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<document id="FB6B4F1045DE198D1248D6CE25B43A1D" ID-DOI="10.1080/00222933.2024.2388791" ID-ISSN="1464-5262" ID-Zenodo-Dep="14241324" ID-ZooBank="18D05DD2- 4B64-4A87-8389-582D5714411C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732836998606" checkinUser="plazi" docAuthor="Sharaf, Mostafa R., Wetterer, James K., Mohamed, AbdulAziz M. A., Georgiadis, Christos, Nasser, Mohamed G. &amp; Aldawood, Abdulrahman S." docDate="2024" docId="03D9FD3BFFB8FF8BFE2DF9B2AE64FCF0" docLanguage="en" docName="JNATHIST.58.41-44.1705-1786.pdf" docOrigin="Journal of Natural History 58 (41 - 44)" docSource="http://dx.doi.org/10.1080/00222933.2024.2388791" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Messor ebeninus Santschi 1927" docType="treatment" docVersion="1" lastPageNumber="1744" masterDocId="FFE08543FF9BFFACFFE7FFABAD31FF9B" masterDocTitle="Filling gaps in global myrmecology: ants of the Kingdom of Bahrain (Hymenoptera: Formicidae)" masterLastPageNumber="1786" masterPageNumber="1705" pageNumber="1740" updateTime="1733158646059" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="C6DAB86A7C28B3DA2AC08454F4298717">Filling gaps in global myrmecology: ants of the Kingdom of Bahrain (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="79B34EFC943A17F0C6958305A66E960D">Sharaf, Mostafa R.</mods:namePart>
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<mods:namePart id="66C028B0283D9E17124214F2EE08633A">Wetterer, James K.</mods:namePart>
<mods:affiliation id="833846BB5208B78A881E9D3E894CC610">Wilkes Honors College, Florida Atlantic University, Jupiter, FL, USA;</mods:affiliation>
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<mods:namePart id="2515828939A701F2DA7ED8F98623DEA9">Mohamed, AbdulAziz M. A.</mods:namePart>
<mods:affiliation id="C269222628C97883964272CDF781AA95">Agriculture Affairs, Ministry of Municipalities Affairs and Agriculture, Kingdom of Bahrain;</mods:affiliation>
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<mods:namePart id="44D796FBD9C0C397DB731FBD03C9B16A">Georgiadis, Christos</mods:namePart>
<mods:affiliation id="40B54435BB9C21E10DFC2B38F3978E03">Section of Zoology-Marine Biology and Zoological Museum, Department of Biology, National and Kapodistrian University of Athens, Athens, Greece;</mods:affiliation>
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<mods:namePart id="82E210E7CF8F44EB188C1D78D5BFBCC7">Nasser, Mohamed G.</mods:namePart>
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<mods:title id="DDD5F969EA44BAE365F429ABAB823BC2">Journal of Natural History</mods:title>
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<mods:date id="E4FB5689301C34D215B37604157F98B5">2024</mods:date>
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<mods:number id="6B6A1224642185D4BE123480FCE5E735">2024-09-16</mods:number>
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<subSubSection id="C36A1FA6FFB8FF8FFE2DF9B2AE68F9AF" box="[458,857,1561,1588]" pageId="35" pageNumber="1740" type="nomenclature">
<paragraph id="8BCF4C2DFFB8FF8FFE2DF9B2AE68F9AF" blockId="35.[458,857,1561,1588]" box="[458,857,1561,1588]" pageId="35" pageNumber="1740">
<heading id="D087FB41FFB8FF8FFE2DF9B2AE68F9AF" box="[458,857,1561,1588]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="1740" reason="5">
<taxonomicName id="4C7037AEFFB8FF8FFE2DF9B2AE68F9AF" ID-CoL="427ZS" authority="Santschi, 1927" authorityName="Santschi" authorityYear="1927" box="[458,857,1561,1588]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="1740" phylum="Arthropoda" rank="species" species="ebeninus">
<emphasis id="B904903FFFB8FF8FFE2DF9B2AF91F9AF" bold="true" box="[458,672,1561,1588]" italics="true" pageId="35" pageNumber="1740">Messor ebeninus</emphasis>
<bibRefCitation id="EFE131DCFFB8FF8FFD4FF9B2AE68F9AF" author="Santschi F" box="[680,857,1561,1588]" pageId="35" pageNumber="1740" pagination="225 - 250" refId="ref23074" refString="Santschi F. 1927. Revision des Messor du groupe instabilis Sm. (Hymenopt.). Bol R Soc Esp Hist Nat. 27: 225 - 250." type="journal article" year="1927">Santschi, 1927</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36A1FA6FFB8FF8FFDFCF9F7AE39F9EC" box="[539,776,1628,1655]" pageId="35" pageNumber="1740" type="description">
<paragraph id="8BCF4C2DFFB8FF8FFDFCF9F7AE39F9EC" blockId="35.[539,776,1628,1655]" box="[539,776,1628,1655]" pageId="35" pageNumber="1740">
(
<figureCitation id="134B50A8FFB8FF8FFDC4F9F7AF92F9EC" box="[547,675,1628,1655]" captionStart="Figure 23" captionStartId="37.[160,225,1754,1777]" captionTargetBox="[394,921,144,1719]" captionTargetId="figure-3@37.[392,924,142,1722]" captionTargetPageId="37" captionText="Figure 23. Messor ebeninus. Worker. (A) Body in profile. (B) Head in full-face view, ANTWEB1041905, AntWeb.org (Michele Esposito). (C) Distribution map." figureDoi="http://doi.org/10.5281/zenodo.14241371" httpUri="https://zenodo.org/record/14241371/files/figure.png" pageId="35" pageNumber="1740">Figures 23</figureCitation>
and
<figureCitation id="134B50A8FFB8FF8FFD05F9F7AE31F9EC" box="[738,768,1628,1655]" captionStart="Figure 24" captionStartId="38.[160,225,1754,1777]" captionTargetBox="[329,986,145,1719]" captionTargetId="figure-3@38.[326,989,142,1722]" captionTargetPageId="38" captionText="Figure 24. Messor ebeninus. Queen. (A) Body in profile. (B) Body in dorsal view. (C) Head in full-face view, ANTWEB1041906, AntWeb.org (Michele Esposito)." figureDoi="http://doi.org/10.5281/zenodo.14241373" httpUri="https://zenodo.org/record/14241373/files/figure.png" pageId="35" pageNumber="1740">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FFB8FF8FFF27F90CA934F95B" box="[192,1029,1703,1728]" pageId="35" pageNumber="1740" type="reference_group">
<paragraph id="8BCF4C2DFFB8FF8FFF27F90CA934F95B" blockId="35.[192,1029,1703,1728]" box="[192,1029,1703,1728]" pageId="35" pageNumber="1740">
<treatmentCitationGroup id="AB606B03FFB8FF8FFF27F90CA934F95B" box="[192,1029,1703,1728]" pageId="35" pageNumber="1740">
<treatmentCitation id="0AD16A3CFFB8FF8FFF27F90CAE3DF95B" author="Santschi F" box="[192,780,1703,1728]" page="229" pageId="35" pageNumber="1740" year="1927">
<taxonomicName id="4C7037AEFFB8FF8FFF27F90CAE3DF95B" ID-CoL="427ZS" authority="Santschi, 1927, p. 229" authorityName="Santschi" authorityPageNumber="229" authorityYear="1927" box="[192,780,1703,1728]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="1740" phylum="Arthropoda" rank="variety" species="semirufus" variety="ebeninus">
<emphasis id="B904903FFFB8FF8FFF27F90CAC4AF95B" box="[192,379,1703,1728]" italics="true" pageId="35" pageNumber="1740">Messor semirufus</emphasis>
var.
<emphasis id="B904903FFFB8FF8FFE56F90CAF23F95B" box="[433,530,1703,1728]" italics="true" pageId="35" pageNumber="1740">ebeninus</emphasis>
<bibRefCitation id="EFE131DCFFB8FF8FFDFEF90CAF8AF95B" author="Santschi F" box="[537,699,1703,1728]" pageId="35" pageNumber="1740" pagination="225 - 250" refId="ref23074" refString="Santschi F. 1927. Revision des Messor du groupe instabilis Sm. (Hymenopt.). Bol R Soc Esp Hist Nat. 27: 225 - 250." type="journal article" year="1927">Santschi, 1927</bibRefCitation>
, p. 229
</taxonomicName>
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(w.)
<collectingCountry id="F3670CBDFFB8FF8FFCA3F90CAE4DF95B" box="[836,892,1703,1728]" name="Syria" pageId="35" pageNumber="1740">Syria</collectingCountry>
. Palaearctic.
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</paragraph>
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<caption id="DF0F1CA5FFBFFF88FF47F971AE50F895" ID-DOI="http://doi.org/10.5281/zenodo.14241369" ID-Zenodo-Dep="14241369" httpUri="https://zenodo.org/record/14241369/files/figure.png" pageId="36" pageNumber="1741" startId="36.[160,227,1754,1777]" targetBox="[368,947,145,1719]" targetPageId="36" targetType="figure">
<paragraph id="8BCF4C2DFFBFFF88FF47F971AE50F895" blockId="36.[160,1157,1754,1806]" pageId="36" pageNumber="1741">
<emphasis id="B904903FFFBFFF88FF47F971AC20F96A" bold="true" box="[160,273,1754,1777]" pageId="36" pageNumber="1741">Figure 22.</emphasis>
<taxonomicName id="4C7037AEFFBFFF88FEC7F971AF21F96A" authorityName="Seifert" authorityYear="2003" box="[288,528,1754,1777]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="1741" phylum="Arthropoda" rank="species" species="rugulosa">
<emphasis id="B904903FFFBFFF88FEC7F971AF21F96A" box="[288,528,1754,1777]" italics="true" pageId="36" pageNumber="1741">Cardiocondyla rugulosa</emphasis>
</taxonomicName>
. Worker. (A) Body in profile. (B) Head in full-face view, ANTWEB1041915, AntWeb.org (Michele Esposito). (C) Distribution map.
</paragraph>
</caption>
<caption id="DF0F1CA5FFBEFF89FF47F971AF9DF895" ID-DOI="http://doi.org/10.5281/zenodo.14241371" ID-Zenodo-Dep="14241371" httpUri="https://zenodo.org/record/14241371/files/figure.png" pageId="37" pageNumber="1742" startId="37.[160,225,1754,1777]" targetBox="[394,921,144,1719]" targetPageId="37" targetType="figure">
<paragraph id="8BCF4C2DFFBEFF89FF47F971AF9DF895" blockId="37.[160,1157,1754,1806]" pageId="37" pageNumber="1742">
<emphasis id="B904903FFFBEFF89FF47F971AC39F96A" bold="true" box="[160,264,1754,1777]" pageId="37" pageNumber="1742">Figure 23.</emphasis>
<taxonomicName id="4C7037AEFFBEFF89FEE8F971AC9CF96A" authorityName="Santschi" authorityYear="1927" box="[271,429,1754,1777]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="1742" phylum="Arthropoda" rank="species" species="ebeninus">
<emphasis id="B904903FFFBEFF89FEE8F971AC9CF96A" box="[271,429,1754,1777]" italics="true" pageId="37" pageNumber="1742">Messor ebeninus</emphasis>
</taxonomicName>
. Worker. (A) Body in profile. (B) Head in full-face view, ANTWEB1041905, AntWeb.org (Michele Esposito). (C) Distribution map.
</paragraph>
</caption>
<caption id="DF0F1CA5FFBDFF8AFF47F971AFF7F895" ID-DOI="http://doi.org/10.5281/zenodo.14241373" ID-Zenodo-Dep="14241373" httpUri="https://zenodo.org/record/14241373/files/figure.png" pageId="38" pageNumber="1743" startId="38.[160,225,1754,1777]" targetBox="[329,986,145,1719]" targetPageId="38" targetType="figure">
<paragraph id="8BCF4C2DFFBDFF8AFF47F971AFF7F895" blockId="38.[160,1156,1754,1806]" pageId="38" pageNumber="1743">
<emphasis id="B904903FFFBDFF8AFF47F971AC39F96A" bold="true" box="[160,264,1754,1777]" pageId="38" pageNumber="1743">Figure 24.</emphasis>
<taxonomicName id="4C7037AEFFBDFF8AFEE8F971AC9FF96A" authorityName="Santschi" authorityYear="1927" box="[271,430,1754,1777]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="1743" phylum="Arthropoda" rank="species" species="ebeninus">
<emphasis id="B904903FFFBDFF8AFEE8F971AC9FF96A" box="[271,430,1754,1777]" italics="true" pageId="38" pageNumber="1743">Messor ebeninus</emphasis>
</taxonomicName>
. Queen. (A) Body in profile. (B) Body in dorsal view. (C) Head in full-face view, ANTWEB1041906, AntWeb.org (Michele Esposito).
</paragraph>
</caption>
<subSubSection id="C36A1FA6FFBCFF8BFF47FF25A95AFE7B" pageId="39" pageNumber="1744" type="diagnosis">
<paragraph id="8BCF4C2DFFBCFF8BFF47FF25AC25FF33" blockId="39.[160,1156,142,480]" box="[160,276,142,168]" pageId="39" pageNumber="1744">
<emphasis id="B904903FFFBCFF8BFF47FF25AC25FF33" bold="true" box="[160,276,142,168]" italics="true" pageId="39" pageNumber="1744">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BCF4C2DFFBCFF8BFF47FF1AA95AFE7B" blockId="39.[160,1156,142,480]" pageId="39" pageNumber="1744">
<emphasis id="B904903FFFBCFF8BFF47FF1AADCCFF50" bold="true" box="[160,253,177,203]" italics="true" pageId="39" pageNumber="1744">Worker.</emphasis>
Colour slightly variable, ranging from uniform dark brown to black, some specimens have head, mesosoma, petiole, and postpetiole brown contrasting with brown or black gaster; underside of head with long J-shaped setae forming a distinct psammophore; propodeum angled, without spines or teeth of any description; posterior margin of head with 35 setae at each side of the midline; mesosoma with several pairs of setae, two or three on promesonotum, one or two pairs on mesonotum, a single pair on propodeum; petiole and postpetiole each with one pair of long setae; first gastral tergite bare except for few pairs of seta on posterior margin; mesosoma partially or completely rugulose; head smooth and shining, in some individuals vertex with faint longitudinal striations.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FFBCFF8BFF47FDB0AC75FDC3" pageId="39" pageNumber="1744" type="materials_examined">
<paragraph id="8BCF4C2DFFBCFF8BFF47FDB0AC4EFDAE" blockId="39.[160,383,539,600]" box="[160,383,539,565]" pageId="39" pageNumber="1744">
<heading id="D087FB41FFBCFF8BFF47FDB0AC4EFDAE" bold="true" box="[160,383,539,565]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="1744" reason="3">
<emphasis id="B904903FFFBCFF8BFF47FDB0AC4EFDAE" bold="true" box="[160,383,539,565]" italics="true" pageId="39" pageNumber="1744">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF4C2DFFBCFF8BFF47FD94AC75FDC3" blockId="39.[160,383,539,600]" box="[160,324,575,600]" pageId="39" pageNumber="1744">Two sites: 3; 8.</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FFBCFF8BFF47FD38AFB4FD68" pageId="39" pageNumber="1744" type="distribution">
<paragraph id="8BCF4C2DFFBCFF8BFF47FD38AFB4FD68" blockId="39.[160,1156,659,755]" pageId="39" pageNumber="1744">
<emphasis id="B904903FFFBCFF8BFF47FD38AC4DFD36" bold="true" box="[160,380,659,685]" italics="true" pageId="39" pageNumber="1744">Geographic range.</emphasis>
This species is widespread in the Middle East, with a broad range of distribution in the Arabian Peninsula (
<bibRefCitation id="EFE131DCFFBCFF8BFDA8FD1CAE2EFD4B" author="Collingwood CA" box="[591,799,695,720]" pageId="39" pageNumber="1744" pagination="230 - 302" refId="ref19269" refString="Collingwood CA. 1985. Hymenoptera: fam. Formicidae of Saudi Arabia. Fauna Saudi Arabia. 7: 230 - 302. doi: 10.5281 / zenodo. 26692." type="journal article" year="1985">Collingwood 1985</bibRefCitation>
;
<bibRefCitation id="EFE131DCFFBCFF8BFCCAFD1CA94FFD4B" author="Collingwood CA &amp; Agosti D" box="[813,1150,695,720]" pageId="39" pageNumber="1744" pagination="300 - 385" refId="ref19301" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
;
<bibRefCitation id="EFE131DCFFBCFF8BFF47FD71AC9BFD68" author="Collingwood CA &amp; Agosti D &amp; Sharaf MR &amp; van Harten A" box="[160,426,730,755]" pageId="39" pageNumber="1744" pagination="405 - 474" refId="ref19333" refString="Collingwood CA, Agosti D, Sharaf MR, van Harten A. 2011. Order Hymenoptera, family Formicidae. Arthropod Fauna UAE. 4: 405 - 474. doi: 10.5281 / zenodo. 1168586." type="journal article" year="2011">
Collingwood
<emphasis id="B904903FFFBCFF8BFED1FD71AC57FD68" box="[310,358,730,755]" italics="true" pageId="39" pageNumber="1744">et al</emphasis>
. 2011
</bibRefCitation>
;
<bibRefCitation id="EFE131DCFFBCFF8BFE50FD71AF49FD68" author="Sharaf MR &amp; Al Dhafer HM &amp; Aldawood AS &amp; Hita Garcia F" box="[439,632,730,755]" pageId="39" pageNumber="1744" refId="ref24023" refString="Sharaf MR, Al Dhafer HM, Aldawood AS, Hita Garcia F. 2018 a. Ants of the Monomorium monomorium species-group (Hymenoptera: Formicidae) in the Arabian Peninsula with description of a new species from southwestern Saudi Arabia. PeerJ. 6: 31. e 4277. doi: 10.7717 / peerj. 4277." type="journal volume" year="2018">
Sharaf
<emphasis id="B904903FFFBCFF8BFDE3FD71AF05FD68" box="[516,564,730,755]" italics="true" pageId="39" pageNumber="1744">et al</emphasis>
. 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FFBCFF8BFF47FC85AE64FCF0" pageId="39" pageNumber="1744" type="biology_ecology">
<paragraph id="8BCF4C2DFFBCFF8BFF47FC85AE64FCF0" blockId="39.[160,1155,814,875]" pageId="39" pageNumber="1744">
<emphasis id="B904903FFFBCFF8BFF47FC85ACADFCD3" bold="true" box="[160,412,814,840]" italics="true" pageId="39" pageNumber="1744">Ecology and biology.</emphasis>
This species, like the majority of species of the genus
<taxonomicName id="4C7037AEFFBCFF8BFBFEFC84A955FCD3" authorityName="Forel" authorityYear="1890" box="[1049,1124,815,840]" class="Insecta" family="Formicidae" genus="Messor" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="1744" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFFBCFF8BFBFEFC84A955FCD3" box="[1049,1124,815,840]" italics="true" pageId="39" pageNumber="1744">Messor</emphasis>
</taxonomicName>
, is a seed harvester. It nests directly in the ground under stones.
</paragraph>
</subSubSection>
</treatment>
</document>