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<document id="ADD091510CA0031DBB13D8ECA1C9C58F" ID-DOI="10.11646/zootaxa.3539.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10096759" ID-ZooBank="0564EF02-FCF9-49A8-BDAC-F432876C16B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629821597637" checkinUser="felipe" docAuthor="Ramos, Magno S. &amp; Azevedo, Celso O." docDate="2012" docId="03D887D08A02FFBF1399F986FC21FA53" docLanguage="en" docName="zt03539p080.pdf" docOrigin="Zootaxa 3539 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3539.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eupsenella kokatha Ramos &amp; Azevedo, sp. nov." docType="treatment" docVersion="4" lastPageNumber="51" masterDocId="FFE1FFA88A33FF8D130EFFB1FFFCFFEC" masterDocTitle="Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="50" updateTime="1699581041301" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B68DCA91466F2AFE2D0E57FA412A8D7A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DD784C9A02A1A4C857EDB8FCEEC064F6">Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae)</mods:title>
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<mods:name id="94DEE075AEA7E3899DF69836648373A9" type="personal">
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<mods:namePart id="F8F9503423D44B2D36C0429C00940B3F">Ramos, Magno S.</mods:namePart>
</mods:name>
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<mods:namePart id="080C7D68FEEC5CCCA1EFF2A61FD5553C">Azevedo, Celso O.</mods:namePart>
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<mods:title id="E0835183183BC0F15ED7868C90F0835B">Zootaxa</mods:title>
</mods:titleInfo>
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<mods:date id="75F210316AC4CF581BF292BDBAF43D02">2012</mods:date>
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<mods:number id="FD2A6E33CCA8B4BB54E4B9D4C729386A">2012-11-05</mods:number>
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<mods:number id="17EE8ECDE806E6E0BC593485F5FE104F">3539</mods:number>
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</mods:location>
<mods:classification id="443BA3026A2D3A4CDB2FA4B7F8B8B373">journal article</mods:classification>
<mods:identifier id="0E524BB28E24267ACC8AC9262148E693" type="DOI">10.11646/zootaxa.3539.1.1</mods:identifier>
<mods:identifier id="A2995D143953B114ABCB00F2188ECEBD" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="C41E157EE2B3CB843572B9AD81799DA8" type="Zenodo-Dep">10096759</mods:identifier>
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<treatment id="03D887D08A02FFBF1399F986FC21FA53" ID-DOI="http://doi.org/10.5281/zenodo.5461881" ID-Zenodo-Dep="5461881" LSID="urn:lsid:plazi:treatment:03D887D08A02FFBF1399F986FC21FA53" httpUri="http://treatment.plazi.org/id/03D887D08A02FFBF1399F986FC21FA53" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<subSubSection id="C36B654D8A02FFBC1399F986FD1DF9BD" box="[151,737,1591,1617]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="8BCE36C68A02FFBC1399F986FD1DF9BD" blockId="49.[151,737,1591,1650]" box="[151,737,1591,1617]" pageId="49" pageNumber="50">
<heading id="D08681AA8A02FFBC1399F986FD1DF9BD" bold="true" box="[151,737,1591,1617]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="B905EAD48A02FFBC1399F986FD1DF9BD" bold="true" box="[151,737,1591,1617]" pageId="49" pageNumber="50">
<taxonomicName id="4C714D458A02FFBC1399F986FD8AF9BD" authority="Ramos &amp; Azevedo" authorityName="Ramos &amp; Azevedo" box="[151,630,1591,1617]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="kokatha" status="sp. nov.">
<emphasis id="B905EAD48A02FFBC1399F986FE7BF9BD" bold="true" box="[151,391,1591,1617]" italics="true" pageId="49" pageNumber="50">Eupsenella kokatha</emphasis>
Ramos &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A23657AF8A02FFBC118DF986FD1DF9BD" box="[643,737,1591,1617]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A02FFBF1399F9EBFE66FB0B" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
<paragraph id="8BCE36C68A02FFBC1399F9EBFEA2F99E" blockId="49.[151,737,1591,1650]" box="[151,350,1626,1650]" pageId="49" pageNumber="50">
(
<figureCitation id="134A2A438A02FFBC1391F9EBFF09F99E" box="[159,245,1626,1650]" captionStart="FIGURES 39–47" captionStartId="65.[151,264,1445,1468]" captionTargetBox="[151,1436,193,1424]" captionTargetId="figure-71@65.[151,1436,193,1424]" captionTargetPageId="65" captionText="FIGURES 39–47. Head in dorsal view. 39. Eupsenella kokatha sp. nov. 40. E. larrakia sp. nov. 41. E. maya sp. nov. 42. E. malgana sp. nov. 43. E. nanda sp. nov. 44. E. nangatara sp. nov. 45. E. pangkala sp. nov. 46. E. wanamara sp. nov. 47. E. agilis. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096769" httpUri="https://zenodo.org/record/10096769/files/figure.png" pageId="49" pageNumber="50">Figs 39</figureCitation>
,
<figureCitation id="134A2A438A02FFBC120FF9EBFEE3F99E" box="[257,287,1626,1650]" captionStart="FIGURES 84–92" captionStartId="70.[151,264,1931,1954]" captionTargetBox="[161,1425,181,1910]" captionTargetId="figure-72@70.[161,1425,181,1910]" captionTargetPageId="70" captionText="FIGURES 84–92. Mesosoma in dorsal view. 84. Eupsenella kokatha sp. nov. 85. E. larrakia sp. nov. 86. E. maya sp. nov. 87." figureDoi="http://doi.org/10.5281/zenodo.10096773" httpUri="https://zenodo.org/record/10096773/files/figure.png" pageId="49" pageNumber="50">84</figureCitation>
,
<figureCitation id="134A2A438A02FFBC1225F9EBFEAAF99E" box="[299,342,1626,1650]" captionStart="FIGURES 123–132" captionStartId="74.[151,263,1911,1934]" captionTargetBox="[151,1436,193,1890]" captionTargetId="figure-77@74.[151,1436,193,1890]" captionTargetPageId="74" captionText="FIGURES 123–132. Right forewing. 123. Eupsenella jagara sp. nov. 124. E. janda sp. nov. 125. E. kabalbara sp. nov. 126. E." figureDoi="http://doi.org/10.5281/zenodo.10096781" httpUri="https://zenodo.org/record/10096781/files/figure.png" pageId="49" pageNumber="50">129</figureCitation>
)
</paragraph>
<paragraph id="8BCE36C68A02FFBC1399F913FC24F956" blockId="49.[151,1437,1698,1938]" box="[151,984,1698,1723]" pageId="49" pageNumber="50">
<emphasis id="B905EAD48A02FFBC1399F913FED4F957" bold="true" box="[151,296,1698,1723]" inLineHeading="true" pageId="49" pageNumber="50" reason="1">Description.</emphasis>
FEMALE (
<typeStatus id="54CA88648A02FFBC12A0F913FDEBF956" box="[430,535,1698,1722]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
). Body length
<quantity id="4C899B238A02FFBC11B7F913FCDCF956" box="[697,800,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="7.29" pageId="49" pageNumber="50" unit="mm" value="7.29">7.29 mm</quantity>
. LFW
<quantity id="4C899B238A02FFBC1063F913FC28F956" box="[877,980,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.81" pageId="49" pageNumber="50" unit="mm" value="4.81">4.81 mm</quantity>
.
</paragraph>
<paragraph id="8BCE36C68A02FFBC13C9F977FBB3F8A6" blockId="49.[151,1437,1698,1938]" pageId="49" pageNumber="50">
<emphasis id="B905EAD48A02FFBC13C9F977FEB5F932" box="[199,329,1734,1758]" italics="true" pageId="49" pageNumber="50">Coloration.</emphasis>
Head dark castaneous almost black; scape dark castaneous; pedicel and flagellum castaneous; mandible with body almost dark castaneous and only apex castaneous; palpi light castaneous; mesosoma dark castaneous almost black; tegula and wing venation light castaneous; procoxa dark castaneous; mesocoxa, metacoxa, trochanters, castaneous; tarsi light castaneous; metasoma dark castaneous.
</paragraph>
<paragraph id="8BCE36C68A02FFBF13C9F8E7FE02FE23" blockId="49.[151,1437,1698,1938]" lastBlockId="50.[151,1437,151,1472]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="B905EAD48A02FFBC13C9F8E7FEF8F882" box="[199,260,1878,1902]" italics="true" pageId="49" pageNumber="50">Head</emphasis>
(
<figureCitation id="134A2A438A02FFBC1218F8E7FE96F882" box="[278,362,1878,1902]" captionStart="FIGURES 39–47" captionStartId="65.[151,264,1445,1468]" captionTargetBox="[151,1436,193,1424]" captionTargetId="figure-71@65.[151,1436,193,1424]" captionTargetPageId="65" captionText="FIGURES 39–47. Head in dorsal view. 39. Eupsenella kokatha sp. nov. 40. E. larrakia sp. nov. 41. E. maya sp. nov. 42. E. malgana sp. nov. 43. E. nanda sp. nov. 44. E. nangatara sp. nov. 45. E. pangkala sp. nov. 46. E. wanamara sp. nov. 47. E. agilis. (Scale bar = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.10096769" httpUri="https://zenodo.org/record/10096769/files/figure.png" pageId="49" pageNumber="50">Fig. 39</figureCitation>
). Head as long as wide, subtriangular in profile; gena strongly concave in profile. Mandible evenly wide, two lowermost teeth larger than upper ones, lower tooth curved inward, lower margin strongly setose. Antenna
<quantity id="4C899B238A01FFBF13F0FF26FE9BFF43" box="[254,359,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" pageId="50" pageNumber="51" unit="mm" value="2.53">2.53 mm</quantity>
; first five antennomeres in ratio of about 19:19:17:15:15; flagellomere 1 1.7 longer than wide, flagellar pubescence dense, with many outstanding erect setae. Region between torulus and eye concave. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.06 LH. Eye protuberant, 0.61 LH, with dense hairs, located more laterally, lower margin without strong depression. Frons coriaceous punctate. WH 1.0 LH. WF 0.58 WH. WF 1.0 HE. OOL 0.89 WOT. VOL 0.26 HE. Distance from posterior ocellus to vertex crest 0.5 DAO. Ocelli large, DAO 0.12 WF, ocellar triangle not compact, distance between posterior ocelli 2.4 DAO, anterior ocellus surpassing imaginary top eye line, posterior ocellus distant from vertex. Vertex straight, long setae, 0.18 LH, temple profile slightly convergent in dorsal view, with corner angulate, not projected postero-laterad. Hipostomal carina arched, high.
</paragraph>
<paragraph id="8BCE36C68A01FFBF13C9FE6CFE6AFC48" blockId="50.[151,1437,151,1472]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FE6CFEC3FE18" box="[199,319,477,500]" italics="true" pageId="50" pageNumber="51">Mesosoma</emphasis>
(
<figureCitation id="134A2A438A01FFBF125BFE6DFE52FE18" box="[341,430,476,500]" captionStart="FIGURES 84–92" captionStartId="70.[151,264,1931,1954]" captionTargetBox="[161,1425,181,1910]" captionTargetId="figure-72@70.[161,1425,181,1910]" captionTargetPageId="70" captionText="FIGURES 84–92. Mesosoma in dorsal view. 84. Eupsenella kokatha sp. nov. 85. E. larrakia sp. nov. 86. E. maya sp. nov. 87." figureDoi="http://doi.org/10.5281/zenodo.10096773" httpUri="https://zenodo.org/record/10096773/files/figure.png" pageId="50" pageNumber="51">Fig. 84</figureCitation>
). Pronotal disc coriaceous-punctate, 0.44 as long as wide, anterior corner rounded, posterior margin concave medially, pronotal disc without short longitudinal furrow. Parapsidal furrow inconspicuous. Notauli convergent posterad. Mesoscutum 1.53 length pronotal disc. Scutellum coriaceouspunctate, 1.27 length pronotal disc, almost flat in profile, posterior margin convex medially, scutellar pit elliptical, narrowed, distant each other 14.0 its own diameter, scutellar furrow absent. Dorsal axillar surface with fovea subrectangular. Metanotum with median region high, coriaceous-puncticulate, as high as scutellum in profile, with fovea elliptical laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.95 length propodeal disc, complete posterad; discal carina 0.1 length propodeal disc, converging posterad, slightly arched; space between median and discal carina confused-rugulose. Mesopleuron with subtegular furrow oval; mesepimeral suture almost inconspicuous posterad; mesopleural epicoxal lobe inconspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression heart-shaped; median suture evident.
</paragraph>
<paragraph id="8BCE36C68A01FFBF13C9FC01FCA9FBBB" blockId="50.[151,1437,151,1472]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FC01FEEFFC2B" box="[199,275,944,967]" italics="true" pageId="50" pageNumber="51">Wings.</emphasis>
Macropterous. Forewing (
<figureCitation id="134A2A438A01FFBF1135FC1EFD66FC2B" box="[571,666,943,967]" captionStart="FIGURES 123–132" captionStartId="74.[151,263,1911,1934]" captionTargetBox="[151,1436,193,1890]" captionTargetId="figure-77@74.[151,1436,193,1890]" captionTargetPageId="74" captionText="FIGURES 123–132. Right forewing. 123. Eupsenella jagara sp. nov. 124. E. janda sp. nov. 125. E. kabalbara sp. nov. 126. E." figureDoi="http://doi.org/10.5281/zenodo.10096781" httpUri="https://zenodo.org/record/10096781/files/figure.png" pageId="50" pageNumber="51">Fig. 129</figureCitation>
) with 1M cell subtriangular; 1M and 1R1 cells not aligned basally; C cell widening distally; 1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs
<emphasis id="B905EAD48A01FFBF1789FC65FB69FC00" box="[1159,1173,980,1004]" italics="true" pageId="50" pageNumber="51">b</emphasis>
straight; 2R1 cell 0.15 LFW; distal margin of Rs
<emphasis id="B905EAD48A01FFBF12B5FC48FE3BFBFC" box="[443,455,1017,1040]" italics="true" pageId="50" pageNumber="51">c</emphasis>
convex; r-rs vein inclined; R1
<emphasis id="B905EAD48A01FFBF102DFC49FCCDFBFC" box="[803,817,1016,1040]" italics="true" pageId="50" pageNumber="51">b</emphasis>
arching distally; Cu
<emphasis id="B905EAD48A01FFBF1717FC49FBDBFBFC" box="[1049,1063,1016,1040]" italics="true" pageId="50" pageNumber="51">b</emphasis>
absent; Rs+M vein 0.85 Rs
<emphasis id="B905EAD48A01FFBF166BFC48FA8FFBFC" box="[1381,1395,1017,1040]" italics="true" pageId="50" pageNumber="51">a</emphasis>
; M vein 1.0 Rs
<emphasis id="B905EAD48A01FFBF1214FBACFED4FBD8" box="[282,296,1053,1076]" italics="true" pageId="50" pageNumber="51">a</emphasis>
; Rs
<emphasis id="B905EAD48A01FFBF1259FBACFE99FBD8" box="[343,357,1053,1076]" italics="true" pageId="50" pageNumber="51">a</emphasis>
slightly inclined toward tegula; stigma rectangular; distal stigmal margin truncate. Hind wing with four distal hamuli, last hamulus discontinuous each other.
</paragraph>
<paragraph id="8BCE36C68A01FFBF13C9FBD4FD02FB73" blockId="50.[151,1437,151,1472]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FBD4FEFFFB90" box="[199,259,1125,1148]" italics="true" pageId="50" pageNumber="51">Legs.</emphasis>
Profemur swollen, 0.48 as wide as long. Metacoxa with median region 0.45 as wide as long. Metacoxal projection in dorsal region conspicuous, apex rounded.
</paragraph>
<paragraph id="8BCE36C68A01FFBF13C9FB1CFB47FB2F" blockId="50.[151,1437,151,1472]" box="[199,1211,1196,1220]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FB1CFEBFFB28" box="[199,323,1197,1220]" italics="true" pageId="50" pageNumber="51">Metasoma.</emphasis>
0.33 as wide as long, except first two tergites, ventral carina of petiole present.
</paragraph>
<paragraph id="8BCE36C68A01FFBF13C9FB7EFE66FB0B" blockId="50.[151,1437,151,1472]" box="[199,410,1231,1255]" pageId="50" pageNumber="51">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A01FFBF13C9FB42FC7DFAC3" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="8BCE36C68A01FFBF13C9FB42FC7DFAC3" blockId="50.[151,1437,151,1472]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FB42FE45FAE0" bold="true" box="[199,441,1267,1292]" inLineHeading="true" pageId="50" pageNumber="51" reason="1">Material examined.</emphasis>
<materialsCitation id="3B193C9B8A01FFBF12C7FB45FC7DFAC3" collectionCode="ANIC" collectorName="Rockpool George, M &amp; H. Frauca" country="Australia" location="AUSTRALIA" pageId="50" pageNumber="51" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B905EAD48A01FFBF12C7FB45FDCCFAE0" box="[457,560,1268,1292]" italics="true" pageId="50" pageNumber="51">
<typeStatus id="54CA88648A01FFBF12C7FB45FDCCFAE0" box="[457,560,1268,1292]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D77FD4F8A01FFBF1131FB45FDA8FAE7" box="[575,596,1268,1291]" pageId="50" pageNumber="51" type="female">♀</specimenCount>
.
<collectingCountry id="F36676568A01FFBF1165FB45FCF0FAE0" box="[619,780,1268,1292]" name="Australia" pageId="50" pageNumber="51">AUSTRALIA</collectingCountry>
, Q[ueensland],
<collectorName id="268453108A01FFBF10C0FB45FB32FAE0" box="[974,1230,1268,1292]" pageId="50" pageNumber="51">Rockpool George, M</collectorName>
[oun]t[ain] Walah Nat[ional] P[ar]k, Biggenden, 22 Aug[ust] 1972,
<collectorName id="268453108A01FFBF11CEFAA9FCD6FAC3" box="[704,810,1304,1327]" pageId="50" pageNumber="51">H.Frauca</collectorName>
,
<collectionCode id="ED60AE038A01FFBF103BFAA6FC81FAC3" box="[821,893,1303,1327]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="50" pageNumber="51">ANIC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A01FFBF13C9FA8AFD8FFAB8" box="[199,627,1339,1364]" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BCE36C68A01FFBF13C9FA8AFD8FFAB8" blockId="50.[151,1437,151,1472]" box="[199,627,1339,1364]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FA8AFE9CFAB8" bold="true" box="[199,352,1339,1364]" inLineHeading="true" pageId="50" pageNumber="51" reason="1">Distribution.</emphasis>
<collectingCountry id="F36676568A01FFBF1269FA8DFE32FAB8" box="[359,462,1340,1364]" name="Australia" pageId="50" pageNumber="51">Australia</collectingCountry>
(
<collectingRegion id="49B5F8248A01FFBF12D3FA8DFD94FAB8" box="[477,616,1340,1364]" country="Australia" name="Queensland" pageId="50" pageNumber="51">Queensland</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B654D8A01FFBF13C9FAD1FC21FA53" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8BCE36C68A01FFBF13C9FAD1FC21FA53" blockId="50.[151,1437,151,1472]" pageId="50" pageNumber="51">
<emphasis id="B905EAD48A01FFBF13C9FAD1FEC7FA94" bold="true" box="[199,315,1376,1400]" inLineHeading="true" pageId="50" pageNumber="51" reason="1">Remarks.</emphasis>
This species is very different of the others by having the vertex crest clearly narrow, the ocelli distant, the eyes large, protuberant, the median clypeal lobe angulate, high, the discal carina of propodeal disc inconspicuous, the Rs
<emphasis id="B905EAD48A01FFBF1282FA19FE66FA2C" box="[396,410,1448,1472]" italics="true" pageId="50" pageNumber="51">b</emphasis>
of forewing straight, the Cu
<emphasis id="B905EAD48A01FFBF11D9FA19FD19FA2C" box="[727,741,1448,1472]" italics="true" pageId="50" pageNumber="51">b</emphasis>
of forewing vestigial.
</paragraph>
</subSubSection>
</treatment>
</document>
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