh, Orlov, Nikolai L., Hoang, Chung Van, Nguyen, Truong Quang, Jin, Jie-Qiong, Rao, Ding-Qi, Hoang, Thao Ngoc, Che, Jing, Murphy, Robert W. &amp; Zhang, Ya-Ping" docDate="2014" docId="03EC2F3C0222E122FF53FD00FF0EFD45" docLanguage="en" docName="zt03784p066.pdf" docOrigin="Zootaxa 3784 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus puhuensis Nguyen, Yang, Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang, Che, Murphy &amp; Zhang, 2014, sp. nov." docType="treatment" docVersion="9" masterDocId="FFD55744022BE12CFFC4FF82FFEBFFAA" masterDocTitle="DNA barcoding of Vietnamese bent-toed geckos (Squamata: Gekkonidae: Cyrtodactylus) and the description of a new species" updateTime="1698299380338" updateUser="ExternalLinkService">
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<mods:title id="188D9DABEDE66D3A03902C48661F2CB5">DNA barcoding of Vietnamese bent-toed geckos (Squamata: Gekkonidae: Cyrtodactylus) and the description of a new species</mods:title>
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<mods:namePart id="162379D19E072EFD405E6BF3E60DE4D1">Nguyen, Sang Ngoc</mods:namePart>
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<mods:namePart id="5802F9037C9614B8005DA1456CBD4361">Yang, Jun-Xiao</mods:namePart>
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<mods:namePart id="79F1D8CACDA26B36AEE3FC4C64349268">Le, Thanh-Ngan Thi</mods:namePart>
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<mods:namePart id="11CBCB30B03E76C53608C6E1410D06E9">Nguyen, Luan Thanh</mods:namePart>
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<mods:namePart id="6FB00B0C2780AC2090AD422D0FE29C73">Hoang, Chung Van</mods:namePart>
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<mods:namePart id="093FCD505DDBF003242D91C14A0C0796">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="27F8A1F72E8499E847BE63034085316B">Jin, Jie-Qiong</mods:namePart>
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<mods:namePart id="42208E316C17F5E677A588249A9F3E15">Rao, Ding-Qi</mods:namePart>
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<mods:namePart id="00ADCE1E1433BDE055426A4BDDF84BC3">Hoang, Thao Ngoc</mods:namePart>
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<mods:namePart id="DCDF4B61F7A06F6A444F17462F4A04B4">Che, Jing</mods:namePart>
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<mods:namePart id="E93F266ECF5E0A3CE3AB5959E9FAE8A7">Zhang, Ya-Ping</mods:namePart>
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<mods:date id="A8BD039EA21E8C529025D48AF30D626E">2014</mods:date>
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<treatment id="03EC2F3C0222E122FF53FD00FF0EFD45" ID-DOI="http://doi.org/10.5281/zenodo.6143217" ID-GBIF-Taxon="119420147" ID-Zenodo-Dep="6143217" LSID="urn:lsid:plazi:treatment:03EC2F3C0222E122FF53FD00FF0EFD45" httpUri="http://treatment.plazi.org/id/03EC2F3C0222E122FF53FD00FF0EFD45" lastPageId="14" pageId="9">
<subSubSection id="C35FCDA10222E125FF53FD00FEFAFD48" pageId="9" type="nomenclature">
<paragraph id="8BFA9E2A0222E125FF53FD00FDCFFD36" blockId="9.[151,548,642,738]" box="[151,548,642,668]" pageId="9">
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<taxonomicName id="4C45E5A90222E125FF53FD00FE2BFD36" box="[151,448,642,668]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380222E125FF53FD00FE2BFD36" bold="true" box="[151,448,642,668]" italics="true" pageId="9">Cyrtodactylus puhuensis</emphasis>
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<taxonomicNameLabel id="A202FF430222E125FE03FD00FDCFFD36" box="[455,548,642,668]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
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Pù Hu bent-toed
<taxonomicName id="4C45E5A90222E125FE91FD27FE73FD14" box="[341,408,677,702]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="9" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
(
<figureCitation id="137E82AF0222E125FF5BFD4BFEE2FD48" box="[159,265,713,738]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="8.[151,250,1878,1900]" captionStartId-1="11.[151,250,888,910]" captionStartId-2="11.[151,250,1862,1884]" captionStartId-3="12.[151,250,1085,1107]" captionTargetBox-0="[178,1387,223,1835]" captionTargetBox-1="[151,1436,193,866]" captionTargetBox-2="[155,1427,963,1835]" captionTargetBox-3="[153,1431,207,1061]" captionTargetId-0="figure@8.[169,1405,193,1840]" captionTargetId-1="figure@11.[151,1436,193,867]" captionTargetId-2="figure@11.[151,1436,960,1841]" captionTargetId-3="figure@12.[151,1436,193,1064]" captionTargetPageId-0="8" captionTargetPageId-1="11" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 3. Bayesian inference tree for species of Cyrtodactylus from Vietnam and adjacent areas. Symbols at nodes show levels of support based on Bayesian posterior probabilities (BPP), and bootstrap (BS) proportions from maximum likelihood (ML) and maximum parsimony (MP), respectively. Asterisks indicate nodes supported 100 % by BI, ML, and MP; a plus sign indicates support of 95 99 % by BI and 70 99 % by ML and MP; and a hyphen an un-supported node. Photos of geckos not proportional to size in life." captionText-1="FIGURE 4. Holotype of Cyrtodactylus puhuensis sp. nov. in life." captionText-2="FIGURE 5. Holotype of Cyrtodactylus puhuensis sp. nov. Precloacal-femoral area (A), ventral surface of tail (B), and lateral (C), dorsal (D), and ventral (E) views of head." captionText-3="FIGURE 6. Holotype of Cyrtodactylus puhuensis sp. nov. dorsal (A) and ventral views (B) in preservative and ventral surface of right hand (C) and foot (D)." httpUri-0="https://zenodo.org/record/230344/files/figure.png" httpUri-1="https://zenodo.org/record/230345/files/figure.png" httpUri-2="https://zenodo.org/record/230346/files/figure.png" httpUri-3="https://zenodo.org/record/230347/files/figure.png" pageId="9">Figs. 36</figureCitation>
)
</paragraph>
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<paragraph id="8BFA9E2A0222E125FF53FC93FAD6FCE4" blockId="9.[151,1436,785,2034]" pageId="9">
<typeStatus id="54FE20880222E125FF53FC93FEEFFC80" box="[151,260,785,810]" pageId="9" type="holotype">
<emphasis id="B93142380222E125FF53FC93FEEFFC80" bold="true" box="[151,260,785,810]" pageId="9">Holotype</emphasis>
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.
<collectionCode id="ED5406EF0222E125FED6FC90FEABFC83" box="[274,320,786,809]" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="9">KIZ</collectionCode>
11665, an adult male from Pù Hu Nature Reserve, Thanh Hoa Province,
<collectingCountry id="F352DEBA0222E125FBBFFC93FB36FC80" box="[1147,1245,785,810]" name="Vietnam" pageId="9">Vietnam</collectingCountry>
,
<geoCoordinate id="EE71F8ED0222E125FB2CFC90FAB0FC83" box="[1256,1371,786,810]" direction="north" orientation="latitude" pageId="9" precision="925" value="20.55">20° 33 N</geoCoordinate>
,
<geoCoordinate id="EE71F8ED0222E125FAA3FC90FF3DFCE7" direction="east" orientation="longitude" pageId="9" precision="925" value="104.88333">104° 53 E</geoCoordinate>
,
<quantity id="4CBD33CF0222E125FF26FCB7FEC0FCE7" box="[226,299,821,846]" metricMagnitude="2" metricUnit="m" metricValue="6.38" pageId="9" unit="m" value="638.0">638 m</quantity>
, collected on
<date id="FFFBB8EA0222E125FE02FCB4FD93FCE4" box="[454,632,822,846]" pageId="9" value="2011-08-21">21 August 2011</date>
by Hoang Van Chung, Yang Jun Xiao and Nguyen Ngoc Sang.
</paragraph>
</subSubSection>
<subSubSection id="C35FCDA10222E125FF03FCDBFEA8FBA8" pageId="9" type="diagnosis">
<paragraph id="8BFA9E2A0222E125FF03FCDBFEA8FBA8" blockId="9.[151,1436,785,2034]" pageId="9">
<emphasis id="B93142380222E125FF03FCDBFEAAFCD8" bold="true" box="[199,321,857,882]" pageId="9">Diagnosis.</emphasis>
Medium sized, snout-vent length
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; body moderately robust; tail
<quantity id="4CBD33CF0222E125FB5CFCDBFAE5FCDB" box="[1176,1294,857,882]" metricMagnitude="-2" metricUnit="m" metricValue="8.259" pageId="9" unit="mm" value="82.59">82.59 mm</quantity>
, longer than snout-vent length, with a series of medium enlarged transversal subcaudals; a series of medium enlarged femoral scales continue from precloacal scales; femoral scales without pores; males with five precloacal pores; supralabials 8; infralabials 10; narrow subdigital lamellae on fourth toe 23; ventral scales 36; and dorsal tubercles smooth or slightly keeled.
</paragraph>
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<subSubSection id="C35FCDA10222E126FF03FB8FFB16FE05" lastPageId="10" pageId="9" type="description">
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<emphasis id="B93142380222E125FF03FB8FFE01FB8C" bold="true" box="[199,490,1037,1062]" pageId="9">
Description of
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.
</emphasis>
In addition to diagnosis. Head relatively depressed, height
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, width 15.00 mm (height/width ratio 0.64), distinct from neck; supralabials 8, continuous with a row of small scales extending to the corner of the mouth, first contacting nostril; infralabials 10; supra- and infralabials nearest to rostral and mental largest in size; rostral large, width
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, height
<quantity id="4CBD33CF0222E125FD19FBF8FCA8FB3B" box="[733,835,1146,1170]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" unit="mm" value="2.4">2.40 mm</quantity>
, vertical sunken groove in middle, contacting nostril; two internasals contact each other; mental nearly triangular, posteriorly contacting two large postmentals that contact one another and first infralabials (
<figureCitation id="137E82AF0222E125FDB6FB43FD5FFB70" box="[626,692,1217,1242]" captionStart="FIGURE 5" captionStartId="11.[151,250,1862,1884]" captionTargetBox="[155,1427,963,1835]" captionTargetId="figure@11.[151,1436,960,1841]" captionTargetPageId="11" captionText="FIGURE 5. Holotype of Cyrtodactylus puhuensis sp. nov. Precloacal-femoral area (A), ventral surface of tail (B), and lateral (C), dorsal (D), and ventral (E) views of head." httpUri="https://zenodo.org/record/230346/files/figure.png" pageId="9">Fig. 5</figureCitation>
CE); gular scales much smaller than ventrals; head scales small, granular, with scattered tubercles in occipital and temporal areas; eye relatively large,
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(eye/head width ratio 0.61), pupil vertical; supraciliary scales large, cover the eye anteriorly, superiorly, and posteriorly; ear opening oval,
<quantity id="4CBD33CF0222E125FF11FAACFED0FAEF" box="[213,315,1326,1350]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="9" unit="mm" value="1.9">1.90 mm</quantity>
, obliquely oriented, much smaller than eyes (ear/eye ratio 0.32); eye-nostril distance 7.00 mm, eyesnout distance
<quantity id="4CBD33CF0222E125FEF9FAD0FE48FAC3" box="[317,419,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="8.83" pageId="9" unit="mm" value="8.83">8.83 mm</quantity>
, eye-ear distance
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, distance between eyes
<quantity id="4CBD33CF0222E125FC1EFAD3FBABFAC3" box="[986,1088,1361,1386]" metricMagnitude="-3" metricUnit="m" metricValue="8.06" pageId="9" unit="mm" value="8.06">8.06 mm</quantity>
, distance between nostrils
<quantity id="4CBD33CF0222E125FAAFFAD0FF2DFA27" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" unit="mm" value="2.4">2.40 mm</quantity>
.
</paragraph>
<paragraph id="8BFA9E2A0222E125FF03FA1BFDB4F923" blockId="9.[151,1436,785,2034]" pageId="9">
Body elongate, trunk length
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, with weak ventral-lateral folds; dorsal scales small, granular, not homogeneous; dorsal and lateral tubercles round, smooth or slightly keeled, not in rows; dorsal tubercles between thighs and base of tail conical and keeled; ventral scales fairly large, smooth, and flattened, about 36 midbody transversal ventrals; dorsal surface of limbs with granular scales and slightly keeled conical tubercles; basal subdigital lamellae narrow and irregular, largest in joints of digits; subdigital lamellae well developed at distal phalanx, as broad as the digit. Subdigital lamellae under fourth finger 18 and toe 23; digits unwebbed; forearm length
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, crus length 16.00 mm.
</paragraph>
<paragraph id="8BFA9E2A0222E125FF03F917FE6DF8C8" blockId="9.[151,1436,785,2034]" pageId="9">
Precloacal scales large, flattened, and imbricate; about 8 large imbricate scales under a series of 8 precloacal scales arranged in Λ-shape that bears five precloacal scales with pores; a series of moderately enlarged femoral scales on each side of thigh extend from the eight precloacal scales (
<figureCitation id="137E82AF0222E125FC50F95FFC3CF95C" box="[916,983,1757,1782]" captionStart="FIGURE 5" captionStartId="11.[151,250,1862,1884]" captionTargetBox="[155,1427,963,1835]" captionTargetId="figure@11.[151,1436,960,1841]" captionTargetPageId="11" captionText="FIGURE 5. Holotype of Cyrtodactylus puhuensis sp. nov. Precloacal-femoral area (A), ventral surface of tail (B), and lateral (C), dorsal (D), and ventral (E) views of head." httpUri="https://zenodo.org/record/230346/files/figure.png" pageId="9">Fig. 5</figureCitation>
A); precloacal groove and femoral pores absent; tail round in cross-section,
<quantity id="4CBD33CF0222E125FDF1F883FD47F8B3" box="[565,684,1793,1818]" metricMagnitude="-2" metricUnit="m" metricValue="8.259" pageId="9" unit="mm" value="82.59">82.59 mm</quantity>
in length, longer than snout-vent length; transversal subcaudals medium size, about a half size of tail width (
<figureCitation id="137E82AF0222E125FD5AF8A7FD0FF894" box="[670,740,1829,1854]" captionStart="FIGURE 5" captionStartId="11.[151,250,1862,1884]" captionTargetBox="[155,1427,963,1835]" captionTargetId="figure@11.[151,1436,960,1841]" captionTargetPageId="11" captionText="FIGURE 5. Holotype of Cyrtodactylus puhuensis sp. nov. Precloacal-femoral area (A), ventral surface of tail (B), and lateral (C), dorsal (D), and ventral (E) views of head." httpUri="https://zenodo.org/record/230346/files/figure.png" pageId="9">Fig. 5</figureCitation>
B); dorsum of proximal tail with conical tubercles extending down to the base tail.
</paragraph>
<paragraph id="8BFA9E2A0222E126FF03F8EFFB17FF7F" blockId="9.[151,1436,785,2034]" lastBlockId="10.[151,1436,151,2028]" lastPageId="10" pageId="9">
<emphasis id="B93142380222E125FF03F8EFFE5CF82F" box="[199,439,1901,1925]" italics="true" pageId="9">
Color in life (
<figureCitation id="137E82AF0222E125FEA6F8ECFE42F82F" box="[354,425,1901,1925]" captionStart="FIGURE 4" captionStartId="11.[151,250,888,910]" captionTargetBox="[151,1436,193,866]" captionTargetId="figure@11.[151,1436,193,867]" captionTargetPageId="11" captionText="FIGURE 4. Holotype of Cyrtodactylus puhuensis sp. nov. in life." httpUri="https://zenodo.org/record/230345/files/figure.png" pageId="9">Fig. 4</figureCitation>
).
</emphasis>
Dorsal and lateral background of body chocolate-black; dorsum with faint and irregular pale-yellowish transverse bands; dorsal tubercles at the transverse bands yellow to white, others colored as background; flanks with yellow to white tubercles; head dark brown with yellowish pattern; eyelids yellow; eye brown, with black vertical pupil; nuchal band chocolate-black, interrupted at nape; upper surface of legs and digits dark brown with yellow to white bands or blotches; unregenerated tail with seven black and white to yellowish alternating rings; white rings much more narrow than black; lower surfaces of flanks and legs white.
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03FF62FAC7FEB7" blockId="10.[151,1436,151,2028]" pageId="10">
<emphasis id="B93142380221E126FF03FF62FD99FF5D" box="[199,626,224,248]" italics="true" pageId="10">
Color in preservative (
<figureCitation id="137E82AF0221E126FE1AFF62FDC1FF52" box="[478,554,224,248]" captionStart="FIGURE 6" captionStartId="12.[151,250,1085,1107]" captionTargetBox="[153,1431,207,1061]" captionTargetId="figure@12.[151,1436,193,1064]" captionTargetPageId="12" captionText="FIGURE 6. Holotype of Cyrtodactylus puhuensis sp. nov. dorsal (A) and ventral views (B) in preservative and ventral surface of right hand (C) and foot (D)." httpUri="https://zenodo.org/record/230347/files/figure.png" pageId="10">Fig. 6</figureCitation>
A, B).
</emphasis>
In preservative, color slightly faded and pattern unchanged; body background black or dark brown, dorsal transverse bands white; dorsum and flanks with white tubercles.
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03FEA5FCFFFEEA" blockId="10.[151,1436,151,2028]" box="[199,788,295,320]" pageId="10">
<emphasis id="B93142380221E126FF03FEA5FE80FEEA" bold="true" box="[199,363,295,320]" pageId="10">DNA barcode</emphasis>
: GenBank Accession No.
<accessionNumber id="941603C90221E126FD50FEA5FCFBFEEA" box="[660,784,295,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF929529" pageId="10">KF929529</accessionNumber>
.
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03FECEFB16FE05" blockId="10.[151,1436,151,2028]" pageId="10">
<emphasis id="B93142380221E126FF03FECEFE86FECF" bold="true" box="[199,365,332,357]" pageId="10">Comparisons.</emphasis>
The new species was compared to other congeners in Indochina by using the following morphological characters: precloacal pores, subcaudal scales, ventral scales, continuous medium enlarged femoral scales, dorsal tubercles, dorsal pattern, nuchal band, and number of digital lamellae under the 4th toe.
</paragraph>
</subSubSection>
<subSubSection id="C35FCDA10221E122FF03FE39FBD7FE5F" lastPageId="14" pageId="10" type="reference_group">
<paragraph id="8BFA9E2A0221E126FF03FE39FEC4FBF8" blockId="10.[151,1436,151,2028]" pageId="10">
<taxonomicName id="4C45E5A90221E126FF03FE39FE08FE79" box="[199,483,443,467]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380221E126FF03FE39FE08FE79" box="[199,483,443,467]" italics="true" pageId="10">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<emphasis id="B93142380221E126FE36FE38FDBFFE79" bold="true" box="[498,596,442,467]" pageId="10">
<taxonomicNameLabel id="A202FF430221E126FE36FE38FDBFFE79" box="[498,596,442,467]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from Vietnamese congeners, from
<taxonomicName id="4C45E5A90221E126FB9BFE39FCC4FE5D" authority="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler 2012" authorityName="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="bidoupimontis">
<emphasis id="B93142380221E126FB9BFE39FAC3FE79" box="[1119,1320,443,467]" italics="true" pageId="10">C. bidoupimontis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FAFCFE39FCC4FE5D" author="Nazarov" pageId="10" refString="Nazarov, R., Poyarkov, N. A., Orlov, N. L., Phung, T. M., Nguyen, T. T., Hoang, D. M. &amp; Ziegler, T. (2012) Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 3302, 1 - 24." type="journal article" year="2012">Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FCFAFE5DFE15FDB0" authority="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler 2012" authorityName="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="bugiamapensis">
<emphasis id="B93142380221E126FCFAFE5DFBE1FE5C" box="[830,1034,478,502]" italics="true" pageId="10">C. bugiamapensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FBD0FE5DFE15FDB0" author="Nazarov" pageId="10" refString="Nazarov, R., Poyarkov, N. A., Orlov, N. L., Phung, T. M., Nguyen, T. T., Hoang, D. M. &amp; Ziegler, T. (2012) Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 3302, 1 - 24." type="journal article" year="2012">Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FDCDFD80FA73FDB0" authority="Geissler, Nazarov, Orlov, Bohme, Phung, Nguyen &amp; Ziegler 2009" authorityName="Geissler, Nazarov, Orlov, Bohme, Phung, Nguyen &amp; Ziegler" authorityYear="2009" box="[521,1432,513,538]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="cattienensis">
<emphasis id="B93142380221E126FDCDFD80FD46FDB3" box="[521,685,514,537]" italics="true" pageId="10">C. cattienensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FD70FD83FA73FDB0" author="Geissler" box="[692,1432,513,538]" pageId="10" refString="Geissler, P., Nazarov, R., Orlov, N. L., Bohme, W., Phung, T. M., Nguyen, T. Q. &amp; Ziegler, T. (2009) A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 2161, 20 - 32." type="journal article" year="2009">Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen &amp; Ziegler 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FF53FDA4FCC2FD94" authority="Heidrich, Rosler, Vu, Bohme &amp; Ziegler 2007" authorityName="Heidrich, Rosler, Vu, Bohme &amp; Ziegler" authorityYear="2007" box="[151,809,549,574]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="cryptus">
<emphasis id="B93142380221E126FF53FDA4FEE5FD97" box="[151,270,550,573]" italics="true" pageId="10">C. cryptus</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FEDCFDA7FCC2FD94" author="Heidrich" box="[280,809,549,574]" pageId="10" refString="Heidrich, A., Rosler, H., Vu, N. T., Bohme, W. &amp; Ziegler, T. (2007) Another new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha - Ke Bang National Park, central Truong Son, Vietnam. Zootaxa, 1445, 35 - 48." type="journal article" year="2007">Heidrich, Rösler, Vu, Böhme &amp; Ziegler 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FCF3FDA4FBECFD94" authority="Ngo 2013" authorityName="Ngo" authorityYear="2013" box="[823,1031,549,574]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="dati">
<emphasis id="B93142380221E126FCF3FDA4FC6DFD97" box="[823,902,549,573]" italics="true" pageId="10">C. dati</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FC55FDA4FBECFD94" author="Ngo" box="[913,1031,550,574]" pageId="10" refString="Ngo, T. V. (2013) Cyrtodactylus dati, a new forest dwelling bent-toed gecko (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 3616, 151 - 164. http: // dx. doi. org / 10.11646 / zootaxa. 3616.2.4" type="journal article" year="2013">Ngo 2013</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FBD1FDA4FA98FD94" authority="Ngo &amp; Bauer 2008" authorityName="Ngo &amp; Bauer" authorityYear="2008" box="[1045,1395,549,574]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="huynhi">
<emphasis id="B93142380221E126FBD1FDA4FB6DFD97" box="[1045,1158,549,573]" italics="true" pageId="10">C. huynhi</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FB54FDA4FA98FD94" author="Ngo" box="[1168,1395,550,574]" pageId="10" refString="Ngo, T. V. &amp; Bauer, A. M. (2008) Descriptions of two new species of Cyrtodactylus Gray 1827 (Squamata: Gekkonidae) endemic to southern Vietnam. Zootaxa, 1715, 27 - 42." type="journal article" year="2008">Ngo &amp; Bauer 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FA46FDA4FEE4FDCA" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B93142380221E126FA46FDA4FEE4FDCA" italics="true" pageId="10">C. irregularis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD4E3DB0221E126FEE7FDCAFE56FDCB" author="Smith" box="[291,445,584,609]" pageId="10" refString="Smith, M. A. (1921 b) New or little-known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 91 (2), 423 - 440. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1921. tb 03271. x" type="journal article" year="1921" yearSuffix="b">Smith 1921b</bibRefCitation>
,
<bibRefCitation id="EFD4E3DB0221E126FE0AFDCAFDE2FDCB" author="Smith" box="[462,521,584,609]" pageId="10" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">1935</bibRefCitation>
),
<taxonomicName id="4C45E5A90221E126FDE7FDCBFAD7FDCB" authority="Rosler, Nguyen, Vu, Ngo &amp; Ziegler 2008" authorityName="Rosler, Nguyen, Vu, Ngo &amp; Ziegler" authorityYear="2008" box="[547,1340,584,609]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="pseudoquadrivirgatus">
<emphasis id="B93142380221E126FDE7FDCBFCD5FDCA" box="[547,830,584,608]" italics="true" pageId="10">C. pseudoquadrivirgatus</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FC8EFDCAFAD7FDCB" author="Rosler" box="[842,1340,584,609]" pageId="10" refString="Rosler, H., Vu, T. N., Nguyen, T. Q., Ngo, T. V. &amp; Ziegler, T. (2008) A new Cyrtodactylus (Squamata: Gekkonidae) from central Vietnam. Hamadryad, 33, 48 - 63." type="journal article" year="2008">Rösler, Nguyen, Vu, Ngo &amp; Ziegler 2008</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C45E5A90221E126FA46FDCBFD4EFD2E" authority="Nazarov, Orlov, Nguyen &amp; Ho 2008" authorityName="Nazarov, Orlov, Nguyen &amp; Ho" authorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B93142380221E126FA46FDCBFF00FD2E" italics="true" pageId="10">C. ziegleri</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FF32FDEFFD4EFD2E" author="Nazarov" box="[246,677,620,645]" pageId="10" refString="Nazarov, A. R., Orlov, N. L., Nguyen, S. N. &amp; Ho, T. C. (2008) Taxonomy of naked-toes geckos Cyrtodactylus irregularis complex of South Vietnam and description of a new species from Chu Yang Sin National Park (Krong Bong District, Dac Lac Province, Vietnam). Russian Journal of Herpetology, 15, 141 - 156." type="journal article" year="2008">Nazarov, Orlov, Nguyen &amp; Ho 2008</bibRefCitation>
</taxonomicName>
, in having a series of moderately enlarged subcaudal scales and continuous precloacal-femoral scales. The new species differs from
<taxonomicName id="4C45E5A90221E126FC6AFD12FB4AFD02" authority="Ngo 2011" authorityName="Ngo" authorityYear="2011" box="[942,1185,656,680]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="martini">
<emphasis id="B93142380221E126FC6AFD12FBCEFD0D" box="[942,1061,656,679]" italics="true" pageId="10">C. martini</emphasis>
Ngo
<quantity id="4CBD33CF0221E126FBAEFD12FB4AFD02" box="[1130,1185,656,680]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="10" unit="in" value="2011.0">2011</quantity>
</taxonomicName>
in having a series of transversal and enlarged subcaudal scales (
<emphasis id="B93142380221E126FD45FD36FD23FD61" box="[641,712,692,715]" italics="true" pageId="10">versus</emphasis>
small or medium ones, not transversely enlarged) and a higher number of precloacal pores (5
<emphasis id="B93142380221E126FE39FD55FDAEFD44" box="[509,581,727,750]" italics="true" pageId="10">versus</emphasis>
<quantity id="4CBD33CF0221E126FD8AFD55FD96FD45" box="[590,637,726,751]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" unit="in" value="4.0">4 in</quantity>
all four males from the
<typeStatus id="54FE20880221E126FC59FD55FC26FD45" box="[925,973,727,751]" pageId="10">type</typeStatus>
series). By having series of moderately enlarged femoral scales continuous to the precloacal scales,
<taxonomicName id="4C45E5A90221E126FC9FFD78FC1BFCB8" box="[859,1008,762,786]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380221E126FC9FFD78FC1BFCB8" box="[859,1008,762,786]" italics="true" pageId="10">C. puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430221E126FC38FD78FBB2FCB9" box="[1020,1113,762,787]" pageId="10" rank="species">
<emphasis id="B93142380221E126FC38FD78FBB2FCB9" bold="true" box="[1020,1113,762,787]" pageId="10">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90221E126FB3EFD78FDC0FC9C" authority="Nguyen, Orlov &amp; Darevsky 2006" authorityName="Nguyen, Orlov &amp; Darevsky" authorityYear="2006" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="badenensis">
<emphasis id="B93142380221E126FB3EFD78FA77FCB8" box="[1274,1436,762,786]" italics="true" pageId="10">C. badenensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FF53FC9CFDC0FC9C" author="Nguyen" box="[151,555,797,822]" pageId="10" refString="Nguyen, S. N, Orlov, N. L. &amp; Darevsky, I. S. (2006) Descriptions of two new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from southern Vietnam. Russian Journal of Herpetology, 13, 215 - 226." type="journal article" year="2006">Nguyen, Orlov &amp; Darevsky 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FDF8FC9CFA9BFC9C" authority="Orlov, Nguyen, Nazarov, Ananjeva &amp; Nguyen 2007" authorityName="Orlov, Nguyen, Nazarov, Ananjeva &amp; Nguyen" authorityYear="2007" box="[572,1392,797,822]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="caovansungi">
<emphasis id="B93142380221E126FDF8FC9CFD1AFC9F" box="[572,753,798,821]" italics="true" pageId="10">C. caovansungi</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FD3BFC9FFA9BFC9C" author="Orlov" box="[767,1392,797,822]" pageId="10" refString="Orlov, N. L., Nguyen, T. Q., Nazarov, R. A., Ananjeva, N. B. &amp; Nguyen, S. N. (2007) A new species of the genus Cyrtodactylus Gray, 1827 and redescription of Cyrtodactylus paradoxus (Darevsky et Szczerbak, 1997) (Squamata: Sauria: Gekkonidae) from South Vietnam. Russian Journal of Herpetology, 14, 145 - 152." type="journal article" year="2007">Orlov, Nguyen, Nazarov, Ananjeva &amp; Nguyen 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FA46FC9CFDC1FCF3" authority="Ngo &amp; Onn 2011" authorityName="Ngo &amp; Onn" authorityYear="2011" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="cucphuongensis">
<emphasis id="B93142380221E126FA46FC9CFEA1FCF3" italics="true" pageId="10">C. cucphuongensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FE92FCC3FDC1FCF3" author="Ngo" box="[342,554,833,857]" pageId="10" refString="Ngo, T. V. &amp; Onn, C. K. (2011) A new karstic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from northern Vietnam. Zootaxa, 3125, 51 - 63." type="journal article" year="2011">Ngo &amp; Onn 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FDFFFCC3FC83FCF3" authority="Ngo 2008" authorityName="Ngo" authorityYear="2008" box="[571,872,833,857]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="eisenmanae">
<emphasis id="B93142380221E126FDFFFCC3FD0EFCF2" box="[571,741,833,856]" italics="true" pageId="10">C. eisenmanae</emphasis>
Ngo 2008
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FCBCFCC3FB6AFCF3" authority="Ngo 2008" authorityName="Ngo" authorityYear="2008" box="[888,1153,833,857]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="grismeri">
<emphasis id="B93142380221E126FCBCFCC3FC16FCF2" box="[888,1021,833,856]" italics="true" pageId="10">C. grismeri</emphasis>
Ngo 2008
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FB55FCC3FE33FCD7" authority="Luu, Nguyen, Do &amp; Ziegler 2011" authorityName="Luu, Nguyen, Do &amp; Ziegler" authorityYear="2011" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="huongsonensis">
<emphasis id="B93142380221E126FB55FCC3FAB7FCF3" box="[1169,1372,833,857]" italics="true" pageId="10">C. huongsonensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FAACFCC3FE33FCD7" author="Luu" pageId="10" refString="Luu, V. Q., Nguyen, T. Q, Do, H. Q. &amp; Ziegler, T. (2011) A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam. Zootaxa, 3129, 39 - 50." type="journal article" year="2011">Luu, Nguyen, Do &amp; Ziegler 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FE20FCE7FBE6FCD7" authority="Ziegler, Phung, Le &amp; Nguyen 2013" authorityName="Ziegler, Phung, Le &amp; Nguyen" authorityYear="2013" box="[484,1037,868,893]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="kingsadai">
<emphasis id="B93142380221E126FE20FCE7FD98FCD6" box="[484,627,868,892]" italics="true" pageId="10">C. kingsadai</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FDBFFCE6FBE6FCD7" author="Ziegler" box="[635,1037,868,893]" pageId="10" refString="Ziegler, T., Phung, T. M., Le, M. D. &amp; Nguyen, T. Q. (2013) A new Cyrtodactylus (Squamata: Gekkonidae) from Phu Yen Province, southern Vietnam. Zootaxa, 3686 (4), 432 - 446. http: // dx. doi. org / 10.11646 / zootaxa. 3686.4.2" type="journal article" year="2013">Ziegler, Phung, Le &amp; Nguyen 2013</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FBDEFCE7FEAEFC0A" authority="Nguyen, Orlov &amp; Darevsky 2006" authorityName="Nguyen, Orlov &amp; Darevsky" authorityYear="2006" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="nigriocularis">
<emphasis id="B93142380221E126FBDEFCE7FB25FCD6" box="[1050,1230,868,892]" italics="true" pageId="10">C. nigriocularis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FB11FCE7FEAEFC0A" author="Nguyen" pageId="10" refString="Nguyen, S. N, Orlov, N. L. &amp; Darevsky, I. S. (2006) Descriptions of two new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from southern Vietnam. Russian Journal of Herpetology, 13, 215 - 226." type="journal article" year="2006">Nguyen, Orlov &amp; Darevsky 2006</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C45E5A90221E126FE44FC0AFD10FC0A" authority="Ngo &amp; Bauer 2008" authorityName="Ngo &amp; Bauer" authorityYear="2008" box="[384,763,904,928]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="takouensis">
<emphasis id="B93142380221E126FE44FC0AFDF3FC0A" box="[384,536,904,928]" italics="true" pageId="10">C. takouensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FDDBFC0AFD10FC0A" author="Ngo" box="[543,763,904,928]" pageId="10" refString="Ngo, T. V. &amp; Bauer, A. M. (2008) Descriptions of two new species of Cyrtodactylus Gray 1827 (Squamata: Gekkonidae) endemic to southern Vietnam. Zootaxa, 1715, 27 - 42." type="journal article" year="2008">Ngo &amp; Bauer 2008</bibRefCitation>
</taxonomicName>
. In addition, the new species differs from
<taxonomicName id="4C45E5A90221E126FB12FC0AFA98FC0A" box="[1238,1395,904,928]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="badenensis">
<emphasis id="B93142380221E126FB12FC0AFA98FC0A" box="[1238,1395,904,928]" italics="true" pageId="10">C. badenensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FA46FC0AFEA1FC69" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="cucphuongensis">
<emphasis id="B93142380221E126FA46FC0AFEA1FC69" italics="true" pageId="10">C. cucphuongensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FE9CFC2EFE17FC69" box="[344,508,940,963]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="eisenmanae">
<emphasis id="B93142380221E126FE9CFC2EFE17FC69" box="[344,508,940,963]" italics="true" pageId="10">C. eisenmanae</emphasis>
</taxonomicName>
,
<emphasis id="B93142380221E126FDCEFC2EFD7BFC69" box="[522,656,940,963]" italics="true" pageId="10">
<taxonomicName id="4C45E5A90221E126FDCEFC2EFD66FC69" box="[522,653,940,963]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="grismeri">C. grismeri</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C45E5A90221E126FD03FC2EFC91FC69" box="[711,890,939,963]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="nigriocularis">
<emphasis id="B93142380221E126FD03FC2EFC91FC69" box="[711,890,939,963]" italics="true" pageId="10">C. nigriocularis</emphasis>
</taxonomicName>
in having 5 precloacal pores (
<emphasis id="B93142380221E126FB08FC2EFAFFFC69" box="[1228,1300,940,963]" italics="true" pageId="10">versus</emphasis>
0, or
<quantity id="4CBD33CF0221E126FA91FC2EFA77FC6E" box="[1365,1436,939,964]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="5.08" metricValueMin="0.0" pageId="10" unit="in" value="1.0" valueMax="2.0" valueMin="0.0">02 in</quantity>
the last species); from
<taxonomicName id="4C45E5A90221E126FE5DFC4DFDB4FC4D" box="[409,607,975,999]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="huongsonensis">
<emphasis id="B93142380221E126FE5DFC4DFDB4FC4D" box="[409,607,975,999]" italics="true" pageId="10">C. huongsonensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C45E5A90221E126FD5CFC4DFCCCFC4D" box="[664,807,975,999]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="kingsadai">
<emphasis id="B93142380221E126FD5CFC4DFCCCFC4D" box="[664,807,975,999]" italics="true" pageId="10">C. kingsadai</emphasis>
</taxonomicName>
by absence of femoral pores (
<emphasis id="B93142380221E126FB44FC52FB23FC4D" box="[1152,1224,976,999]" italics="true" pageId="10">versus</emphasis>
17 or 37 femoral pores on both thighs, respectively); and from
<taxonomicName id="4C45E5A90221E126FD5EFC70FCA2FBA0" box="[666,841,1010,1034]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="caovansungi">
<emphasis id="B93142380221E126FD5EFC70FCA2FBA0" box="[666,841,1010,1034]" italics="true" pageId="10">C. caovansungi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C45E5A90221E126FC45FC70FAF5FBA0" authority="Ngo &amp; Grismer 2010" authorityName="Ngo &amp; Grismer" authorityYear="2010" box="[897,1310,1010,1035]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="bichnganae">
<emphasis id="B93142380221E126FC45FC70FBCDFBA0" box="[897,1062,1010,1034]" italics="true" pageId="10">C. bichnganae</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FBE9FC71FAF5FBA0" author="Ngo" box="[1069,1310,1010,1035]" pageId="10" refString="Ngo, T. V, Grismer, L. L. &amp; Grismer, J. L. (2010) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 2604, 37 - 51." type="journal article" year="2010">Ngo &amp; Grismer 2010</bibRefCitation>
</taxonomicName>
by lacking femoral pores (
<emphasis id="B93142380221E126FE83FB94FE64FB87" box="[327,399,1046,1069]" italics="true" pageId="10">versus</emphasis>
3, or 9 on each thigh, respectively) and having a less precloacal pores (5
<emphasis id="B93142380221E126FB2BFB94FADDFB87" box="[1263,1334,1046,1069]" italics="true" pageId="10">versus</emphasis>
9 or 10, respectively).
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03FBDFFCF0FAE0" blockId="10.[151,1436,151,2028]" pageId="10">
Our examination of the
<typeStatus id="54FE20880221E126FE10FBDFFDECFBDF" box="[468,519,1117,1141]" pageId="10">type</typeStatus>
,
<typeStatus id="54FE20880221E126FDD7FBDFFD98FBDF" box="[531,627,1117,1141]" pageId="10" type="paratype">paratype</typeStatus>
and additional specimens collected from the
<typeStatus id="54FE20880221E126FBB6FBDFFB49FBDF" box="[1138,1186,1117,1141]" pageId="10">type</typeStatus>
locality (Appendix 1) showed
<taxonomicName id="4C45E5A90221E126FF32FB03FBDFFB32" authority="Hoang, Orlov, Ananjeva, Johns, Hoang &amp; Dau 2007" authorityName="Hoang, Orlov, Ananjeva, Johns, Hoang &amp; Dau" authorityYear="2007" box="[246,1076,1152,1177]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="chauquangensis">
<emphasis id="B93142380221E126FF32FB03FE25FB32" box="[246,462,1152,1176]" italics="true" pageId="10">C. chauquangensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FE1CFB03FBDFFB32" author="Hoang" box="[472,1076,1152,1177]" pageId="10" refString="Hoang, Q. X., Orlov, N. L., Ananjeva, N. B., Johns, A. G., Hoang, T. N. &amp; Dau, V. Q. (2007) Description of a new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from the karst of north central Vietnam. Russian Journal of Herpetology, 14, 98 - 106." type="journal article" year="2007">Hoang, Orlov, Ananjeva, Johns, Hoang &amp; Dau 2007</bibRefCitation>
</taxonomicName>
differed from the new species. First,
<taxonomicName id="4C45E5A90221E126FF18FB26FE5DFB16" box="[220,438,1188,1212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="chauquangensis">
<emphasis id="B93142380221E126FF18FB26FE5DFB16" box="[220,438,1188,1212]" italics="true" pageId="10">C. chauquangensis</emphasis>
</taxonomicName>
has a uniform transversal butterfly-shaped pattern on dorsum (
<figureCitation id="137E82AF0221E126FB58FB21FB08FB16" box="[1180,1251,1187,1212]" captionStart="FIGURE 7" captionStartId="12.[151,250,1920,1942]" captionTargetBox="[151,1436,1188,1898]" captionTargetId="figure@12.[151,1436,1188,1898]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtodactylus chauquangensis showing butterfly-shaped pattern on dorsum." httpUri="https://zenodo.org/record/230348/files/figure.png" pageId="10">Fig. 7</figureCitation>
) while the new species shows faint, thin, and irregular dorsal bands. Second, femoral scale rows in
<taxonomicName id="4C45E5A90221E126FB55FB45FA86FB75" box="[1169,1389,1223,1247]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="chauquangensis">
<emphasis id="B93142380221E126FB55FB45FA86FB75" box="[1169,1389,1223,1247]" italics="true" pageId="10">C. chauquangensis</emphasis>
</taxonomicName>
are undeveloped; they are the same size as other femoral scales. The new species has series of moderately enlarged femoral scales (
<figureCitation id="137E82AF0221E126FE8EFA8FFE65FA8C" box="[330,398,1293,1318]" captionStart="FIGURE 5" captionStartId="11.[151,250,1862,1884]" captionTargetBox="[155,1427,963,1835]" captionTargetId="figure@11.[151,1436,960,1841]" captionTargetPageId="11" captionText="FIGURE 5. Holotype of Cyrtodactylus puhuensis sp. nov. Precloacal-femoral area (A), ventral surface of tail (B), and lateral (C), dorsal (D), and ventral (E) views of head." httpUri="https://zenodo.org/record/230346/files/figure.png" pageId="10">Fig. 5</figureCitation>
A). Third,
<taxonomicName id="4C45E5A90221E126FDCDFA8CFD0AFA8C" box="[521,737,1294,1318]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="chauquangensis">
<emphasis id="B93142380221E126FDCDFA8CFD0AFA8C" box="[521,737,1294,1318]" italics="true" pageId="10">C. chauquangensis</emphasis>
</taxonomicName>
has 67 precloacal pores, except for topotype CQ526, which has 8 pores. In contrast, the new species has 5 pores only.
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03FAD7FE8BF9CC" blockId="10.[151,1436,151,2028]" pageId="10">
The new
<taxonomicName id="4C45E5A90221E126FEEAFAD7FE9AFAC4" box="[302,369,1365,1390]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
differs from
<taxonomicName id="4C45E5A90221E126FDC0FAD7FCB9FAC7" authority="(Smith 1921a)" baseAuthorityName="Smith" baseAuthorityYear="1921" box="[516,850,1365,1390]" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="condorensis">
<emphasis id="B93142380221E126FDC0FAD7FD40FAC7" box="[516,683,1365,1389]" italics="true" pageId="10">C. condorensis</emphasis>
(
<bibRefCitation id="EFD4E3DB0221E126FD7DFAD7FCA0FAC7" author="Smith" box="[697,843,1365,1390]" pageId="10" refString="Smith, M. A. (1921 a) Reptiles and Batrachians collected on Pulo Condore. Journal of the Natural History Society of Siam, 4, 93 - 97." type="journal article" year="1921" yearSuffix="a">Smith 1921a</bibRefCitation>
)
</taxonomicName>
by the distribution of precloacal pores (in a chevron
<emphasis id="B93142380221E126FF53FAFBFF34FA3A" box="[151,223,1401,1424]" italics="true" pageId="10">versus</emphasis>
<quantity id="4CBD33CF0221E126FF2CFAFBFED9FA3B" box="[232,306,1400,1425]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.778" metricValueMin="1.016" pageId="10" unit="in" value="5.5" valueMax="7.0" valueMin="4.0">47 in</quantity>
two juxtaposed rows) and dorsal tubercles (smooth or slightly keeled
<emphasis id="B93142380221E126FBA1FAFBFB47FA3A" box="[1125,1196,1401,1424]" italics="true" pageId="10">versus</emphasis>
subtrihedral) (
<bibRefCitation id="EFD4E3DB0221E126FA93FAFAFF39FA1F" author="Smith" pageId="10" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
;
<bibRefCitation id="EFD4E3DB0221E126FF24FA1FFE30FA1E" author="Ngo" box="[224,475,1436,1461]" pageId="10" refString="Ngo, T. V. &amp; Grismer, L. L. (2012) A new endemic species of Cyrtodactylus (Squamata: Gekkonidae) from Tho Chu Island, southeastern Vietnam. Zootaxa, 3228, 48 - 60." type="journal article" year="2012">Ngo &amp; Grismer 2012</bibRefCitation>
); from
<taxonomicName id="4C45E5A90221E126FDF4FA1EFBC6FA1E" authority="Darevsky &amp; Szczerbak 1997" authorityName="Darevsky &amp; Szczerbak" authorityYear="1997" box="[560,1069,1436,1461]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="paradoxus">
<emphasis id="B93142380221E126FDF4FA1EFD22FA1E" box="[560,713,1436,1460]" italics="true" pageId="10">C. paradoxus</emphasis>
(
<bibRefCitation id="EFD4E3DB0221E126FD1EFA1EFBCEFA1E" author="Darevsky" box="[730,1061,1436,1461]" pageId="10" refString="Darevsky, I. S. &amp; Szczerbak, N. N. (1997) A new gecko of the genus Gonydactylus (Sauria: Gekkonidae) with a key to the species from Vietnam. Asiatic Herpetological Research, 7, 19 - 22." type="journal article" year="1997">Darevsky &amp; Szczerbak 1997</bibRefCitation>
)
</taxonomicName>
by having more and differently arranged precloacal pores (
<quantity id="4CBD33CF0221E126FE16FA3DFDEAFA72" box="[466,513,1471,1496]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" unit="in" value="5.0">5 in</quantity>
chevron
<emphasis id="B93142380221E126FDB5FA42FD53FA7D" box="[625,696,1472,1495]" italics="true" pageId="10">versus</emphasis>
<quantity id="4CBD33CF0221E126FD00FA3DFCE4FA72" box="[708,783,1471,1496]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="10.16" metricValueMin="0.0" pageId="10" unit="in" value="2.0" valueMax="4.0" valueMin="0.0">04 in</quantity>
two juxtaposed rows) (Orlov
<emphasis id="B93142380221E126FBB7FA42FB44FA7D" box="[1139,1199,1471,1495]" italics="true" pageId="10">et al.</emphasis>
2007); and from
<taxonomicName id="4C45E5A90221E126FA46FA42FDF0FA51" authority="Ngo &amp; Grismer 2012" authorityName="Ngo &amp; Grismer" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="thochuensis">
<emphasis id="B93142380221E126FA46FA42FEF0FA51" italics="true" pageId="10">C. thochuensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FEE0FA61FDF0FA51" author="Ngo" box="[292,539,1506,1531]" pageId="10" refString="Ngo, T. V. &amp; Grismer, L. L. (2012) A new endemic species of Cyrtodactylus (Squamata: Gekkonidae) from Tho Chu Island, southeastern Vietnam. Zootaxa, 3228, 48 - 60." type="journal article" year="2012">Ngo &amp; Grismer 2012</bibRefCitation>
</taxonomicName>
by the chevron arrangement of its precloacal pores (
<emphasis id="B93142380221E126FBBBFA61FB2CFA50" box="[1151,1223,1507,1530]" italics="true" pageId="10">versus</emphasis>
in two juxtaposed rows), smooth or slightly keeled dorsal tubercles (
<emphasis id="B93142380221E126FD26F985FCC2F9B4" box="[738,809,1543,1566]" italics="true" pageId="10">versus</emphasis>
strongly keeled), normal sized scales on heel (
<emphasis id="B93142380221E126FA90F985FA70F9B4" box="[1364,1435,1543,1566]" italics="true" pageId="10">versus</emphasis>
enlarged), and having the dorsal caudal tubercles extending to the base of the tail only (
<emphasis id="B93142380221E126FB57F9A8FB30F9EB" box="[1171,1243,1578,1601]" italics="true" pageId="10">versus</emphasis>
to about 1/3 the length of the tail).
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03F9F3FA82F8BC" blockId="10.[151,1436,151,2028]" pageId="10">
<taxonomicName id="4C45E5A90221E126FF03F9F3FE36F923" box="[199,477,1649,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380221E126FF03F9F3FE36F923" box="[199,477,1649,1673]" italics="true" pageId="10">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430221E126FE21F9F2FDAAF923" box="[485,577,1648,1673]" pageId="10" rank="species">
<emphasis id="B93142380221E126FE21F9F2FDAAF923" bold="true" box="[485,577,1648,1673]" pageId="10">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90221E126FD1EF9F3FB03F923" authority="Ngo, Grismer &amp; Grismer 2008" authorityName="Ngo, Grismer &amp; Grismer" authorityYear="2008" box="[730,1256,1648,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="hontreensis">
<emphasis id="B93142380221E126FD1EF9F3FC96F923" box="[730,893,1649,1673]" italics="true" pageId="10">C. hontreensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FC42F9F3FB03F923" author="Ngo" box="[902,1256,1648,1673]" pageId="10" refString="Ngo, T. V., Grismer, L. L. &amp; Grismer, J. L. (2008) A new endemic cave dwelling species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 1967, 53 - 62." type="journal article" year="2008">Ngo, Grismer &amp; Grismer 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90221E126FB31F9F3FA77F923" box="[1269,1436,1649,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93142380221E126FB31F9F3FA77F923" box="[1269,1436,1649,1673]" italics="true" pageId="10">C. intermedius</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD4E3DB0221E126FF5AF916FECDF906" author="Smith" box="[158,294,1684,1709]" pageId="10" refString="Smith, M. A. (1917) Descriptions of new reptiles and a new batrachian from Siam. Journal of the Natural History Society of Siam, 2, 221 - 225." type="journal article" year="1917">Smith 1917</bibRefCitation>
,
<bibRefCitation id="EFD4E3DB0221E126FEF6F916FE86F907" author="Smith" box="[306,365,1684,1709]" pageId="10" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">1935</bibRefCitation>
), and
<taxonomicName id="4C45E5A90221E126FE77F916FC38F906" authority="Ngo, Grismer &amp; Grismer 2010" authorityName="Ngo, Grismer &amp; Grismer" authorityYear="2010" box="[435,979,1684,1709]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="phuquocensis">
<emphasis id="B93142380221E126FE77F916FD86F906" box="[435,621,1684,1708]" italics="true" pageId="10">C. phuquocensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FDB2F917FC38F906" author="Ngo" box="[630,979,1684,1709]" pageId="10" refString="Ngo, T. V, Grismer, L. L. &amp; Grismer, J. L. (2010) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 2604, 37 - 51." type="journal article" year="2010">
Ngo, Grismer &amp; Grismer
<quantity id="4CBD33CF0221E126FC5FF916FC38F906" box="[923,979,1684,1708]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="10" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in having faint, irregular pale yellowish dorsal bands. In contrast,
<taxonomicName id="4C45E5A90221E126FE72F93AFDBCF965" box="[438,599,1719,1743]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="hontreensis">
<emphasis id="B93142380221E126FE72F93AFDBCF965" box="[438,599,1719,1743]" italics="true" pageId="10">C. hontreensis</emphasis>
</taxonomicName>
has 3 broad bands between limbs and
<taxonomicName id="4C45E5A90221E126FBC3F93AFB47F965" box="[1031,1196,1719,1743]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93142380221E126FBC3F93AFB47F965" box="[1031,1196,1719,1743]" italics="true" pageId="10">C. intermedius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C45E5A90221E126FB27F93AFA77F965" box="[1251,1436,1719,1743]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="phuquocensis">
<emphasis id="B93142380221E126FB27F93AFA77F965" box="[1251,1436,1719,1743]" italics="true" pageId="10">C. phuquocensis</emphasis>
</taxonomicName>
have 4. The new species also differs in its absence of nuchal band, which is always present in the other species, and in having 5 precloacal pores
<emphasis id="B93142380221E126FE1DF97DFDCBF8BC" box="[473,544,1791,1814]" italics="true" pageId="10">versus</emphasis>
<quantity id="4CBD33CF0221E126FDECF97DFD84F8BD" box="[552,623,1790,1815]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="10" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">78 in</quantity>
<taxonomicName id="4C45E5A90221E126FDB2F97DFCFDF8BC" box="[630,790,1790,1814]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="hontreensis">
<emphasis id="B93142380221E126FDB2F97DFCFDF8BC" box="[630,790,1790,1814]" italics="true" pageId="10">C. hontreensis</emphasis>
</taxonomicName>
,
<quantity id="4CBD33CF0221E126FCE0F97DFC91F8BD" box="[804,890,1790,1815]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="10" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">810 in</quantity>
<taxonomicName id="4C45E5A90221E126FC44F97DFBCEF8BC" box="[896,1061,1790,1814]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93142380221E126FC44F97DFBCEF8BC" box="[896,1061,1790,1814]" italics="true" pageId="10">C. intermedius</emphasis>
</taxonomicName>
and
<quantity id="4CBD33CF0221E126FB98F97CFB49F8BD" box="[1116,1186,1790,1815]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="10" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">68 in</quantity>
<taxonomicName id="4C45E5A90221E126FB6DF97DFA89F8BC" box="[1193,1378,1790,1814]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="phuquocensis">
<emphasis id="B93142380221E126FB6DF97DFA89F8BC" box="[1193,1378,1790,1814]" italics="true" pageId="10">C. phuquocensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA9E2A0221E126FF03F8A0FD77F846" blockId="10.[151,1436,151,2028]" pageId="10">
The new species differs from
<taxonomicName id="4C45E5A90221E126FDDEF8A0FB00F890" authority="Ziegler, Rosler, Herrmann &amp; Vu 2003" authorityName="Ziegler, Rosler, Herrmann &amp; Vu" authorityYear="2003" box="[538,1259,1826,1851]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="phongnhakebangensis">
<emphasis id="B93142380221E126FDDEF8A0FCDFF890" box="[538,820,1826,1850]" italics="true" pageId="10">C. phongnhakebangensis</emphasis>
Ziegler, Rösler, Herrmann &amp; Vu 2003
</taxonomicName>
and
<taxonomicName id="4C45E5A90221E126FAE0F8A0FC6AF8F4" authority="Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh &amp; Schmitz 2010" authorityName="Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh &amp; Schmitz" authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="roesleri">
<emphasis id="B93142380221E126FAE0F8A0FA77F890" box="[1316,1436,1826,1850]" italics="true" pageId="10">C. roesleri</emphasis>
Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh &amp; Schmitz
<quantity id="4CBD33CF0221E126FC8DF8C4FC6AF8F4" box="[841,897,1862,1886]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="10" unit="in" value="2010.0">2010</quantity>
</taxonomicName>
in lacking femoral pores (
<emphasis id="B93142380221E126FB6CF8C4FB1BF8F7" box="[1192,1264,1862,1885]" italics="true" pageId="10">versus</emphasis>
2742 and 20 28, respectively, precloaacal-femoral pores situated as one continuous row). The new
<taxonomicName id="4C45E5A90221E126FB5DF8EAFB37F82B" box="[1177,1244,1896,1921]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
differs from
<taxonomicName id="4C45E5A90221E126FA46F8EBFDECF80F" authority="Ngo &amp; Onn 2010" authorityName="Ngo &amp; Onn" authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" phylum="Chordata" rank="species" species="yangbayensis">
<emphasis id="B93142380221E126FA46F8EBFEC5F80E" italics="true" pageId="10">C. yangbayensis</emphasis>
<bibRefCitation id="EFD4E3DB0221E126FEFCF80FFDECF80F" author="Ngo" box="[312,519,1933,1957]" pageId="10" refString="Ngo, T. V &amp; Onn, C. K. (2010) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Khanh Hoa Province, southern Vietnam. Zootaxa, 2504, 47 - 60." type="journal article" year="2010">
Ngo &amp; Onn
<quantity id="4CBD33CF0221E126FE0BF80FFDECF80F" box="[463,519,1933,1957]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="10" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in possessing fewer precloacal pores (5
<emphasis id="B93142380221E126FC2CF80FFBDBF80E" box="[1000,1072,1933,1956]" italics="true" pageId="10">versus</emphasis>
68), more subdigital lamellae under the fourth toe (23
<emphasis id="B93142380221E126FE6DF833FE1AF862" box="[425,497,1969,1992]" italics="true" pageId="10">versus</emphasis>
1517), medium sized scales on heel (
<emphasis id="B93142380221E126FC62F833FC05F862" box="[934,1006,1969,1992]" italics="true" pageId="10">versus</emphasis>
910 enlarged scales), and seven pale rings on the tail (
<emphasis id="B93142380221E126FE92F856FE75F841" box="[342,414,2004,2027]" italics="true" pageId="10">versus</emphasis>
911 irregular bands).
</paragraph>
<caption id="DF3ACEA20220E127FF53FCFAFCD8FC24" box="[151,819,888,910]" httpUri="https://zenodo.org/record/230345/files/figure.png" pageId="11" targetBox="[151,1436,193,866]" targetPageId="11">
<paragraph id="8BFA9E2A0220E127FF53FCFAFCD8FC24" blockId="11.[151,819,888,910]" box="[151,819,888,910]" pageId="11">
<emphasis id="B93142380220E127FF53FCFAFEF9FC24" bold="true" box="[151,274,888,910]" pageId="11">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="4C45E5A90220E127FE5DFCFAFD79FC24" box="[409,658,888,910]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380220E127FE5DFCFAFD79FC24" box="[409,658,888,910]" italics="true" pageId="11">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430220E127FD5DFCFAFD02FC24" box="[665,745,888,910]" pageId="11" rank="species">
<emphasis id="B93142380220E127FD5DFCFAFD02FC24" bold="true" box="[665,745,888,910]" pageId="11">sp. nov.</emphasis>
</taxonomicNameLabel>
in life.
</paragraph>
</caption>
<caption id="DF3ACEA20220E127FF53F8C4FD80F8D1" httpUri="https://zenodo.org/record/230346/files/figure.png" pageId="11" targetBox="[155,1427,963,1835]" targetPageId="11">
<paragraph id="8BFA9E2A0220E127FF53F8C4FD80F8D1" blockId="11.[151,1435,1862,1915]" pageId="11">
<emphasis id="B93142380220E127FF53F8C4FEF8F8F6" bold="true" box="[151,275,1862,1884]" pageId="11">FIGURE 5.</emphasis>
Holotype of
<taxonomicName id="4C45E5A90220E127FE58F8C4FD7DF8F6" box="[412,662,1862,1884]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380220E127FE58F8C4FD7DF8F6" box="[412,662,1862,1884]" italics="true" pageId="11">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430220E127FD5AF8C4FD04F8F6" box="[670,751,1862,1884]" pageId="11" rank="species">
<emphasis id="B93142380220E127FD5AF8C4FD04F8F6" bold="true" box="[670,751,1862,1884]" pageId="11">sp. nov.</emphasis>
</taxonomicNameLabel>
Precloacal-femoral area (A), ventral surface of tail (B), and lateral (C), dorsal (D), and ventral (E) views of head.
</paragraph>
</caption>
<caption id="DF3ACEA20227E120FF53FBBFFE27FBD8" httpUri="https://zenodo.org/record/230347/files/figure.png" pageId="12" targetBox="[153,1431,207,1061]" targetPageId="12">
<paragraph id="8BFA9E2A0227E120FF53FBBFFE27FBD8" blockId="12.[151,1436,1085,1138]" pageId="12">
<emphasis id="B93142380227E120FF53FBBFFEF9FBF9" bold="true" box="[151,274,1085,1107]" pageId="12">FIGURE 6.</emphasis>
Holotype of
<taxonomicName id="4C45E5A90227E120FE5DFBBFFD79FBF9" box="[409,658,1085,1107]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380227E120FE5DFBBFFD79FBF9" box="[409,658,1085,1107]" italics="true" pageId="12">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430227E120FD5DFBBFFD02FBF9" box="[665,745,1085,1107]" pageId="12" rank="species">
<emphasis id="B93142380227E120FD5DFBBFFD02FBF9" bold="true" box="[665,745,1085,1107]" pageId="12">sp. nov.</emphasis>
</taxonomicNameLabel>
dorsal (A) and ventral views (B) in preservative and ventral surface of right hand (C) and foot (D).
</paragraph>
</caption>
<caption id="DF3ACEA20227E120FF53F802FBF5F83C" box="[151,1054,1920,1942]" httpUri="https://zenodo.org/record/230348/files/figure.png" pageId="12" targetBox="[151,1436,1188,1898]" targetPageId="12">
<paragraph id="8BFA9E2A0227E120FF53F802FBF5F83C" blockId="12.[151,1054,1920,1942]" box="[151,1054,1920,1942]" pageId="12">
<emphasis id="B93142380227E120FF53F802FEF8F83C" bold="true" box="[151,275,1920,1942]" pageId="12">FIGURE 7.</emphasis>
<taxonomicName id="4C45E5A90227E120FEDDF802FDBBF83C" box="[281,592,1920,1942]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="chauquangensis">
<emphasis id="B93142380227E120FEDDF802FDBBF83C" box="[281,592,1920,1942]" italics="true" pageId="12">Cyrtodactylus chauquangensis</emphasis>
</taxonomicName>
showing butterfly-shaped pattern on dorsum.
</paragraph>
</caption>
<paragraph id="8BFA9E2A0226E121FF03FF1AFDCFFE5E" blockId="13.[151,1436,151,2015]" pageId="13">
For Laotian congeners,
<taxonomicName id="4C45E5A90226E121FE18FF1AFD1FFF1A" box="[476,756,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380226E121FE18FF1AFD1FFF1A" box="[476,756,152,176]" italics="true" pageId="13">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430226E121FD3BFF15FCB7FF1A" box="[767,860,151,176]" pageId="13" rank="species">
<emphasis id="B93142380226E121FD3BFF15FCB7FF1A" bold="true" box="[767,860,151,176]" pageId="13">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90226E121FC38FF1AFF39FF7E" authority="David, Teynie &amp; Ohler 2004" authorityName="David, Teynie &amp; Ohler" authorityYear="2004" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="B93142380226E121FC38FF1AFB6FFF1A" box="[1020,1156,152,176]" italics="true" pageId="13">C. buchardi</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FB4BFF15FF39FF7E" author="David" pageId="13" refString="David, P., Teynie, A. &amp; Ohler, A. (2004) A new species of Cyrtodactylus Gray 1827 (Reptilia: Squamata: Gekkonidae) from southern Laos. The Raffles Bulletin of Zoology, 52, 621 - 627." type="journal article" year="2004">David, Teynié &amp; Ohler 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FF20FF3EFBAFFF7E" authority="Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler 2011" authorityName="Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler" authorityYear="2011" box="[228,1092,188,213]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="pageli">
<emphasis id="B93142380226E121FF20FF3EFEB9FF7E" box="[228,338,188,212]" italics="true" pageId="13">C. pageli</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FEA4FF3EFBAFFF7E" author="Schneider" box="[352,1092,188,213]" pageId="13" refString="Schneider, N., Nguyen, T. Q., Schmitz, A., Kingsada, P., Auer, M. &amp; Ziegler, T. (2011) A new species of karst dwelling Cyrtodactylus (Squamata: Gekkonidae) from northwestern Laos. Zootaxa, 2930, 1 - 21." type="journal article" year="2011">Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler 2011</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C45E5A90226E121FB4AFF3EFD89FF52" authority="Nguyen, Kingsada, Rosler, Auer &amp; Ziegler 2010" authorityName="Nguyen, Kingsada, Rosler, Auer &amp; Ziegler" authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="wayakonei">
<emphasis id="B93142380226E121FB4AFF3EFAC6FF7E" box="[1166,1325,188,212]" italics="true" pageId="13">C. wayakonei</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FAF8FF3FFD89FF52" author="Nguyen" pageId="13" refString="Nguyen, T. Q., Kingsada, P., Rosler, H., Auer, M. &amp; Ziegler, T. (2010) A new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos. Zootaxa, 2652, 1 - 16." type="journal article" year="2010">
Nguyen, Kingsada, Rösler, Auer &amp; Ziegler
<quantity id="4CBD33CF0226E121FDEEFF62FD89FF52" box="[554,610,224,248]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="13" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in having a series of enlarged femoral scales (
<emphasis id="B93142380226E121FB4EFF63FB39FF52" box="[1162,1234,225,248]" italics="true" pageId="13">versus</emphasis>
without enlarged femoral scales). The new species also differs from
<taxonomicName id="4C45E5A90226E121FD25FE86FC83FEB6" box="[737,872,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="B93142380226E121FD25FE86FC83FEB6" box="[737,872,260,284]" italics="true" pageId="13">C. buchardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C45E5A90226E121FC66FE86FBD6FEB6" box="[930,1085,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="wayakonei">
<emphasis id="B93142380226E121FC66FE86FBD6FEB6" box="[930,1085,260,284]" italics="true" pageId="13">C. wayakonei</emphasis>
</taxonomicName>
in having enlarged transversal subcaudal scales (
<emphasis id="B93142380226E121FEADFEABFE5AFEEA" box="[361,433,297,320]" italics="true" pageId="13">versus</emphasis>
small or medium scales, not transversally enlarged); and from
<taxonomicName id="4C45E5A90226E121FB5FFEAAFAEDFEEA" box="[1179,1286,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="pageli">
<emphasis id="B93142380226E121FB5FFEAAFAEDFEEA" box="[1179,1286,296,320]" italics="true" pageId="13">C. pageli</emphasis>
</taxonomicName>
in lacking a nuchal band (
<emphasis id="B93142380226E121FEF6FECFFE91FECE" box="[306,378,333,356]" italics="true" pageId="13">versus</emphasis>
dark nuchal band).
<taxonomicName id="4C45E5A90226E121FD99FECEFC98FECE" box="[605,883,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380226E121FD99FECEFC98FECE" box="[605,883,332,356]" italics="true" pageId="13">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430226E121FCBFFECEFC3EFECF" box="[891,981,332,357]" pageId="13" rank="species">
<emphasis id="B93142380226E121FCBFFECEFC3EFECF" bold="true" box="[891,981,332,357]" pageId="13">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90226E121FBA9FECEFA80FECE" authority="Ulber 1993" authorityName="Ulber" authorityYear="1993" box="[1133,1387,332,357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="jarujini">
<emphasis id="B93142380226E121FBA9FECEFB08FECE" box="[1133,1251,332,356]" italics="true" pageId="13">C. jarujini</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FB2FFECEFA80FECE" author="Ulber" box="[1259,1387,332,357]" pageId="13" refString="Ulber, T. (1993) Bemerkungen uber cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen Aus Dem Zoologischen Museum in Berlin, 69, 187 - 200. http: // dx. doi. org / 10.1002 / mmnz. 19930690202" type="journal article" year="1993">Ulber 1993</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C45E5A90226E121FF53FEF2FDB2FE22" authority="Ngo &amp; Pauwels 2010" authorityName="Ngo &amp; Pauwels" authorityYear="2010" box="[151,601,367,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="lomyenensis">
<emphasis id="B93142380226E121FF53FEF2FEACFE22" box="[151,327,368,392]" italics="true" pageId="13">C. lomyenensis</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FE90FEF2FDB2FE22" author="Ngo" box="[340,601,367,392]" pageId="13" refString="Ngo, T. V. &amp; Pauwels, O. (2010) A new cave-dwelling species of Cyrtodactylus Gray, 1827 (Squamata: Gekkodidae) from Khammouane Province, southern Laos. Zootaxa, 2730, 44 - 56." type="journal article" year="2010">
Ngo &amp; Pauwels
<quantity id="4CBD33CF0226E121FDE5FEF2FDB2FE22" box="[545,601,368,392]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="13" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in lacking femoral pores (5254 and 3940, respectively, continuous precloacal-femoral pores). The new
<taxonomicName id="4C45E5A90226E121FDEBFE16FD99FE07" box="[559,626,404,429]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
differs from
<taxonomicName id="4C45E5A90226E121FCC3FE16FBD4FE06" authority="Ulber 1993" authorityName="Ulber" authorityYear="1993" box="[775,1087,404,429]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="B93142380226E121FCC3FE16FC53FE06" box="[775,952,404,428]" italics="true" pageId="13">C. interdigitalis</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FC04FE16FBD4FE06" author="Ulber" box="[960,1087,404,429]" pageId="13" refString="Ulber, T. (1993) Bemerkungen uber cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen Aus Dem Zoologischen Museum in Berlin, 69, 187 - 200. http: // dx. doi. org / 10.1002 / mmnz. 19930690202" type="journal article" year="1993">Ulber 1993</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C45E5A90226E121FBB2FE16FE2EFE7A" authority="David, Nguyen, Schneider &amp; Ziegler 2011" authorityName="David, Nguyen, Schneider &amp; Ziegler" authorityYear="2011" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="teyniei">
<emphasis id="B93142380226E121FBB2FE16FB09FE06" box="[1142,1250,404,428]" italics="true" pageId="13">C. teyniei</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FB2DFE16FE2EFE7A" author="David" pageId="13" refString="David, P., Nguyen, T. Q., Schneider, N. &amp; Ziegler, T. (2011) A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae). Zootaxa, 2833, 29 - 40." type="journal article" year="2011">
David, Nguyen, Schneider &amp; Ziegler
<quantity id="4CBD33CF0226E121FE4AFE3AFE2EFE7A" box="[398,453,440,464]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="13" unit="in" value="2011.0">2011</quantity>
</bibRefCitation>
</taxonomicName>
in having fewer precloacal pores (5
<emphasis id="B93142380226E121FCBDFE3BFC2AFE7A" box="[889,961,441,464]" italics="true" pageId="13">versus</emphasis>
14) and also differs from the former by lacking femoral pores (
<emphasis id="B93142380226E121FE5FFE5FFE08FE5E" box="[411,483,477,500]" italics="true" pageId="13">versus</emphasis>
89).
</paragraph>
<paragraph id="8BFA9E2A0226E121FF03FD82FEC4FCBF" blockId="13.[151,1436,151,2015]" pageId="13">
From
<collectingCountry id="F352DEBA0226E121FECFFD82FE93FDB2" box="[267,376,512,536]" name="Myanmar" pageId="13">Myanmar</collectingCountry>
congeners,
<taxonomicName id="4C45E5A90226E121FE3BFD82FCFFFDB2" box="[511,788,512,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380226E121FE3BFD82FCFFFDB2" box="[511,788,512,536]" italics="true" pageId="13">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430226E121FCDFFE7DFC9EFDB2" box="[795,885,511,536]" pageId="13" rank="species">
<emphasis id="B93142380226E121FCDFFE7DFC9EFDB2" bold="true" box="[795,885,511,536]" pageId="13">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90226E121FBCEFD82FA9CFDB2" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[1034,1399,512,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="B93142380226E121FBCEFD82FB01FDB2" box="[1034,1258,512,536]" italics="true" pageId="13">C. ayeyarwadyensis</emphasis>
Bauer 2003
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FA46FD82FE5CFD96" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="brevidactylus">
<emphasis id="B93142380226E121FA46FD82FEC6FD96" italics="true" pageId="13">C. brevidactylus</emphasis>
Bauer 2003
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FE06FDA6FD41FD96" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[450,682,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="gansi">
<emphasis id="B93142380226E121FE06FDA6FDCBFD96" box="[450,544,548,572]" italics="true" pageId="13">C. gansi</emphasis>
Bauer 2003
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FD70FDA6FBCAFD96" authority="Mahony 2009" authorityName="Mahony" authorityYear="2009" box="[692,1057,548,573]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="mandalayensis">
<emphasis id="B93142380226E121FD70FDA6FC91FD96" box="[692,890,548,572]" italics="true" pageId="13">C. mandalayensis</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FC46FDA6FBCAFD96" author="Mahony" box="[898,1057,548,573]" pageId="13" refString="Mahony, S. (2009) Taxonomic status of Cyrtodactylus khasiensis tamaiensis (Smith, 1940) and description of a new species allied to C. chrysopylos Bauer, 2003 from Myanmar (Reptilia: Gekkonidae). Hamadryad, 34, 62 - 74." type="journal article" year="2009">Mahony 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FBEFFDA6FA83FD96" authority="Mahony 2009" authorityName="Mahony" authorityYear="2009" box="[1067,1384,548,573]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="tamaiensis">
<emphasis id="B93142380226E121FBEFFDA6FB28FD96" box="[1067,1219,548,572]" italics="true" pageId="13">C. tamaiensis</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FB0EFDA6FA83FD96" author="Mahony" box="[1226,1384,548,573]" pageId="13" refString="Mahony, S. (2009) Taxonomic status of Cyrtodactylus khasiensis tamaiensis (Smith, 1940) and description of a new species allied to C. chrysopylos Bauer, 2003 from Myanmar (Reptilia: Gekkonidae). Hamadryad, 34, 62 - 74." type="journal article" year="2009">Mahony 2009</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C45E5A90226E121FF53FDCAFE5CFDCA" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[151,439,584,608]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="wakeorum">
<emphasis id="B93142380226E121FF53FDCAFEC0FDCA" box="[151,299,584,608]" italics="true" pageId="13">C. wakeorum</emphasis>
Bauer 2003
</taxonomicName>
, in having moderately enlarged transversal subcaudal scales. The new species also differs from
<taxonomicName id="4C45E5A90226E121FF10FDEEFE5FFD2E" box="[212,436,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="B93142380226E121FF10FDEEFE5FFD2E" box="[212,436,620,644]" italics="true" pageId="13">C. ayeyarwadyensis</emphasis>
</taxonomicName>
,
<emphasis id="B93142380226E121FE06FDEEFD92FD2E" box="[450,633,620,644]" italics="true" pageId="13">
C,
<taxonomicName id="4C45E5A90226E121FE27FDEEFD92FD2E" box="[483,633,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="brevidactylus">brevidactylus</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C45E5A90226E121FD73FDEEFCFFFD2E" box="[695,788,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="gansi">
<emphasis id="B93142380226E121FD73FDEEFCFFFD2E" box="[695,788,620,644]" italics="true" pageId="13">C. gansi</emphasis>
</taxonomicName>
in having few precloacal pores (
<emphasis id="B93142380226E121FB40FDEFFB27FD2E" box="[1156,1228,621,644]" italics="true" pageId="13">versus</emphasis>
1028, 8, and 16 29, respectively); from
<taxonomicName id="4C45E5A90226E121FE59FD12FE11FD0D" box="[413,506,656,679]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="gansi">
<emphasis id="B93142380226E121FE59FD12FE11FD0D" box="[413,506,656,679]" italics="true" pageId="13">C. gansi</emphasis>
</taxonomicName>
in lacking a shallow precloacal groove in males; and from
<taxonomicName id="4C45E5A90226E121FB4AFD12FACDFD0D" box="[1166,1318,656,679]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="tamaiensis">
<emphasis id="B93142380226E121FB4AFD12FACDFD0D" box="[1166,1318,656,679]" italics="true" pageId="13">C. tamaiensis</emphasis>
</taxonomicName>
in lacking femoral pores (
<emphasis id="B93142380226E121FE8BFD37FE7DFD66" box="[335,406,693,716]" italics="true" pageId="13">versus</emphasis>
40 precloacal-femoral pores in an almost continuous series,
<bibRefCitation id="EFD4E3DB0226E121FBBCFD36FAF4FD66" author="Mahony" box="[1144,1311,692,717]" pageId="13" refString="Mahony, S. (2009) Taxonomic status of Cyrtodactylus khasiensis tamaiensis (Smith, 1940) and description of a new species allied to C. chrysopylos Bauer, 2003 from Myanmar (Reptilia: Gekkonidae). Hamadryad, 34, 62 - 74." type="journal article" year="2009">Mahony 2009</bibRefCitation>
). From
<taxonomicName id="4C45E5A90226E121FA46FD36FEA1FD5A" authority="Gray" authorityName="Gray" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="B93142380226E121FA46FD36FEE1FD5A" italics="true" pageId="13">C. pulchellus</emphasis>
Gray
</taxonomicName>
and
<taxonomicName id="4C45E5A90226E121FE47FD5AFD7FFD5A" authority="Blyth 1861" authorityName="Blyth" authorityYear="1861" box="[387,660,727,752]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="rubidus">
<emphasis id="B93142380226E121FE47FD5AFE11FD5A" box="[387,506,728,752]" italics="true" pageId="13">C. rubidus</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FDCFFD55FD67FD5A" author="Blyth" box="[523,652,727,752]" pageId="13" refString="Blyth, E. (1861) Report of the Curator. Journal of the Asiatic Society of Bengal, 29, 87 - 115." type="journal article" year="1861">Blyth 1861</bibRefCitation>
)
</taxonomicName>
the new species differs in lacking a precloacal groove (
<bibRefCitation id="EFD4E3DB0226E121FACBFD55FA7CFD5A" author="Smith" box="[1295,1431,727,752]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
;
<bibRefCitation id="EFD4E3DB0226E121FF53FD7EFEC9FCBF" author="Taylor" box="[151,290,764,789]" pageId="13" refString="Taylor, E. H. (1963) The lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
).
</paragraph>
<paragraph id="8BFA9E2A0226E121FF03FCA2FC04FAFF" blockId="13.[151,1436,151,2015]" pageId="13">
<taxonomicName id="4C45E5A90226E121FF03FCA2FE36FC92" box="[199,477,800,824]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380226E121FF03FCA2FE36FC92" box="[199,477,800,824]" italics="true" pageId="13">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430226E121FE21FC9DFDAAFC92" box="[485,577,799,824]" pageId="13" rank="species">
<emphasis id="B93142380226E121FE21FC9DFDAAFC92" bold="true" box="[485,577,799,824]" pageId="13">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90226E121FD1FFCA2FC07FC92" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[731,1004,800,824]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="aequalis">
<emphasis id="B93142380226E121FD1FFCA2FCB7FC92" box="[731,860,800,824]" italics="true" pageId="13">C. aequalis</emphasis>
Bauer 2003
</taxonomicName>
, in lacking femoral pores (
<emphasis id="B93142380226E121FAE5FCA3FA83FC92" box="[1313,1384,801,824]" italics="true" pageId="13">versus</emphasis>
34 minute femoral pores) and possesing more ventral scales (36
<emphasis id="B93142380226E121FC85FCC7FC62FCF6" box="[833,905,837,860]" italics="true" pageId="13">versus</emphasis>
24); from
<taxonomicName id="4C45E5A90226E121FBC4FCC6FAC3FCF6" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[1024,1320,836,860]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="B93142380226E121FBC4FCC6FB75FCF6" box="[1024,1182,836,860]" italics="true" pageId="13">C. annandalei</emphasis>
Bauer 2003
</taxonomicName>
, in having less precloacal pores (5
<emphasis id="B93142380226E121FE6DFCEBFE1AFC2A" box="[425,497,873,896]" italics="true" pageId="13">versus</emphasis>
1112), lacking femoral pores (
<emphasis id="B93142380226E121FC9BFCEBFC4CFC2A" box="[863,935,873,896]" italics="true" pageId="13">versus</emphasis>
1011), and having a dorsal pattern of faint and irregular pale yellowish bands (
<emphasis id="B93142380226E121FD80FC0FFD67FC0E" box="[580,652,909,932]" italics="true" pageId="13">versus</emphasis>
six dark bands); from
<taxonomicName id="4C45E5A90226E121FC60FC0EFB0CFC0E" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[932,1255,908,932]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="chrysopylos">
<emphasis id="B93142380226E121FC60FC0EFBA4FC0E" box="[932,1103,908,932]" italics="true" pageId="13">C. chrysopylos</emphasis>
Bauer 2003
</taxonomicName>
, in having less precloacal pores (5
<emphasis id="B93142380226E121FEBAFC33FE2EFC62" box="[382,453,945,968]" italics="true" pageId="13">versus</emphasis>
10 and a single, pored scale posterior to precloacal series); from
<taxonomicName id="4C45E5A90226E121FB08FC32FECDFC47" authority="Annandale" authorityName="Annandale" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="consobrinoides">
<emphasis id="B93142380226E121FB08FC32FA77FC62" box="[1228,1436,944,968]" italics="true" pageId="13">C. consobrinoides</emphasis>
(Annandale)
</taxonomicName>
, in having more ventral scales (36
<emphasis id="B93142380226E121FD02FC57FCE6FC46" box="[710,781,981,1004]" italics="true" pageId="13">versus</emphasis>
2430), lacking a nuchal band, and possessing a dorsal pattern of faint and irregular pale yellowish bands (
<emphasis id="B93142380226E121FD30FC7BFCD0FBBA" box="[756,827,1017,1040]" italics="true" pageId="13">versus</emphasis>
67 narrow dark cross-bars or transverse markings,
<bibRefCitation id="EFD4E3DB0226E121FF53FB9EFEF7FB9F" author="Smith" box="[151,284,1052,1077]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
); from
<taxonomicName id="4C45E5A90226E121FEA9FB9EFD60FB9E" authority="Boulenger 1893" authorityName="Boulenger" authorityYear="1893" box="[365,651,1052,1077]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="feae">
<emphasis id="B93142380226E121FEA9FB9EFE56FB9E" box="[365,445,1052,1076]" italics="true" pageId="13">C. feae</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FE0FFB9EFD68FB9E" author="Boulenger" box="[459,643,1052,1077]" pageId="13" refString="Boulenger, G. A. (1893) Concluding report on the reptiles and batrachians obtained in Burma by Signor L. Fea, dealing with a collection made in Pegu and Karin Hills in 1887 - 1888. Annali del Museo civico di storia naturale di Genova, 2, 303 - 347." type="journal article" year="1893">Boulenger 1893</bibRefCitation>
)
</taxonomicName>
, in lacking femoral pores (Bauer 2003), and having a dorsal pattern of faint and irregular pale yellowish bands (
<emphasis id="B93142380226E121FDB2FBC3FD56FBF2" box="[630,701,1089,1112]" italics="true" pageId="13">versus</emphasis>
four dark cross-bars,
<bibRefCitation id="EFD4E3DB0226E121FC7AFBBDFBACFBF2" author="Smith" box="[958,1095,1087,1112]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
); from
<taxonomicName id="4C45E5A90226E121FB5BFBC2FF31FBD7" authority="Theoblad 1876" authorityName="Theoblad" authorityYear="1876" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="oldhami">
<emphasis id="B93142380226E121FB5BFBC2FAF4FBF2" box="[1183,1311,1088,1112]" italics="true" pageId="13">C. oldhami</emphasis>
(Theoblad 1876)
</taxonomicName>
, in having more precloacal pores (5
<emphasis id="B93142380226E121FDBDFBE7FD2BFBD6" box="[633,704,1125,1148]" italics="true" pageId="13">versus</emphasis>
04), absence of a nuchal band, and a having a dorsal pattern of faint and irregular pale yellowish bands
<emphasis id="B93142380226E121FD9FFB0BFD49FB0A" box="[603,674,1161,1184]" italics="true" pageId="13">versus</emphasis>
white spots) (
<bibRefCitation id="EFD4E3DB0226E121FC85FB05FC22FB0A" author="Smith" box="[833,969,1159,1184]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
); from
<taxonomicName id="4C45E5A90226E121FBDFFB0AFA91FB0A" authority="Boulenger 1893" authorityName="Boulenger" authorityYear="1893" box="[1051,1402,1159,1184]" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B93142380226E121FBDFFB0AFB41FB35" box="[1051,1194,1160,1183]" italics="true" pageId="13">C. peguensis</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FB7DFB05FA9AFB0A" author="Boulenger" box="[1209,1393,1159,1184]" pageId="13" refString="Boulenger, G. A. (1893) Concluding report on the reptiles and batrachians obtained in Burma by Signor L. Fea, dealing with a collection made in Pegu and Karin Hills in 1887 - 1888. Annali del Museo civico di storia naturale di Genova, 2, 303 - 347." type="journal article" year="1893">Boulenger 1893</bibRefCitation>
)
</taxonomicName>
, in having smaller number of precloacal pores (5
<emphasis id="B93142380226E121FD79FB2FFCEEFB6E" box="[701,773,1197,1220]" italics="true" pageId="13">versus</emphasis>
79) and continuous series of precloacal-femoral scales (
<bibRefCitation id="EFD4E3DB0226E121FF5AFB4DFECCFB42" author="Smith" box="[158,295,1231,1256]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
;
<bibRefCitation id="EFD4E3DB0226E121FEF1FB4DFE29FB42" author="Taylor" box="[309,450,1231,1256]" pageId="13" refString="Taylor, E. H. (1963) The lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
); from
<taxonomicName id="4C45E5A90226E121FDD3FB52FCF2FB42" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[535,793,1232,1256]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="russelli">
<emphasis id="B93142380226E121FDD3FB52FD67FB42" box="[535,652,1232,1256]" italics="true" pageId="13">C. russelli</emphasis>
Bauer 2003
</taxonomicName>
and
<taxonomicName id="4C45E5A90226E121FC97FB52FB9EFB42" authority="Bauer 2003" authorityName="Bauer" authorityYear="2003" box="[851,1141,1232,1256]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="slowinskii">
<emphasis id="B93142380226E121FC97FB52FC0DFB42" box="[851,998,1232,1256]" italics="true" pageId="13">C. slowinskii</emphasis>
Bauer 2003
</taxonomicName>
, in having less precloacal pores (5
<emphasis id="B93142380226E121FEC5FB77FEA3FAA6" box="[257,328,1269,1292]" italics="true" pageId="13">versus</emphasis>
15 and 9, respectively) and lacking femoral pores (
<emphasis id="B93142380226E121FC74FB77FC13FAA6" box="[944,1016,1269,1292]" italics="true" pageId="13">versus</emphasis>
1619 and 11 pores on each thigh, respectively); and from
<taxonomicName id="4C45E5A90226E121FE74FA9AFD06FA9A" authority="Blyth 1859" authorityName="Blyth" authorityYear="1859" box="[432,749,1303,1328]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="variegatus">
<emphasis id="B93142380226E121FE74FA9AFDA0FA85" box="[432,587,1304,1327]" italics="true" pageId="13">C. variegatus</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FD9AFA95FD0EFA9A" author="Blyth" box="[606,741,1303,1328]" pageId="13" refString="Blyth, E. (1859) Of reptiles, Acanthosaura armata, Gray; and a beautiful gecko from the interior. Journal of the Asiatic Society of Bengal, 28, 279." type="journal article" year="1859">Blyth 1859</bibRefCitation>
)
</taxonomicName>
, in having more ventral scales (36
<emphasis id="B93142380226E121FB56FA9BFB31FA9A" box="[1170,1242,1305,1328]" italics="true" pageId="13">versus</emphasis>
22) and lacking femoral pores (
<emphasis id="B93142380226E121FE86FABFFE62FAFE" box="[322,393,1341,1364]" italics="true" pageId="13">versus</emphasis>
32 continuous precloacal-femoral pores (
<bibRefCitation id="EFD4E3DB0226E121FC9FFABEFC09FAFF" author="Smith" box="[859,994,1340,1365]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
).
</paragraph>
<paragraph id="8BFA9E2A0226E122FF03FAE2FAE8FECF" blockId="13.[151,1436,151,2015]" lastBlockId="14.[151,1436,151,752]" lastPageId="14" pageId="13">
For
<collectingCountry id="F352DEBA0226E121FF3DFADDFEC7FAD2" box="[249,300,1375,1400]" name="Thailand" pageId="13">Thai</collectingCountry>
congeners,
<taxonomicName id="4C45E5A90226E121FE78FAE2FD3EFAD2" box="[444,725,1376,1400]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380226E121FE78FAE2FD3EFAD2" box="[444,725,1376,1400]" italics="true" pageId="13">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430226E121FD24FADDFCD4FAD2" box="[736,831,1375,1400]" pageId="13" rank="species">
<emphasis id="B93142380226E121FD24FADDFCD4FAD2" bold="true" box="[736,831,1375,1400]" pageId="13">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C45E5A90226E121FC24FAE2FAA8FAD2" authority="Taylor 1962" authorityName="Taylor" authorityYear="1962" box="[992,1347,1375,1400]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="B93142380226E121FC24FAE2FB40FAD2" box="[992,1195,1376,1400]" italics="true" pageId="13">C. quadrivirgatus</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FB72FADDFAA8FAD2" author="Taylor" box="[1206,1347,1375,1400]" pageId="13" refString="Taylor, E. H. (1962) New oriental reptiles. The University of Kansas Science Bulletin. 43, 209 - 263." type="journal article" year="1962">Taylor 1962</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C45E5A90226E121FA46FAE2FE47FA37" authority="(Smith 1921b)" baseAuthorityName="Smith" baseAuthorityYear="1921" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B93142380226E121FA46FAE2FEE8FA36" italics="true" pageId="13">C. angularis</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FED4FA06FE4FFA37" author="Smith" box="[272,420,1412,1437]" pageId="13" refString="Smith, M. A. (1921 b) New or little-known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 91 (2), 423 - 440. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1921. tb 03271. x" type="journal article" year="1921" yearSuffix="b">Smith 1921b</bibRefCitation>
)
</taxonomicName>
, in having moderately enlarged subcaudal scales (
<emphasis id="B93142380226E121FC18FA07FBC8FA36" box="[988,1059,1413,1436]" italics="true" pageId="13">versus</emphasis>
small;
<bibRefCitation id="EFD4E3DB0226E121FBB0FA06FB12FA37" author="Smith" box="[1140,1273,1412,1437]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
;
<bibRefCitation id="EFD4E3DB0226E121FAC1FA06FA7BFA37" author="Taylor" box="[1285,1424,1412,1437]" pageId="13" refString="Taylor, E. H. (1963) The lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
). The new
<taxonomicName id="4C45E5A90226E121FEC4FA25FEA8FA6A" box="[256,323,1447,1472]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
differs from
<taxonomicName id="4C45E5A90226E121FE1CFA2AFA73FA6A" authority="Sumontha, Pauwels, Kunya, Nitikul, Samphanthamit &amp; Grismer 2012" authorityName="Sumontha, Pauwels, Kunya, Nitikul, Samphanthamit &amp; Grismer" authorityYear="2012" box="[472,1432,1447,1472]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="phuketensis">
<emphasis id="B93142380226E121FE1CFA2AFD97FA6A" box="[472,636,1448,1472]" italics="true" pageId="13">C. phuketensis</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FD47FA25FA73FA6A" author="Sumontha" box="[643,1432,1447,1472]" pageId="13" refString="Sumontha, M., Pauwels, O. S. G., Kunya, K., Nitikul, A., Samphanthamit, P. &amp; Grismer, L. L. (2012) A new forest-dwelling gecko from Phuket Island, southern Thailand, related to Cyrtodactylus macrotunerculatus (Squamata: Gekkonidae). Zootaxa, 3522, 61 - 72." type="journal article" year="2012">Sumontha, Pauwels, Kunya, Nitikul, Samphanthamit &amp; Grismer 2012</bibRefCitation>
</taxonomicName>
, in lacking a precloacal groove and femoral pores (
<emphasis id="B93142380226E121FD1BFA4FFCCDFA4E" box="[735,806,1485,1508]" italics="true" pageId="13">versus</emphasis>
3336 continuous precloacal-femoral pores). The new species differs from the following species in lacking femoral pores:
<taxonomicName id="4C45E5A90226E121FC4AFA72FC53F986" authority="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels 2012" authorityName="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="astrum">
<emphasis id="B93142380226E121FC4AFA72FC17F9A2" box="[910,1020,1520,1544]" italics="true" pageId="13">C. astrum</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FBC7FA6DFC53F986" author="Grismer" pageId="13" refString="Grismer, L. L., Wood, P. L., Quah, E. S. H., Anuar, S., Muin, M. A., Sumontha, M., Ahmad, N., Bauer, A. M., Wangkulangkul, S., Grismer, J. L. &amp; Pauwels, O. S. G. (2012) A phylogeny and taxonomy of the Thai-Malay Peninsula bent-toed geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520, 1 - 55." type="journal article" year="2012">Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FC01F996FEF0F9FA" authority="Sumontha, Panitvong &amp; Deein 2010" authorityName="Sumontha, Panitvong &amp; Deein" authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="auribalteatus">
<emphasis id="B93142380226E121FC01F996FB95F986" box="[965,1150,1556,1580]" italics="true" pageId="13">C. auribalteatus</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FB4CF996FEF0F9FA" author="Sumontha" pageId="13" refString="Sumontha, M., Panitvong, N. &amp; Deein, G. (2010) Cyrtodactylus auribalteatus (Squamata: Gekkonidae), a new cave-dwelling gecko from Phitsanulok Province, Thailand. Zootaxa, 2370, 53 - 64." type="journal article" year="2010">Sumontha, Panitvong &amp; Deein 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FEE2F9BAFD69F9FA" authority="Smith 1923" authorityName="Smith" authorityYear="1923" box="[294,642,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="B93142380226E121FEE2F9BAFE0EF9FA" box="[294,485,1592,1616]" italics="true" pageId="13">C. brevipalmatus</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FE30F9B5FD91F9FA" author="Smith" box="[500,634,1591,1616]" pageId="13" refString="Smith, M. A. (1923) Notes on reptiles and batrachians from Siam and Indo-China (No. 2). Journal of the Natural History Society of Siam, 6, 47 - 53." type="journal article" year="1923">Smith 1923</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FD4AF9BAFB2DF9FA" authority="Bauer, Sumontha &amp; Pauwels 2003" authorityName="Bauer, Sumontha &amp; Pauwels" authorityYear="2003" box="[654,1222,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="chanhomeae">
<emphasis id="B93142380226E121FD4AF9BAFCD0F9FA" box="[654,827,1592,1616]" italics="true" pageId="13">C. chanhomeae</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FC87F9BAFB2DF9FA" author="Bauer" box="[835,1222,1591,1616]" pageId="13" refString="Bauer, A. M., Sumontha, M. &amp; Pauwels, O. S. G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Thailand. Zootaxa, 376, 1 - 18." type="journal article" year="2003">Bauer, Sumontha &amp; Pauwels 2003</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FB14F9BAFC56F9DE" authority="Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome &amp; Kunya 2010" authorityName="Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome &amp; Kunya" authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="dumnuii">
<emphasis id="B93142380226E121FB14F9BAFAA6F9FA" box="[1232,1357,1592,1616]" italics="true" pageId="13">C. dumnuii</emphasis>
Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome &amp; Kunya 2010
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FC0AF9DEFCB8F932" authority="Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome &amp; Kunya 2009" authorityName="Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome &amp; Kunya" authorityYear="2009" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="erythrops">
<emphasis id="B93142380226E121FC0AF9DEFB8BF9DE" box="[974,1120,1628,1652]" italics="true" pageId="13">C. erythrops</emphasis>
Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome &amp;
<bibRefCitation id="EFD4E3DB0226E121FD06F903FCB8F932" author="Kunya" box="[706,851,1664,1689]" pageId="13" refString="Bauer, A. M., Kunya, K., Sumontha, M, Niyomwan, P., Nanitvong, P. &amp; Pauwels, O. S. G. (2009) Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand. Zootaxa, 2124, 51 - 62." type="journal article" year="2009">Kunya 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FCA7F902FC3EF916" authority="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels 2012" authorityName="Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="lekaguli">
<emphasis id="B93142380226E121FCA7F902FC08F932" box="[867,995,1664,1688]" italics="true" pageId="13">C. lekaguli</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FC2AF9FDFC3EF916" author="Grismer" pageId="13" refString="Grismer, L. L., Wood, P. L., Quah, E. S. H., Anuar, S., Muin, M. A., Sumontha, M., Ahmad, N., Bauer, A. M., Wangkulangkul, S., Grismer, J. L. &amp; Pauwels, O. S. G. (2012) A phylogeny and taxonomy of the Thai-Malay Peninsula bent-toed geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520, 1 - 55." type="journal article" year="2012">Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer &amp; Pauwels 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C45E5A90226E121FC23F926FAD1F916" authority="Gray 1831" authorityName="Gray" authorityYear="1831" box="[999,1338,1700,1724]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B93142380226E121FC23F926FB73F916" box="[999,1176,1700,1724]" italics="true" pageId="13">C. marmoratus</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FB6AF926FADBF916" author="Gray" box="[1198,1328,1700,1724]" pageId="13" refString="Gray, J. E. (1831) A synopsis of the species of class Reptilia. In: Griffith, E. &amp; Pidgeon, E. (Eds.), The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed. Whittaker, Treacher and Co., London, pp. 1 - 481 + 1 - 110." type="book chapter" year="1831">Gray 1831</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C45E5A90226E121FA46F926FD7BF94A" authority="Bauer, Sumontha &amp; Pauwels 2003" authorityName="Bauer, Sumontha &amp; Pauwels" authorityYear="2003" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="trigroides">
<emphasis id="B93142380226E121FA46F926FEEFF94A" italics="true" pageId="13">C. trigroides</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FEC8F94BFD7BF94A" author="Bauer" box="[268,656,1735,1760]" pageId="13" refString="Bauer, A. M., Sumontha, M. &amp; Pauwels, O. S. G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Thailand. Zootaxa, 376, 1 - 18." type="journal article" year="2003">Bauer, Sumontha &amp; Pauwels 2003</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFD4E3DB0226E121FD64F945FCCCF94A" author="Smith" box="[672,807,1735,1760]" pageId="13" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
;
<bibRefCitation id="EFD4E3DB0226E121FCF0F945FC2BF94A" author="Taylor" box="[820,960,1735,1760]" pageId="13" refString="Taylor, E. H. (1963) The lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
).
<taxonomicName id="4C45E5A90226E121FC10F94AFB02F94A" box="[980,1257,1736,1760]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380226E121FC10F94AFB02F94A" box="[980,1257,1736,1760]" italics="true" pageId="13">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430226E121FB35F945FAA0F94A" box="[1265,1355,1735,1760]" pageId="13" rank="species">
<emphasis id="B93142380226E121FB35F945FAA0F94A" bold="true" box="[1265,1355,1735,1760]" pageId="13">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C45E5A90226E121FEB4F96EFCB9F8AE" authority="Ulber &amp; Grossmann 1991" authorityName="Ulber &amp; Grossmann" authorityYear="1991" box="[368,850,1772,1797]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="papilionoides">
<emphasis id="B93142380226E121FEB4F96EFDC2F8AE" box="[368,553,1772,1796]" italics="true" pageId="13">C. papilionoides</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FDEBF96EFCB9F8AE" author="Ulber" box="[559,850,1772,1797]" pageId="13" refString="Ulber, T. &amp; Grossmann, W. (1991) Ein weiterer neuer Gecko aus Zentral-Thailand: Cyrtodactylus papilionoides sp. nov. (Reptilia: Sauria: Gekkonidae). Sauria, 13, 13 - 22." type="journal article" year="1991">Ulber &amp; Grossmann 1991</bibRefCitation>
</taxonomicName>
, in having a dorsal pattern of faint and irregular pale yellowish bands (
<emphasis id="B93142380226E121FE98F893FE4FF882" box="[348,420,1809,1832]" italics="true" pageId="13">versus</emphasis>
serrated, broad, dark bands with pale margins or transversal rows of blotches), and having a continuous series of 36 digital lamellae under the 4th toes (
<emphasis id="B93142380226E121FCB5F8BAFC53F8E5" box="[881,952,1848,1871]" italics="true" pageId="13">versus</emphasis>
an interrupted series of 1216 lamellae; Heidrick
<emphasis id="B93142380226E121FEC6F8D9FED7F8D9" box="[258,316,1883,1907]" italics="true" pageId="13">et al.</emphasis>
2007; Nguyen
<emphasis id="B93142380226E121FE2AF8D9FDCCF8D9" box="[494,551,1883,1907]" italics="true" pageId="13">et al.</emphasis>
2010); from
<taxonomicName id="4C45E5A90226E121FD05F8D9FBFBF8D9" authority="Peters 1871" authorityName="Peters" authorityYear="1871" box="[705,1040,1883,1907]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="B93142380226E121FD05F8D9FC86F8D9" box="[705,877,1883,1907]" italics="true" pageId="13">C. consobrinus</emphasis>
(
<bibRefCitation id="EFD4E3DB0226E121FCB9F8D9FBECF8D9" author="Peters" box="[893,1031,1883,1907]" pageId="13" refString="Peters, W. (1871) Uber neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzuglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua. Monatsbericht der Koniglich Preußische Akademie der Wissenschaften zu Berlin, 1871, 566 - 581." type="journal article" year="1871">Peters 1871</bibRefCitation>
)
</taxonomicName>
, in having less precloacal pores (5
<emphasis id="B93142380226E121FF53F802FF34F83D" box="[151,223,1920,1943]" italics="true" pageId="13">versus</emphasis>
911) and a dorsal pattern of faint and irregular pale yellowish bands (
<emphasis id="B93142380226E121FC32F802FBD5F83D" box="[1014,1086,1920,1943]" italics="true" pageId="13">versus</emphasis>
8 dark, transverse cross-bands,
<bibRefCitation id="EFD4E3DB0226E121FF53F820FEB8F811" author="Boulenger" box="[151,339,1954,1979]" pageId="13" refString="Boulenger, G. A. (1885) Catalogue of the Lizard in the British Museum (Natural History). Vol. 1. Trustees, London, 512 pp." type="book" year="1885">Boulenger 1885</bibRefCitation>
); from
<taxonomicName id="4C45E5A90226E121FE6BF821FBA1F811" authority="Pauwels, Sumontha, Latinne &amp; Grismer 2013" authorityName="Pauwels, Sumontha, Latinne &amp; Grismer" authorityYear="2013" box="[431,1098,1954,1979]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="sanook">
<emphasis id="B93142380226E121FE6BF821FDC8F811" box="[431,547,1955,1979]" italics="true" pageId="13">C. sanook</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FDF4F820FBA1F811" author="Pauwels" box="[560,1098,1954,1979]" pageId="13" refString="Pauwels, O. S. G., Sumontha, M., Latinne, A. &amp; Grismer, L. L. (2013) Cyrtodactylus sanook (Squamata: Gekkonidae), a new cave-dwelling gecko from Chumphon Province, southern Thailand. Zootaxa, 3635 (3), 275 - 285. http: // dx. doi. org / 10.11646 / zootaxa. 3635.3.7" type="journal article" year="2013">Pauwels, Sumontha, Latinne &amp; Grismer 2013</bibRefCitation>
</taxonomicName>
, in having medium enlarged femoral scale rows (
<emphasis id="B93142380226E121FE42F84AFE26F875" box="[390,461,1992,2015]" italics="true" pageId="13">versus</emphasis>
well-developed); from
<taxonomicName id="4C45E5A90226E121FD26F845FACBF875" authority="Bauer, Pauwels &amp; Chanhome 2002" authorityName="Bauer, Pauwels &amp; Chanhome" authorityYear="2002" box="[738,1312,1990,2015]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="sumonthai">
<emphasis id="B93142380226E121FD26F845FC90F875" box="[738,891,1991,2015]" italics="true" pageId="13">C. sumonthai</emphasis>
<bibRefCitation id="EFD4E3DB0226E121FC41F84AFACBF875" author="Bauer" box="[901,1312,1990,2015]" pageId="13" refString="Bauer, A. M., Pauwels, O. S. G. &amp; Chanhome, L. (2002) A new species of cave-dwelling Cyrtodactylus (Squamata: Gekkonidae) from Thailand. The Natural History Journal of Chulalongkorn University, 2, 19 - 29." type="journal article" year="2002">Bauer, Pauwels &amp; Chanhome 2002</bibRefCitation>
</taxonomicName>
, in having more precloacal pores (5
<emphasis id="B93142380225E122FE78FF1BFDE8FF1A" box="[444,515,153,176]" italics="true" pageId="14">versus</emphasis>
2 minute pores), and a series of moderately enlarged femoral scales (
<emphasis id="B93142380225E122FAEDFF1BFA9BFF1A" box="[1321,1392,153,176]" italics="true" pageId="14">versus</emphasis>
the absence of enlarged femoral scales); from
<taxonomicName id="4C45E5A90225E122FDBFFF3EFD3CFF7E" box="[635,727,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="surin">
<emphasis id="B93142380225E122FDBFFF3EFD3CFF7E" box="[635,727,188,212]" italics="true" pageId="14">C. surin</emphasis>
</taxonomicName>
<bibRefCitation id="EFD4E3DB0225E122FD24FF3EFBF9FF7E" author="Chan-ard" box="[736,1042,188,213]" pageId="14" refString="Chan-ard, T. &amp; Makchai, S. (2011) A new insular species of Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae), from the Surin Islands, Phang-nga Province, southern Thailand. The Thailand Natural History Museum Journal, 5, 7 - 15." type="journal article" year="2011">Chan-ard &amp; Makchai 2011</bibRefCitation>
, in having more ventral scales (36
<emphasis id="B93142380225E122FF53FF63FF34FF52" box="[151,223,225,248]" italics="true" pageId="14">versus</emphasis>
25) and smooth or slightly keeled dorsal tubercles (
<emphasis id="B93142380225E122FCFDFF63FC6AFF52" box="[825,897,225,248]" italics="true" pageId="14">versus</emphasis>
very large, trihedral, and keeled); and from
<taxonomicName id="4C45E5A90225E122FA46FF62FCDDFEB6" authority="Pauwels, Bauer, Sumontha &amp; Chanhome 2004" authorityName="Pauwels, Bauer, Sumontha &amp; Chanhome" authorityYear="2004" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="thirakhupti">
<emphasis id="B93142380225E122FA46FF62FEFFFEB6" italics="true" pageId="14">C. thirakhupti</emphasis>
<bibRefCitation id="EFD4E3DB0225E122FED9FE86FCDDFEB6" author="Pauwels" box="[285,822,260,285]" pageId="14" refString="Pauwels, O. S. G., Bauer, A. M., Sumontha, M. &amp; Chanhome, L. (2004) Cyrtodactylus thirakhupti (Squamata: Gekkonidae), a new cave-dwelling gecko from southern Thailand. Zootaxa, 772, 1 - 11." type="journal article" year="2004">Pauwels, Bauer, Sumontha &amp; Chanhome 2004</bibRefCitation>
</taxonomicName>
, in having fewer precloacal pores (5
<emphasis id="B93142380225E122FB21FE87FAC7FEB6" box="[1253,1324,261,284]" italics="true" pageId="14">versus</emphasis>
89) and lacking femoral pores (
<emphasis id="B93142380225E122FE5BFEABFE0DFEEA" box="[415,486,297,320]" italics="true" pageId="14">versus</emphasis>
57). Other
<collectingCountry id="F352DEBA0225E122FDBDFEA5FD47FEEA" box="[633,684,295,320]" name="Thailand" pageId="14">Thai</collectingCountry>
species, including
<taxonomicName id="4C45E5A90225E122FC43FEAAFBD3FEEA" box="[903,1080,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="B93142380225E122FC43FEAAFBD3FEEA" box="[903,1080,296,320]" italics="true" pageId="14">C. interdigitalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90225E122FB8CFEAAFB05FEEA" box="[1096,1262,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93142380225E122FB8CFEAAFB05FEEA" box="[1096,1262,296,320]" italics="true" pageId="14">C. intermedius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90225E122FB39FEAAFA98FE95" box="[1277,1395,296,319]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="jarujini">
<emphasis id="B93142380225E122FB39FEAAFA98FE95" box="[1277,1395,296,319]" italics="true" pageId="14">C. jarujini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90225E122FA46FEAAFF18FECE" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="oldhami">
<emphasis id="B93142380225E122FA46FEAAFF18FECE" italics="true" pageId="14">C. oldhami</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90225E122FEC5FECEFE64FECE" box="[257,399,332,356]" class="Reptilia" family="Phyllodactylidae" genus="Gymnodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B93142380225E122FEC5FECEFE64FECE" box="[257,399,332,356]" italics="true" pageId="14">C. peguensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C45E5A90225E122FE59FECEFDDAFECE" box="[413,561,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="B93142380225E122FE59FECEFDDAFECE" box="[413,561,332,356]" italics="true" pageId="14">C. pulchellus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C45E5A90225E122FDABFECEFCEEFECE" box="[623,773,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="variegatus">
<emphasis id="B93142380225E122FDABFECEFCEEFECE" box="[623,773,332,356]" italics="true" pageId="14">C. variegatus</emphasis>
</taxonomicName>
, were compared to the new species as above.
</paragraph>
<paragraph id="8BFA9E2A0225E122FF03FEF2FBD7FE5F" blockId="14.[151,1436,151,752]" pageId="14">
Two described congeners occur in
<collectingCountry id="F352DEBA0225E122FDACFEEDFD0BFE22" box="[616,736,367,392]" name="Cambodia" pageId="14">Cambodia</collectingCountry>
:
<taxonomicName id="4C45E5A90225E122FD34FEF2FC70FE22" box="[752,923,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B93142380225E122FD34FEF2FC70FE22" box="[752,923,368,392]" italics="true" pageId="14">C. intermedius</emphasis>
</taxonomicName>
(Grismer
<emphasis id="B93142380225E122FBDDFEF3FBBCFE22" box="[1049,1111,368,392]" italics="true" pageId="14">et al.</emphasis>
2008;
<bibRefCitation id="EFD4E3DB0225E122FB74FEEDFAD3FE22" author="Smith" box="[1200,1336,367,392]" pageId="14" refString="Smith, M. A. (1935) The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Vol. II-Sauria. Taylor and Francis Ltd., London, 541 pp." type="book" year="1935">Smith 1935</bibRefCitation>
) and
<taxonomicName id="4C45E5A90225E122FA46FEF2FD93FE06" authority="Stuart et al. 2010" authorityName="Stuart et al." authorityYear="2010" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="pseudoquadrivirgatus">
<emphasis id="B93142380225E122FA46FEF2FE67FE06" italics="true" pageId="14">C. pseudoquadrivirgatus</emphasis>
(
<bibRefCitation id="EFD4E3DB0225E122FE5AFE16FD9AFE06" author="Stuart" box="[414,625,404,428]" pageId="14" refString="Stuart, B. L., Rowley, J. J. L., Thy, N., Emmett, D. A. &amp; Sitha, S. (2010) Significant new records of amphibians and reptiles from Virachey National Park, northeastern Cambodia. Cambodian Journal of Natural History, 2010, 38 - 47." type="journal article" year="2010">
Stuart
<emphasis id="B93142380225E122FE29FE17FDC9FE06" box="[493,546,404,428]" italics="true" pageId="14">et al</emphasis>
. 2010
</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C45E5A90225E122FD42FE16FC3DFE06" box="[646,982,404,428]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="bugiamapensis">
<emphasis id="B93142380225E122FD42FE16FC3DFE06" box="[646,982,404,428]" italics="true" pageId="14">Cyrtodactylus bugiamapensis</emphasis>
</taxonomicName>
should occur in
<collectingCountry id="F352DEBA0225E122FB64FE16FAF8FE07" box="[1184,1299,404,429]" name="Cambodia" pageId="14">Cambodia</collectingCountry>
because its habitat extends continuously from the
<typeStatus id="54FE20880225E122FDA7FE3AFD78FE7A" box="[611,659,440,464]" pageId="14">type</typeStatus>
locality to the border between
<collectingCountry id="F352DEBA0225E122FBD0FE35FB98FE7A" box="[1044,1139,439,464]" name="Vietnam" pageId="14">Vietnam</collectingCountry>
and
<collectingCountry id="F352DEBA0225E122FB71FE35FAC3FE7A" box="[1205,1320,439,464]" name="Cambodia" pageId="14">Cambodia</collectingCountry>
(see also
<bibRefCitation id="EFD4E3DB0225E122FF53FE5FFE92FE5E" author="Nazarov" box="[151,377,476,500]" pageId="14" refString="Nazarov, R., Poyarkov, N. A., Orlov, N. L., Phung, T. M., Nguyen, T. T., Hoang, D. M. &amp; Ziegler, T. (2012) Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 3302, 1 - 24." type="journal article" year="2012">
Nazarov
<emphasis id="B93142380225E122FF38FE5FFEDEFE5E" box="[252,309,476,500]" italics="true" pageId="14">et al.</emphasis>
2012
</bibRefCitation>
). These three species were compared above to the new species.
</paragraph>
</subSubSection>
<subSubSection id="C35FCDA10225E122FF03FE7DFE49FD97" pageId="14" type="etymology">
<paragraph id="8BFA9E2A0225E122FF03FE7DFE49FD97" blockId="14.[151,1436,151,752]" pageId="14">
<emphasis id="B93142380225E122FF03FE7DFEADFDB2" bold="true" box="[199,326,511,536]" pageId="14">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C45E5A90225E122FDFBFD82FD44FDB2" box="[575,687,512,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="puhuensis">
<emphasis id="B93142380225E122FDFBFD82FD44FDB2" box="[575,687,512,536]" italics="true" pageId="14">puhuensis</emphasis>
</taxonomicName>
refers to the
<typeStatus id="54FE20880225E122FC8EFD82FC91FDB2" box="[842,890,512,536]" pageId="14">type</typeStatus>
locality, Pù Hu Nature Reserve, where the new species was discovered.
</paragraph>
</subSubSection>
<subSubSection id="C35FCDA10225E122FF03FDC5FF0EFD45" pageId="14" type="distribution">
<paragraph id="8BFA9E2A0225E122FF03FDC5FF0EFD45" blockId="14.[151,1436,151,752]" pageId="14">
<emphasis id="B93142380225E122FF03FDC5FE05FDCA" bold="true" box="[199,494,583,608]" pageId="14">Distribution and Habitat</emphasis>
.
<taxonomicName id="4C45E5A90225E122FE38FDCAFCFBFDCA" box="[508,784,584,608]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="puhuensis" status="sp. nov.">
<emphasis id="B93142380225E122FE38FDCAFCFBFDCA" box="[508,784,584,608]" italics="true" pageId="14">Cyrtodactylus puhuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202FF430225E122FCD2FDC5FC9BFDCA" box="[790,880,583,608]" pageId="14" rank="species">
<emphasis id="B93142380225E122FCD2FDC5FC9BFDCA" bold="true" box="[790,880,583,608]" pageId="14">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the Pù Hu Nature Reserve, in northwestern Thanh Hóa Province, northern
<collectingCountry id="F352DEBA0225E122FD54FDEEFD04FD2F" box="[656,751,620,645]" name="Vietnam" pageId="14">Vietnam</collectingCountry>
(
<figureCitation id="137E82AF0225E122FD3BFDEEFCAEFD2E" box="[767,837,620,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1716,1738]" captionTargetBox="[151,1436,303,1694]" captionTargetId="figure@4.[151,1436,303,1694]" captionTargetPageId="4" captionText="FIGURE 2. Sampling localities of Cyrtodactylus in Vietnam and adjacent areas included in this study (circles). Red circle shows the location the new species described herein, and blue circles show locations of sympatry. Triangles represent remaining Vietnamese congeners. Numbers for each locality refer to Table 1." httpUri="https://zenodo.org/record/230343/files/figure.png" pageId="14">Fig. 2</figureCitation>
). The single known specimen of this new
<taxonomicName id="4C45E5A90225E122FAE1FDEEFA83FD2F" box="[1317,1384,620,645]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
was found at night on a large tree (diameter about
<quantity id="4CBD33CF0225E122FD7AFD0DFCEEFD02" box="[702,773,655,680]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" unit="m" value="1.5">1.5 m</quantity>
), at about
<quantity id="4CBD33CF0225E122FC42FD0DFC21FD0D" box="[902,970,655,680]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="14" unit="m" value="2.5">2.5 m</quantity>
above the forest floor. The habitat was evergreen forest intermixed with bamboo. Limestone was not observed in the area and the recorded elevation was
<specimenCount id="9D4355A30225E122FF53FD55FF0EFD45" box="[151,229,727,752]" pageId="14" type="generic">
<quantity id="4CBD33CF0225E122FF53FD55FF0AFD45" box="[151,225,727,752]" metricMagnitude="2" metricUnit="m" metricValue="6.38" pageId="14" unit="m" value="638.0">638 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>