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<mods:title id="E4639E4F56A1515600B84477D01656EB">Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)</mods:title>
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<mods:namePart id="C5FE817EA16930615A0B1C0211F2F396">Ramos, Magno S.</mods:namePart>
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<treatment id="03D887D08A3AFF841399FC46FD5EF870" ID-DOI="http://doi.org/10.5281/zenodo.5461825" ID-Zenodo-Dep="5461825" LSID="urn:lsid:plazi:treatment:03D887D08A3AFF841399FC46FD5EF870" httpUri="http://treatment.plazi.org/id/03D887D08A3AFF841399FC46FD5EF870" lastPageNumber="10" pageId="9" pageNumber="10">
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<paragraph id="8BCE36C68A3AFF841399FC46FE13FBFD" blockId="9.[151,495,1015,1041]" box="[151,495,1015,1041]" pageId="9" pageNumber="10">
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<taxonomicName id="4C714D458A3AFF841399FC46FE13FBFD" ID-CoL="4GGW" ID-ENA="1380409" authority="Westwood, 1874" authorityName="Westwood" authorityYear="1874" box="[151,495,1015,1041]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAD48A3AFF841399FC46FE13FBFD" bold="true" box="[151,495,1015,1041]" pageId="9" pageNumber="10">
<emphasis id="B905EAD48A3AFF841399FC46FEE2FBFD" bold="true" box="[151,286,1015,1041]" italics="true" pageId="9" pageNumber="10">Eupsenella</emphasis>
<bibRefCitation id="EFE04B378A3AFF84122BFC46FE13FBFD" author="Westwood" box="[293,495,1015,1041]" firstAuthor="Westwood" pageId="9" pageNumber="10" refId="ref65716" refString="Westwood, J. O. (1874) Thesaurus Entomologicus Oxoniensis; or, illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the university of oxford by the rev. f. w. hope, m. a., d. c. l., f. r. s., &amp; C. with forty plates from deawings by the author. Clarendon Press, Oxford. 205 p. + 40 plates." type="book" year="1874">Westwood, 1874</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BCE36C68A3AFF841399FB8EFC89FB7D" blockId="9.[151,1436,1086,1169]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB6111E88A3AFF841399FB8EFC89FB7D" pageId="9" pageNumber="10">
<treatmentCitation id="0AD010D78A3AFF841399FB8EFE1AFBB8" author="Westwood, J. O." box="[151,486,1086,1109]" page="168" pageId="9" pageNumber="10" year="1874">
<taxonomicName id="4C714D458A3AFF841399FB8EFE1AFBB8" ID-CoL="4GGW" authority="Westwood 1874: 168" authorityName="Westwood" authorityPageNumber="168" authorityYear="1874" box="[151,486,1086,1109]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAD48A3AFF841399FB8EFEFBFBB9" box="[151,263,1087,1109]" italics="true" pageId="9" pageNumber="10">Eupsenella</emphasis>
<bibRefCitation id="EFE04B378A3AFF841200FB8FFE1AFBB8" author="Westwood" box="[270,486,1086,1108]" firstAuthor="Westwood" page="168" pageId="9" pageNumber="10" refId="ref65716" refString="Westwood, J. O. (1874) Thesaurus Entomologicus Oxoniensis; or, illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the university of oxford by the rev. f. w. hope, m. a., d. c. l., f. r. s., &amp; C. with forty plates from deawings by the author. Clarendon Press, Oxford. 205 p. + 40 plates." type="book" year="1874">Westwood 1874: 168</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 30, fig. 6;
<bibRefCitation id="EFE04B378A3AFF841175FB8FFCEBFBB8" author="Ashmead" box="[635,791,1086,1108]" firstAuthor="Ashmead" pageId="9" pageNumber="10" pagination="1 - 472" refId="ref64604" refString="Ashmead, W. H. (1893) A monograph of the North American Proctotrypidae. Bethylibae. Bulletin of the United States National Museum. 45, 1 - 472." type="journal article" year="1893">Ashmead 1893</bibRefCitation>
: pl 4,
<figureCitation id="134A2A438A3AFF84105AFB8FFC6CFBB8" box="[852,912,1086,1108]" captionStart="FIGURES 1–2" captionStartId="61.[151,264,1938,1961]" captionTargetBox="[151,1436,921,1913]" captionTargetId="figure-282@61.[151,1436,917,1913]" captionTargetPageId="61" captionText="FIGURES 1–2. Forewing. 1–2 Eupsenella barada sp. nov.; 1. Cells of forewing. 2. Veins of forewing." figureDoi="http://doi.org/10.5281/zenodo.10096761" httpUri="https://zenodo.org/record/10096761/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
;
<treatmentCitation id="0AD010D78A3AFF841093FB8FFB99FBB8" author="Dalla Torre, C. G. de" box="[925,1125,1086,1108]" page="5" pageId="9" pageNumber="10" year="1898">
<bibRefCitation id="EFE04B378A3AFF841093FB8FFB99FBB8" author="Dalla Torre" box="[925,1125,1086,1108]" firstAuthor="Dalla Torre" page="5" pageId="9" pageNumber="10" refId="ref64933" refString="Dalla Torre, C. G. de (1898) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Chalcididae et Proctotrupidae., 5. Suptibus Guilelmi Engelmann. Lipsiae. 598 p." type="book" year="1898">Dalla Torre 1898: 5</bibRefCitation>
</treatmentCitation>
, 555;
<treatmentCitation id="0AD010D78A3AFF8417AAFB8FFAB2FBB8" author="Kieffer, J. J." box="[1188,1358,1086,1108]" page="12" pageId="9" pageNumber="10" year="1908">
<bibRefCitation id="EFE04B378A3AFF8417AAFB8FFAB2FBB8" author="Kieffer" box="[1188,1358,1086,1108]" firstAuthor="Kieffer" page="12" pageId="9" pageNumber="10" refId="ref65371" refString="Kieffer, J. J. (1908) Bethylidae. Fascicule 76. In: Wytsman, P. Genera Insectorum, 50 p." type="book" year="1908">Kieffer 1908: 12</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="134A2A438A3AFF841657FB8FFA69FBB8" box="[1369,1429,1086,1108]" captionStart="FIGURES 1–2" captionStartId="61.[151,264,1938,1961]" captionTargetBox="[151,1436,921,1913]" captionTargetId="figure-282@61.[151,1436,917,1913]" captionTargetPageId="61" captionText="FIGURES 1–2. Forewing. 1–2 Eupsenella barada sp. nov.; 1. Cells of forewing. 2. Veins of forewing." figureDoi="http://doi.org/10.5281/zenodo.10096761" httpUri="https://zenodo.org/record/10096761/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
; Kieffer 1914: 556–558, Fig. 198;
<treatmentCitation id="0AD010D78A3AFF841111FBECFD1CFB9F" author="Dodd, A. P." box="[543,736,1117,1139]" page="18" pageId="9" pageNumber="10" year="1916">
<bibRefCitation id="EFE04B378A3AFF841111FBECFD1CFB9F" author="Dodd" box="[543,736,1117,1139]" firstAuthor="Dodd" page="18 - 19" pageId="9" pageNumber="10" pagination="9 - 32" refId="ref65048" refString="Dodd, A. P. (1916) Australian Hymenoptera: Proctotrypoidea. No. 4. Royal Society of South Australia. Transactions and Proceedings, 40, 9 - 32." type="journal article" year="1916">Dodd 1916: 18–19</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD010D78A3AFF8411E5FBEFFC4EFB9F" author="Lameere, A." box="[747,946,1117,1139]" page="440" pageId="9" pageNumber="10" year="1938">
<bibRefCitation id="EFE04B378A3AFF8411E5FBEFFC4EFB9F" author="Lameere" box="[747,946,1117,1139]" firstAuthor="Lameere" page="440" pageId="9" pageNumber="10" refId="ref65397" refString="Lameere, A. (1938) Precis de ZOO / Ogie • 5 • Les 1 nsectes Superieurs • StegOpteres, Dipteres, LepidOpteres, CO / eOpteres, HymenOpteres • Bruxelles, Universit, 536 pp." type="book" year="1938">Lameere 1938: 440</bibRefCitation>
</treatmentCitation>
, Fig. 510;
<treatmentCitation id="0AD010D78A3AFF84172EFBECFA64FB9F" author="Polaszek, A. &amp; Krombein, K. V." box="[1056,1432,1117,1139]" page="91" pageId="9" pageNumber="10" year="1994">
<bibRefCitation id="EFE04B378A3AFF84172EFBECFA64FB9F" author="Polaszek &amp; Krombein" box="[1056,1432,1117,1139]" firstAuthor="Polaszek" page="91 - 105" pageId="9" pageNumber="10" pagination="91 - 105" refId="ref65526" refString="Polaszek, A. &amp; Krombein, K. V. (1994) The genera of Bethylinae (Hymenoptera: Bethylinae). Journal of Hymenoptera Research, 3, 91 - 105." type="journal article" year="1994">Polaszek &amp; Krombein 1994: 91–105</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD010D78A3AFF8413C9FBCAFDF5FB7D" author="Gordh, G. &amp; Harris, A." box="[199,521,1147,1169]" page="530" pageId="9" pageNumber="10" year="1996">
<bibRefCitation id="EFE04B378A3AFF8413C9FBCAFDF5FB7D" author="Gordh &amp; Harris" box="[199,521,1147,1169]" firstAuthor="Gordh" page="530 - 534" pageId="9" pageNumber="10" pagination="529 - 536" refId="ref65238" refString="Gordh, G. &amp; Harris, A. (1996) New records and a new species of Eupsenella (Hymenoptera: Bethylidae) in New Zealand. Journal of The Royal Society of New Zealand, 26, 529 - 536." type="journal article" year="1996">Gordh &amp; Harris 1996: 530–534</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="134A2A438A3AFF84111CFBCAFD85FB7D" box="[530,633,1147,1169]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–14" captionStartId-0="61.[151,264,1938,1961]" captionStartId-1="62.[151,263,1843,1866]" captionTargetBox-0="[151,1436,921,1913]" captionTargetBox-1="[151,1436,193,1822]" captionTargetId-0="figure-282@61.[151,1436,917,1913]" captionTargetId-1="figure-83@62.[151,1436,193,1822]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURES 1–2. Forewing. 1–2 Eupsenella barada sp. nov.; 1. Cells of forewing. 2. Veins of forewing." captionText-1="FIGURES 3–14. Head in dorsal view. 3. Eupsenella ceciliae. 4. E.diemenensis. 5. E. flavifemorata. 6. E. insulana female. 7. E. insulana male. 8. E. reticulata. 9. E. ajabatha sp. nov. 10. E. alawa sp. nov. 11. E. alura sp. nov. 12. E. antakirinja sp. nov. 13. E. araba sp. nov. 14. E. arabana sp. nov. (Scale bar = 300 µm)." figureDoi-0="http://doi.org/10.5281/zenodo.10096761" figureDoi-1="http://doi.org/10.5281/zenodo.10096763" httpUri-0="https://zenodo.org/record/10096761/files/figure.png" httpUri-1="https://zenodo.org/record/10096763/files/figure.png" pageId="9" pageNumber="10">Figs 1–13</figureCitation>
;
<treatmentCitation id="0AD010D78A3AFF84118AFBCDFC8CFB7D" author="Terayama, M." box="[644,880,1147,1169]" page="39" pageId="9" pageNumber="10" year="2004">
<bibRefCitation id="EFE04B378A3AFF84118AFBCDFC8CFB7D" author="Terayama" box="[644,880,1147,1169]" firstAuthor="Terayama" page="39 - 41" pageId="9" pageNumber="10" pagination="39 - 41" refId="ref65636" refString="Terayama, M. (2004) Descriptions of new taxa and distribution records of the family Bethylidae (Insecta, Hymenoptera) II. Subfamily Bethylinae and fossil Taxa. Academic reports, Faculty Engineering. Tokyo Polytechnic University, 27, 39 - 41." type="journal article" year="2004">Terayama 2004: 39–41</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A3AFF841399FB0BFCD9FB23" box="[151,805,1209,1231]" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BCE36C68A3AFF841399FB0BFCD9FB23" blockId="9.[151,805,1209,1231]" box="[151,805,1209,1231]" pageId="9" pageNumber="10">
<typeStatus id="54CA88648A3AFF841399FB0BFF37FB23" box="[151,203,1210,1231]" pageId="9" pageNumber="10">Type</typeStatus>
species:
<taxonomicName id="4C714D458A3AFF841228FB08FD72FB23" authority="Westwood, 1874" authorityName="Westwood" authorityYear="1874" box="[294,654,1209,1231]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="agillis">
<emphasis id="B905EAD48A3AFF841228FB08FE20FB23" box="[294,476,1209,1231]" italics="true" pageId="9" pageNumber="10">Eupsenella agillis</emphasis>
<bibRefCitation id="EFE04B378A3AFF8412ECFB08FD72FB23" author="Westwood" box="[482,654,1209,1231]" firstAuthor="Westwood" pageId="9" pageNumber="10" refId="ref65716" refString="Westwood, J. O. (1874) Thesaurus Entomologicus Oxoniensis; or, illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the university of oxford by the rev. f. w. hope, m. a., d. c. l., f. r. s., &amp; C. with forty plates from deawings by the author. Clarendon Press, Oxford. 205 p. + 40 plates." type="book" year="1874">Westwood, 1874</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A3AFF841399FB4AFCF3F89B" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BCE36C68A3AFF841399FB4AFE3EF8E0" blockId="9.[151,1436,1275,1948]" pageId="9" pageNumber="10">
<emphasis id="B905EAD48A3AFF841399FB4AFEEFFAF8" bold="true" box="[151,275,1275,1300]" inLineHeading="true" pageId="9" pageNumber="10" reason="1">Diagnosis.</emphasis>
Median clypeal lobe truncate, angulate or subangulate, median carina present or absent, extending back into frons. Lateral lobe of clypeus almost inconspicuous. Mandible with four sharpened apical teeth. Antenna with 13 articules; flagellar pubescence suberect. Torulus not projected. Hipostomal carina present and well defined. Pronotal collar regular-sized, evenly narrow. Notauli present, well defined, complete, extending from anterior margin of the posterior margin of mesoscutum. Scutellar pit present. Scutellar furrow absent. Propodeal disc with median carina, posterior transverse carina absent and lateral carina present, followed by depressed narrow area, foveolate. Declivity of propodeum without median carina. Mesopleuron with mesopleural pit deep. Prosternum expanded, 0.7 to 1.0 area of procoxa and excavated medially. Mesofurcal pit present. Metasoma in ventral view with ventral carina of petiole. Forewing with six closed cells, (R, 1Cu, C, 1M, 1R1 and 2R1), except in brachypterous species, stigma regular-sized, except in brachypterous species. Hind wing with incipient anal vein and jugal lobe, hamuli located approximately medially on anterior margin. Male slightly smaller than female. Hypopygium puncticulate, with posterior margin bidentate and angulate. Genitalia with one or two parameres, paramere not fused to basiparamere; dorsal arm of paramere as long as ventral arm; apical lobe of aedeagus expanded; apex of apical lobe of aedeagus; not aligned with paremere apex; apical margin of cuspis rounded.
</paragraph>
<paragraph id="8BCE36C68A3AFF8413C9F8A6FCF3F89B" blockId="9.[151,1436,1275,1948]" pageId="9" pageNumber="10">
<emphasis id="B905EAD48A3AFF8413C9F8A6FE6FF8DC" bold="true" box="[199,403,1815,1840]" inLineHeading="true" pageId="9" pageNumber="10" reason="1">Species included.</emphasis>
<taxonomicName id="4C714D458A3AFF841294F8A9FD1FF8C3" authority="Westwood" authorityName="Westwood" authorityYear="1874" box="[410,739,1815,1840]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="agillis">
<emphasis id="B905EAD48A3AFF841294F8A9FD98F8DC" box="[410,612,1816,1840]" italics="true" pageId="9" pageNumber="10">Eupsenella agillis</emphasis>
Westwood
</taxonomicName>
,
<taxonomicName id="4C714D458A3AFF8411E1F8A8FC1CF8DC" authority="Terayama" authorityName="Terayama" authorityYear="2004" box="[751,992,1816,1840]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ceciliae">
<emphasis id="B905EAD48A3AFF8411E1F8A8FC99F8DC" box="[751,869,1816,1840]" italics="true" pageId="9" pageNumber="10">E. ceciliae</emphasis>
Terayama
</taxonomicName>
,
<taxonomicName id="4C714D458A3AFF8410E7F8A8FB26F8C3" authority="Dodd" authorityName="Dodd" authorityYear="1916" box="[1001,1242,1815,1840]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="diemenensis">
<emphasis id="B905EAD48A3AFF8410E7F8A8FB6EF8DC" box="[1001,1170,1816,1840]" italics="true" pageId="9" pageNumber="10">E. diemenensis</emphasis>
Dodd
</taxonomicName>
,
<taxonomicName id="4C714D458A3AFF8417EBF8A8FEF6F8B8" authority="Terayama" authorityName="Terayama" authorityYear="2004" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="flavifemorata">
<emphasis id="B905EAD48A3AFF8417EBF8A8FA60F8DC" box="[1253,1436,1816,1840]" italics="true" pageId="9" pageNumber="10">E. flavifemorata</emphasis>
Terayama
</taxonomicName>
,
<taxonomicName id="4C714D458A3AFF841219F88CFD9EF8B8" authority="Gordh &amp; Harris" authorityName="Gordh &amp; Harris" authorityYear="1996" box="[279,610,1852,1876]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B905EAD48A3AFF841219F88CFE66F8B8" box="[279,410,1852,1876]" italics="true" pageId="9" pageNumber="10">E. insulana</emphasis>
Gordh &amp; Harris
</taxonomicName>
,
<taxonomicName id="4C714D458A3AFF84117CF88CFCB0F8B8" authority="Kieffer" authorityName="Kieffer" authorityYear="1911" box="[626,844,1852,1876]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="proxima">
<emphasis id="B905EAD48A3AFF84117CF88CFD0DF8B8" box="[626,753,1853,1876]" italics="true" pageId="9" pageNumber="10">E. proxima</emphasis>
Kieffer
</taxonomicName>
(analysis only through the available literature),
<taxonomicName id="4C714D458A3AFF84168AF88CFE87F894" authority="Terayama" authorityName="Terayama" authorityYear="2004" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="B905EAD48A3AFF84168AF88CFEFFF894" italics="true" pageId="9" pageNumber="10">E. reticulata</emphasis>
Terayama
</taxonomicName>
, and 38 new species described here.
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A3AFF8413C9F832FD5EF870" box="[199,674,1923,1948]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BCE36C68A3AFF8413C9F832FD5EF870" blockId="9.[151,1436,1275,1948]" box="[199,674,1923,1948]" pageId="9" pageNumber="10">
<emphasis id="B905EAD48A3AFF8413C9F832FE9CF870" bold="true" box="[199,352,1923,1948]" inLineHeading="true" pageId="9" pageNumber="10" reason="1">Distribution.</emphasis>
<collectingCountry id="F36676568A3AFF841269F835FE32F870" box="[359,462,1924,1948]" name="Australia" pageId="9" pageNumber="10">Australia</collectingCountry>
and
<collectingCountry id="F36676568A3AFF84110AF835FD62F870" box="[516,670,1924,1948]" name="New Zealand" pageId="9" pageNumber="10">New Zealand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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