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<document id="0A2AF4683E625227C843E56C3ED3F733" ID-DOI="10.11646/zootaxa.3539.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10096759" ID-ZooBank="0564EF02-FCF9-49A8-BDAC-F432876C16B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629821597637" checkinUser="felipe" docAuthor="Ramos, Magno S. &amp; Azevedo, Celso O." docDate="2012" docId="03D887D08A27FF981399FACBFA60F878" docLanguage="en" docName="zt03539p080.pdf" docOrigin="Zootaxa 3539 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3539.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eupsenella arabana Ramos &amp; Azevedo, sp. nov." docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="FFE1FFA88A33FF8D130EFFB1FFFCFFEC" masterDocTitle="Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="21" updateTime="1699581041301" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="04EC81888EA22D34D3E934C4F50A8B0E">Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)</mods:title>
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<mods:namePart id="030E6E21D3F266B6B3348026A8F5A474">Ramos, Magno S.</mods:namePart>
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<mods:namePart id="23842D4BBF02A674DD364321B646DA5C">Azevedo, Celso O.</mods:namePart>
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<treatment id="03D887D08A27FF981399FACBFA60F878" LSID="urn:lsid:plazi:treatment:03D887D08A27FF981399FACBFA60F878" httpUri="http://treatment.plazi.org/id/03D887D08A27FF981399FACBFA60F878" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="C36B654D8A27FF991399FACBFD18FA78" box="[151,740,1402,1428]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BCE36C68A27FF991399FACBFD18FA78" blockId="20.[151,740,1402,1461]" box="[151,740,1402,1428]" pageId="20" pageNumber="21">
<heading id="D08681AA8A27FF991399FACBFD18FA78" bold="true" box="[151,740,1402,1428]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B905EAD48A27FF991399FACBFD18FA78" bold="true" box="[151,740,1402,1428]" pageId="20" pageNumber="21">
<taxonomicName id="4C714D458A27FF991399FACBFD85FA78" authority="Ramos &amp; Azevedo" authorityName="Ramos &amp; Azevedo" box="[151,633,1402,1428]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arabana" status="sp. nov.">
<emphasis id="B905EAD48A27FF991399FACBFE76FA78" bold="true" box="[151,394,1402,1428]" italics="true" pageId="20" pageNumber="21">Eupsenella arabana</emphasis>
Ramos &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A23657AF8A27FF991189FACBFD18FA78" box="[647,740,1402,1428]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A27FF981399FA2CFE47FB0B" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="8BCE36C68A27FF991399FA2CFDFBFA59" blockId="20.[151,740,1402,1461]" box="[151,519,1437,1461]" pageId="20" pageNumber="21">
(
<figureCitation id="134A2A438A27FF991391FA2CFF09FA59" box="[159,245,1437,1461]" captionStart="FIGURES 314" captionStartId="62.[151,263,1843,1866]" captionTargetBox="[151,1436,193,1822]" captionTargetId="figure-83@62.[151,1436,193,1822]" captionTargetPageId="62" captionText="FIGURES 314. Head in dorsal view. 3. Eupsenella ceciliae. 4. E.diemenensis. 5. E. flavifemorata. 6. E. insulana female. 7. E. insulana male. 8. E. reticulata. 9. E. ajabatha sp. nov. 10. E. alawa sp. nov. 11. E. alura sp. nov. 12. E. antakirinja sp. nov. 13. E. araba sp. nov. 14. E. arabana sp. nov. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096763" httpUri="https://zenodo.org/record/10096763/files/figure.png" pageId="20" pageNumber="21">Figs 14</figureCitation>
, 59, 104, 144, 163165)
</paragraph>
<paragraph id="8BCE36C68A27FF991399FA54FCC0FA11" blockId="20.[151,1436,1509,1930]" box="[151,828,1509,1534]" pageId="20" pageNumber="21">
<emphasis id="B905EAD48A27FF991399FA54FED4FA12" bold="true" box="[151,296,1509,1534]" inLineHeading="true" pageId="20" pageNumber="21" reason="1">Description.</emphasis>
MALE. Body length
<quantity id="4C899B238A27FF991113FA54FD78FA11" box="[541,644,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" pageId="20" pageNumber="21" unit="mm" value="4.85">4.85 mm</quantity>
. LFW
<quantity id="4C899B238A27FF9911DFFA54FCC4FA11" box="[721,824,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="3.58" pageId="20" pageNumber="21" unit="mm" value="3.58">3.58 mm</quantity>
.
</paragraph>
<paragraph id="8BCE36C68A27FF9913C9F9B8FC0DF961" blockId="20.[151,1436,1509,1930]" pageId="20" pageNumber="21">
<emphasis id="B905EAD48A27FF9913C9F9B8FEB5F9CD" box="[199,329,1545,1569]" italics="true" pageId="20" pageNumber="21">Coloration.</emphasis>
Head dark castaneous almost black; scape, pedicel and flagellum castaneous; mandible light castaneous with teeth dark castaneous; palpi castaneous; mesosoma dark castaneous almost black; tegula castaneous; wing venation light castaneous; procoxa, mesocoxa, and metacoxa castaneous; trochanters light castaneous; femora castaneous; tarsi light castaneous; metasoma castaneous.
</paragraph>
<paragraph id="8BCE36C68A27FF9913C9F928FA60F865" blockId="20.[151,1436,1509,1930]" pageId="20" pageNumber="21">
<emphasis id="B905EAD48A27FF9913C9F928FEF8F95D" box="[199,260,1689,1713]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
(
<figureCitation id="134A2A438A27FF99121DF928FE99F95D" box="[275,357,1689,1713]" captionStart="FIGURES 314" captionStartId="62.[151,263,1843,1866]" captionTargetBox="[151,1436,193,1822]" captionTargetId="figure-83@62.[151,1436,193,1822]" captionTargetPageId="62" captionText="FIGURES 314. Head in dorsal view. 3. Eupsenella ceciliae. 4. E.diemenensis. 5. E. flavifemorata. 6. E. insulana female. 7. E. insulana male. 8. E. reticulata. 9. E. ajabatha sp. nov. 10. E. alawa sp. nov. 11. E. alura sp. nov. 12. E. antakirinja sp. nov. 13. E. araba sp. nov. 14. E. arabana sp. nov. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096763" httpUri="https://zenodo.org/record/10096763/files/figure.png" pageId="20" pageNumber="21">Fig. 14</figureCitation>
). Head wider than long, globoid in profile; gena weakly concave in profile. Mandible with base wider than apex, two lowermost teeth larger than upper ones, lower tooth curved inward, lower margin strongly setose. Clypeus coriaceous, median clypeal lobe angulate, median concavity of median clypeal lobe absent, 0.08 LH, well marked off from frons, median carina present, 0.24 LH. Malar space inconspicuous, 0.18 mandibular base. Antenna
<quantity id="4C899B238A27FF99124CF898FE62F8AD" box="[322,414,1833,1857]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="21" unit="mm" value="1.2">1.2 mm</quantity>
; first five antennomeres in ratio of about 12:12:12:12:12; flagellomere 1 1.5 longer than wide, flagellar pubescence dense, with few outstanding erect setae. Region between torulus and eye concave. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.10 LH. Eye protuberant, 0.58 LH,
</paragraph>
<paragraph id="8BCE36C68A27FF991399F824FA60F819" blockId="20.[151,1436,1941,2037]" pageId="20" pageNumber="21">with sparse hairs, located more laterally, lower margin without strong depression. Frons coriaceous-punctate. WH 1.24 LH. WF 0.62 WH. WF 1.3 HE. OOL 0.72 WOT. VOL 0.4 HE. Distance from posterior ocellus to vertex crest 0.33 DAO. Ocelli large, DAO 0.15 WF, ocellar triangle not compact, distance between posterior ocelli 1.90 DAO,</paragraph>
<paragraph id="8BCE36C68A26FF981399FF26FDA4FF1B" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">anterior ocellus not surpassing imaginary top eye line, posterior ocellus almost reaching vertex. Vertex straight, short setae, 0.10 LH, temple profile slightly convergent in dorsal view, with corner convex, not projected posterolaterad. Hipostomal carina arched, high.</paragraph>
<paragraph id="8BCE36C68A26FF9813C9FEB4FC1EFD4B" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FEB4FEC3FEF0" box="[199,319,261,284]" italics="true" pageId="21" pageNumber="22">Mesosoma</emphasis>
(Fig. 59). Pronotal disc coriaceous-punctate, 0.37 as long as wide, anterior corner rounded, posterior margin almost straight, pronotal disc without short longitudinal furrow. Parapsidal furrow wholly conspicuous. Notauli parallel to each other. Mesoscutum 1.65 length pronotal disc. Scutellum coriaceous-punctate, 1.2 length pronotal disc, almost flat in profile, posterior margin convex medially, scutellar pit elliptical, dilated, distant each other 5.0 its own diameter. Dorsal axillar surface with fovea oval. Metanotum with median region high, coriaceous-puncticulate, as high as scutellum in profile, with fovea oval laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.83 length propodeal disc, incomplete posterad; discal carina 0.23 length propodeal disc, parallel to median carina, straight; space between median and discal carina confused-rugulose. Mesopleuron with subtegular furrow oval; mesepimeral suture wholly conspicuous; mesopleural epicoxal lobe conspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression heart-shaped; median suture evident.
</paragraph>
<paragraph id="8BCE36C68A26FF9813C9FD04FAD2FCB0" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FD04FEEFFD20" box="[199,275,693,716]" italics="true" pageId="21" pageNumber="22">Wings.</emphasis>
Macropterous. Forewing (
<figureCitation id="134A2A438A26FF98114AFD05FD59FD20" box="[580,677,692,716]" captionStart="FIGURES 103112" captionStartId="72.[151,263,1899,1922]" captionTargetBox="[151,1436,193,1878]" captionTargetId="figure-77@72.[151,1436,193,1878]" captionTargetPageId="72" captionText="FIGURES 103112. Right forewing. 103. Eupsenella araba sp. nov. 104. E. arabana sp. nov. 105. E. baada sp. nov. 106. E. barada sp. nov. 107. E. barna sp. nov. 108. E. batjala sp. nov. 109. E. bilingara sp. nov. 110. E. bugulmara sp. nov. 111. E." figureDoi="http://doi.org/10.5281/zenodo.10096777" httpUri="https://zenodo.org/record/10096777/files/figure.png" pageId="21" pageNumber="22">Fig. 104</figureCitation>
) with 1M cell subtriangular; 1M and 1R1 cells aligned basally; C cell widening distally;1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs
<emphasis id="B905EAD48A26FF98178BFD69FB6FFD1C" box="[1157,1171,728,752]" italics="true" pageId="21" pageNumber="22">b</emphasis>
straight; 2R1 cell 0.16 LFW; distal margin of Rs
<emphasis id="B905EAD48A26FF9812B0FD4CFE36FCF8" box="[446,458,765,788]" italics="true" pageId="21" pageNumber="22">c</emphasis>
convex; r-rs vein inclined; R1
<emphasis id="B905EAD48A26FF981024FD4DFCC4FCF8" box="[810,824,764,788]" italics="true" pageId="21" pageNumber="22">b</emphasis>
entirely on anterior margin of forewing; Cu
<emphasis id="B905EAD48A26FF98163BFD4DFABFFCF8" box="[1333,1347,764,788]" italics="true" pageId="21" pageNumber="22">b</emphasis>
absent; Rs+M vein 1.5 Rs
<emphasis id="B905EAD48A26FF981265FC90FE85FCD4" box="[363,377,801,824]" italics="true" pageId="21" pageNumber="22">a</emphasis>
; M vein 1.4 Rs
<emphasis id="B905EAD48A26FF98113CFC90FDBCFCD4" box="[562,576,801,824]" italics="true" pageId="21" pageNumber="22">a</emphasis>
; Rs
<emphasis id="B905EAD48A26FF981161FC90FD81FCD4" box="[623,637,801,824]" italics="true" pageId="21" pageNumber="22">a</emphasis>
subperpendicular to anterior margin of forewing; stigma rectangular; distal stigmal margin truncate. Hind wing with four distal hamuli, last hamulus discontinuous each other.
</paragraph>
<paragraph id="8BCE36C68A26FF9813C9FCD8FCEFFC48" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FCD8FEFFFC6C" box="[199,259,873,896]" italics="true" pageId="21" pageNumber="22">Legs.</emphasis>
Profemur swollen, 0.4 as wide as long. Metacoxa with median region 0.53 as wide as long. Metacoxal projection in dorsal region conspicuous, apex sharpened.
</paragraph>
<paragraph id="8BCE36C68A26FF9813C9FC00FE68FB28" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FC00FEB9FC24" box="[199,325,945,968]" italics="true" pageId="21" pageNumber="22">Metasoma.</emphasis>
0.27 as wide as long, with few setae, ventral carina of petiole present. Hypopygium (
<figureCitation id="134A2A438A26FF98163EFC1EFA6FFC2B" box="[1328,1427,943,967]" captionStart="FIGURES 138144" captionStartId="76.[151,263,1495,1518]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure-60@76.[151,1436,193,1474]" captionTargetPageId="76" captionText="FIGURES 138144. Hypopygium. 138. Eupsenella insulana male. 139. E. ajabatha sp. nov. 140. E. alawa sp. nov. 141. E. alura sp. nov. 142. E. antakirinja sp. nov. 143. E. araba sp. nov. 144. E. arabana sp. nov. 6. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096785" httpUri="https://zenodo.org/record/10096785/files/figure.png" pageId="21" pageNumber="22">Fig. 144</figureCitation>
) with posterior margin strongly angulate; teeth of posterior margin straight, teeth distant each other, almost evenly thick; lateral stalk present, but almost vestigial. Genitalia (
<figureCitation id="134A2A438A26FF9810AFFC46FBBFFBE3" box="[929,1091,1015,1039]" captionStart="FIGURES 163165" captionStartId="79.[151,264,868,891]" captionTargetBox="[156,1420,191,840]" captionTargetId="figure-41@79.[151,1436,165,847]" captionTargetPageId="79" captionText="FIGURES 163165. Male genitalia. 163165 Eupsenella arabana sp. nov.; 163. Ventral view; 164. Dorsal view; 165. Lateral view. (Scale bar = 250 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096791" httpUri="https://zenodo.org/record/10096791/files/figure.png" pageId="21" pageNumber="22">Figs 163165</figureCitation>
) with paramere doble, apical margin in lateral view without concavity, inner margin of ventral arm not angulate in lateral view; basiparamere with apical margin angulate medially; volsella aligned with aedeagus apex; cuspis 0.36 length of apical lobe of aedeagus, 1.40 width of apical lobe of aedeagus; digitus 0.50 length of apical lobe of aedeagus, upper surface spinose; aedeagus with median region as wide as base; apical lobe 0.45 aedeagus length, fold of inner margin present; apex straight.
</paragraph>
<paragraph id="8BCE36C68A26FF9813C9FB7EFE47FB0B" blockId="21.[151,1437,151,1940]" box="[199,443,1231,1255]" pageId="21" pageNumber="22">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A26FF9813C9FB42FB71FAC3" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BCE36C68A26FF9813C9FB42FB71FAC3" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FB42FE47FAE0" bold="true" box="[199,443,1267,1292]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Material examined.</emphasis>
<materialsCitation id="3B193C9B8A26FF9812C2FB45FB71FAC3" collectionCode="ANIC" collectorName="A. Walford-Huggins" country="Australia" location="Black Mountain" municipality="Mountain Road" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B905EAD48A26FF9812C2FB45FDCFFAE0" box="[460,563,1268,1292]" italics="true" pageId="21" pageNumber="22">
<typeStatus id="54CA88648A26FF9812C2FB45FDCFFAE0" box="[460,563,1268,1292]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D77FD4F8A26FF98114AFB45FDA5FAE7" box="[580,601,1268,1291]" pageId="21" pageNumber="22" type="male"></specimenCount>
.
<collectingCountry id="F36676568A26FF98117FFB45FCEFFAE0" box="[625,787,1268,1292]" name="Australia" pageId="21" pageNumber="22">AUSTRALIA</collectingCountry>
, Q[ueens]L[an]D,
<location id="8EAE601D8A26FF9810F4FB45FB46FAE0" LSID="urn:lsid:plazi:treatment:03D887D08A27FF981399FACBFA60F878:8EAE601D8A26FF9810F4FB45FB46FAE0" box="[1018,1210,1268,1292]" country="Australia" municipality="Mountain Road" name="Black Mountain" pageId="21" pageNumber="22">
Black
<collectingMunicipality id="6BAAACBC8A26FF981743FB45FB46FAE0" box="[1101,1210,1268,1292]" pageId="21" pageNumber="22">Mountain</collectingMunicipality>
</location>
Road, Julatten, 5 Sept[ember]10 Oct[ober] 1987,
<collectorName id="268453108A26FF981102FAA6FD08FAC3" box="[524,756,1303,1327]" pageId="21" pageNumber="22">A. Walford-Huggins</collectorName>
, rainf[orest], interc[ept] trap,
<collectionCode id="ED60AE038A26FF98174FFAA6FB75FAC3" box="[1089,1161,1303,1327]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="22">ANIC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A26FF9813C9FA8AFD8FFAB8" box="[199,627,1339,1364]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BCE36C68A26FF9813C9FA8AFD8FFAB8" blockId="21.[151,1437,151,1940]" box="[199,627,1339,1364]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FA8AFE9CFAB8" bold="true" box="[199,352,1339,1364]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Distribution.</emphasis>
<collectingCountry id="F36676568A26FF981269FA8DFE32FAB8" box="[359,462,1340,1364]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
(
<collectingRegion id="49B5F8248A26FF9812D3FA8DFD94FAB8" box="[477,616,1340,1364]" country="Australia" name="Queensland" pageId="21" pageNumber="22">Queensland</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A26FF9813C9FAD1FA60F878" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BCE36C68A26FF9813C9FAD1FDF2F998" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
<emphasis id="B905EAD48A26FF9813C9FAD1FEC7FA94" bold="true" box="[199,315,1376,1400]" inLineHeading="true" pageId="21" pageNumber="22" reason="1">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C714D458A26FF981152FAD0FD20FA94" authorityName="Gordh &amp; Harris" authorityYear="1996" box="[604,732,1376,1400]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B905EAD48A26FF981152FAD0FD20FA94" box="[604,732,1376,1400]" italics="true" pageId="21" pageNumber="22">E. insulana</emphasis>
</taxonomicName>
by having the median clypeal lobe angulate, the median carina of propodeal disc present, the hind wing with four distal hamuli, the posterior margin of hypopygium angulate, but it differs from it by having the propodeal disc discal carina absent, the 1M cell of forewing triangular, the Rs
<emphasis id="B905EAD48A26FF981660FA19FA80FA2C" box="[1390,1404,1448,1472]" italics="true" pageId="21" pageNumber="22">b</emphasis>
of forewing aligned with Cu
<emphasis id="B905EAD48A26FF9812B8FA7CFE38FA08" box="[438,452,1485,1508]" italics="true" pageId="21" pageNumber="22">a</emphasis>
, the anterior margin of basiparamere angulate medially, the teeth of posterior margin of hypopygium slightly narrowed toward apex, whereas
<taxonomicName id="4C714D458A26FF981005FA40FC73F9E4" authorityName="Gordh &amp; Harris" authorityYear="1996" box="[779,911,1520,1544]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B905EAD48A26FF981005FA40FC73F9E4" box="[779,911,1520,1544]" italics="true" pageId="21" pageNumber="22">E. insulana</emphasis>
</taxonomicName>
has with the discal carina of propodeal disc present, but almost inconspicuous, the 1M cell of forewing subpentagonal, the Rs
<emphasis id="B905EAD48A26FF981733F9A5FBB7F9C0" box="[1085,1099,1556,1580]" italics="true" pageId="21" pageNumber="22">b</emphasis>
of forewing not aligned with Cu
<emphasis id="B905EAD48A26FF9813B9F988FF39F9BC" box="[183,197,1593,1616]" italics="true" pageId="21" pageNumber="22">a</emphasis>
, the anterior margin of basiparamere not angulate medially in lateral view, the teeth of posterior margin of hypopygium almost evenly thick.
</paragraph>
<paragraph id="8BCE36C68A26FF9813C9F931FA60F878" blockId="21.[151,1437,151,1940]" pageId="21" pageNumber="22">
It is also similar to
<taxonomicName id="4C714D458A26FF9812A2F930FDB6F974" box="[428,586,1664,1688]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="antakirinja" status="sp. nov.">
<emphasis id="B905EAD48A26FF9812A2F930FDB6F974" box="[428,586,1664,1688]" italics="true" pageId="21" pageNumber="22">E. antakirinja</emphasis>
</taxonomicName>
<emphasis id="B905EAD48A26FF98115AF931FD4CF974" bold="true" box="[596,688,1664,1688]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A23657AF8A26FF98115AF931FD4CF974" box="[596,688,1664,1688]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
because both species present the eyes large and protuberant, the ocelli large, vertex crest straight, the median clypeal lobe angulate, the median carina of median clypeal lobe present and elevated, the median carina of propodeal disc long, the Rs
<emphasis id="B905EAD48A26FF9810BEF979FC42F90C" box="[944,958,1736,1760]" italics="true" pageId="21" pageNumber="22">b</emphasis>
of forewing aligned basally with Cu
<emphasis id="B905EAD48A26FF981653F978FA97F90C" box="[1373,1387,1737,1760]" italics="true" pageId="21" pageNumber="22">a</emphasis>
, the genitalia with paramere doble. However this species has the area between the anterior margin of eyes and demarkation of median clypeal lobe short, the teeth of posterior margin of hypopygium almost evenly thick, the inner margin of ventral arm not angulate, the digitus long, the apical lobe of aedeagus long, the apical lobe of aedeagus with fold in inner margin present, whereas
<taxonomicName id="4C714D458A26FF98100BF8E8FC59F89C" box="[773,933,1880,1904]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="antakirinja" status="sp. nov.">
<emphasis id="B905EAD48A26FF98100BF8E8FC59F89C" box="[773,933,1880,1904]" italics="true" pageId="21" pageNumber="22">E. antakirinja</emphasis>
</taxonomicName>
<emphasis id="B905EAD48A26FF9810BEF8E9FBF1F89C" bold="true" box="[944,1037,1880,1904]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A23657AF8A26FF9810BEF8E9FBF1F89C" box="[944,1037,1880,1904]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the area between the anterior margin of eyes and demarkation of median clypeal lobe long, the teeth of posterior margin of hypopigium
</paragraph>
</subSubSection>
</treatment>
</document>