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<document id="4E774EB3977D931E5F4E04B58EA4AB50" ID-CLB-Dataset="9257" ID-DOI="10.5252/zoosystema2018v40a23" ID-GBIF-Dataset="c35887df-f50e-4c13-8b29-5f23d0e1df75" ID-ISSN="1638-9387" ID-Zenodo-Dep="4336783" ID-ZooBank="urn:lsid:zoobank.org:pub:50057A30-3C33-49A5-9E23-9658C23D2EDD" IM.bibliography_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1608156498336" checkinUser="felipe" docAuthor="Dubois, Alain &amp; Ohler, Annemarie" docDate="2018" docId="03D887C75D05FFA9648755C3FD4DF8C7" docLanguage="en" docName="Zoosystema.40.23.501-506.pdf" docOrigin="Zoosystema 40 (23)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Hyla prasina Burmeister 1856" docType="treatment" docVersion="9" lastPageNumber="502" masterDocId="FFE1FFBF5D06FFAA6403551BFFE6FFCB" masterDocTitle="The Hyla quoyi-Hyla prasina case (Amphibia, Anura), with comments on bibliographic and taxonomic databases and on Article 23.9 of the Code" masterLastPageNumber="506" masterPageNumber="501" pageNumber="502" updateTime="1688567976675" updateUser="tatiana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F5398292767013DE8ADF3C49CE641DED">The Hyla quoyi-Hyla prasina case (Amphibia, Anura), with comments on bibliographic and taxonomic databases and on Article 23.9 of the Code</mods:title>
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<mods:namePart id="3A063E11D29622C5ABA86E58ADC4B55D">Dubois, Alain</mods:namePart>
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<mods:namePart id="20249E7B05396EB93CE2E6E286AF24FD">Ohler, Annemarie</mods:namePart>
<mods:affiliation id="B303E96DE9DA1AA1F0D9E9FF7A4B532E">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE, case postale 30, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) adubois @ mnhn. fr annemarie. ohler @ mnhn. fr</mods:affiliation>
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<subSubSection id="C36B655A5D05FFA9648755C3FE9CFF39" box="[132,378,216,242]" pageId="3" pageNumber="502" type="multiple">
<paragraph id="8BCE36D15D05FFA9648755C3FE9CFF39" blockId="3.[132,641,215,242]" box="[132,378,216,242]" pageId="3" pageNumber="502">
THE
<emphasis id="B905EAC35D05FFA964C555C3FE9CFF39" box="[198,378,216,242]" italics="true" pageId="3" pageNumber="502">
<taxonomicName id="4C714D525D05FFA964C555C3FE88FF39" ID-CoL="3ND8R" authorityName="Bory de Saint-Vincent" authorityYear="1828" box="[198,366,216,242]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="quoyi">HYLA QUOYI</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36B655A5D05FFA9657955C3FD67FF3A" box="[378,641,215,242]" pageId="3" pageNumber="502" type="nomenclature">
<paragraph id="8BCE36D15D05FFA9657955C3FD67FF3A" blockId="3.[132,641,215,242]" box="[378,641,215,242]" pageId="3" pageNumber="502">
<heading id="D08681BD5D05FFA9657955C3FD67FF3A" box="[378,641,215,242]" fontSize="11" level="1" pageId="3" pageNumber="502" reason="1">
<taxonomicName id="4C714D525D05FFA9657955C3FDD0FF39" ID-CoL="3ND7L" authorityName="Burmeister" authorityYear="1856" box="[378,566,216,242]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA9657955C3FDD0FF39" box="[378,566,216,242]" italics="true" pageId="3" pageNumber="502">HYLA PRASINA</emphasis>
</taxonomicName>
CASE
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B655A5D05FFA964875403FD4DF8C7" pageId="3" pageNumber="502" type="description">
<paragraph id="8BCE36D15D05FFA964875403FCE1FE1B" blockId="3.[130,777,279,1804]" pageId="3" pageNumber="502">
Bory de
<collectingCountry id="F36676415D05FFA964E7540CFE9CFEFA" box="[228,378,279,305]" name="Saint Vincent and the Grenadines" pageId="3" pageNumber="502">Saint-Vincent</collectingCountry>
(1828) introduced the nomen
<taxonomicName id="4C714D525D05FFA966D7540CFF5AFE9A" authorityName="Bory de Saint-Vincent" authorityYear="1828" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="quoyi">
<emphasis id="B905EAC35D05FFA966D7540CFF5AFE9A" italics="true" pageId="3" pageNumber="502">Hyla quoyi</emphasis>
</taxonomicName>
for a new frog species from the vicinity of
<collectingRegion id="49B5F8335D05FFA9669A542CFF5AFEBA" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="502">Rio de Janeiro</collectingRegion>
(southeastern
<collectingCountry id="F36676415D05FFA9655B544CFE7EFEB9" box="[344,408,343,370]" name="Brazil" pageId="3" pageNumber="502">Brazil</collectingCountry>
) in a plate of the
<emphasis id="B905EAC35D05FFA966535443FE63FE5A" italics="true" pageId="3" pageNumber="502">Dictionnaire classique dHistoire naturelle</emphasis>
. This book, and the new nomina it contained, were ignored by all subsequent authors, starting with
<bibRefCitation id="EFE04B205D05FFA964BF54ADFE3CFE1A" author="DUMERIL A. - M. - C. &amp; BIBRON G." box="[188,474,438,465]" pageId="3" pageNumber="502" pagination="1 - 792" refId="ref5938" refString="DUMERIL A. - M. - C. &amp; BIBRON G. 1841. - Erpetologie generale ou Histoire naturelle complete des Reptiles. Tome 8. Roret, Paris: [i-iii] + i-ii + 1 - 792." type="book chapter" year="1841">Duméril &amp; Bibron (1841)</bibRefCitation>
, for one century and a half.
</paragraph>
<paragraph id="8BCE36D15D05FFA9649854CDFD2FFC84" blockId="3.[130,777,279,1804]" pageId="3" pageNumber="502">
<bibRefCitation id="EFE04B205D05FFA9649854CDFE7DFE3A" author="BURMEISTER H." box="[155,411,470,497]" pageId="3" pageNumber="502" pagination="1 - 115" refId="ref5118" refString="BURMEISTER H. 1856. - Erlauterungen zur Fauna Brasiliens, enthaltend Abbildungen und ausfuhrliche Beschreibungen neuer oder ungenugend bekannter Thier-Arten. Georg Reimer, Berlin: i-x + 1 - 115 + 32 plates." type="book chapter" year="1856">Burmeister (1856: 106)</bibRefCitation>
described
<taxonomicName id="4C714D525D05FFA9660C54CDFD75FE3B" authorityName="Burmeister" authorityYear="1856" box="[527,659,470,496]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA9660C54CDFD75FE3B" box="[527,659,470,496]" italics="true" pageId="3" pageNumber="502">Hyla prasina</emphasis>
</taxonomicName>
, also from the region of Rio. This nomen was first synonymised with
<taxonomicName id="4C714D525D05FFA96485570DFDAEFDFB" authority="Dumeril &amp; Bibron, 1841" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[134,584,534,560]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B905EAC35D05FFA96485570DFEC6FDFB" box="[134,288,534,560]" italics="true" pageId="3" pageNumber="502">Hyla pulchella</emphasis>
<bibRefCitation id="EFE04B205D05FFA9652A570DFDAEFDFB" author="DUMERIL A. - M. - C. &amp; BIBRON G." box="[297,584,534,560]" pageId="3" pageNumber="502" pagination="1 - 792" refId="ref5938" refString="DUMERIL A. - M. - C. &amp; BIBRON G. 1841. - Erpetologie generale ou Histoire naturelle complete des Reptiles. Tome 8. Roret, Paris: [i-iii] + i-ii + 1 - 792." type="book chapter" year="1841">Duméril &amp; Bibron, 1841</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFE04B205D05FFA96675570DFEE9FD9B" author="STEINDACHNER F." pageId="3" pageNumber="502" pagination="239 - 288" refId="ref6943" refString="STEINDACHNER F. 1864. - Batrachologische Mittheilungen. Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien 14: 239 - 288." type="journal article" year="1864">Steindachner (1864: 241)</bibRefCitation>
, then resurrected by
<bibRefCitation id="EFE04B205D05FFA965FD572DFD32FD9B" author="BARRIO A." box="[510,724,566,593]" pageId="3" pageNumber="502" pagination="115 - 128" refId="ref4768" refString="BARRIO A. 1965. - Las subespecies de Hyla pulchella Dumeril y Bibron (Anura, Hylidae). Physis 25: 115 - 128." type="journal article" year="1965">Barrio (1965: 117)</bibRefCitation>
as a subspecies of
<taxonomicName id="4C714D525D05FFA96519574DFE56FDBB" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[282,432,598,624]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="B905EAC35D05FFA96519574DFE56FDBB" box="[282,432,598,624]" italics="true" pageId="3" pageNumber="502">Hyla pulchella</emphasis>
</taxonomicName>
and finally reinstated at species rank by
<bibRefCitation id="EFE04B205D05FFA964DD576DFE6BFD5B" author="LUTZ B." box="[222,397,630,656]" pageId="3" pageNumber="502" pagination="1 - 260" refId="ref6496" refString="LUTZ B. - 1973 Brazilian species of Hyla. University of Texas Press, Austin: i-xviii + 1 - 260." type="book chapter" year="1973">Lutz (1973: 83)</bibRefCitation>
. This species was transferred to the genus
<taxonomicName id="4C714D525D05FFA964CB578DFE25FD7B" authority="Wagler, 1830" authorityName="Wagler" authorityYear="1830" box="[200,451,662,688]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="genus">
<emphasis id="B905EAC35D05FFA964CB578DFEC9FD7B" box="[200,303,662,688]" italics="true" pageId="3" pageNumber="502">Hypsiboas</emphasis>
Wagler, 1830
</taxonomicName>
by
<bibRefCitation id="EFE04B205D05FFA965EA578DFCE4FD7B" author="FAIVOVICH J. &amp; HADDAD C. F. B. &amp; GARCIA P. C. A. &amp; FROST D. R. &amp; CAMPBELL J. A. &amp; WHEELER W. C." box="[489,770,661,688]" pageId="3" pageNumber="502" pagination="1 - 240" refId="ref6135" refString="FAIVOVICH J., HADDAD C. F. B., GARCIA P. C. A., FROST D. R., CAMPBELL J. A. &amp; WHEELER W. C. 2005. - Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of natural History 294: 1 - 240." type="journal article" year="2005">
Faivovich
<emphasis id="B905EAC35D05FFA96656578DFD6EFD7B" box="[597,648,662,688]" italics="true" pageId="3" pageNumber="502">et al.</emphasis>
(2005: 88)
</bibRefCitation>
, and the combination
<taxonomicName id="4C714D525D05FFA9656957AEFDCDFD04" baseAuthorityName="Burmeister" baseAuthorityYear="1856" box="[362,555,693,719]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasinus">
<emphasis id="B905EAC35D05FFA9656957AEFDCDFD04" box="[362,555,693,719]" italics="true" pageId="3" pageNumber="502">Hypsiboas prasinus</emphasis>
</taxonomicName>
was universally used in the following years.
<bibRefCitation id="EFE04B205D05FFA9657B57CEFD98FD3B" author="WIENS J. J. &amp; FETZNER J. W. J. &amp; PARKINSON C. L. &amp; REEDER T. W." box="[376,638,725,752]" pageId="3" pageNumber="502" pagination="778 - 807" refId="ref6965" refString="WIENS J. J., FETZNER J. W. J., PARKINSON C. L. &amp; REEDER T. W. 2005. - Hylid frog phylogeny and sampling strategies for speciose clades. Systematic Biology 54: 778 - 807." type="journal article" year="2005">
Wiens
<emphasis id="B905EAC35D05FFA965C057CDFE11FD24" box="[451,503,725,751]" italics="true" pageId="3" pageNumber="502">et al.</emphasis>
(2005: 789)
</bibRefCitation>
showed that the nomen
<taxonomicName id="4C714D525D05FFA9650557EDFE29FCDB" authority="Gray, 1825" authorityName="Gray" authorityYear="1825" box="[262,463,757,784]" class="Amphibia" family="Hylidae" genus="Boana" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="genus">
<emphasis id="B905EAC35D05FFA9650557EDFEAFFCC4" box="[262,329,758,783]" italics="true" pageId="3" pageNumber="502">Boana</emphasis>
Gray, 1825
</taxonomicName>
was available for this genus but did not use the combination
<taxonomicName id="4C714D525D05FFA965F0560DFD6FFCE4" baseAuthorityName="Burmeister" baseAuthorityYear="1856" box="[499,649,790,815]" class="Amphibia" family="Hylidae" genus="Boana" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA965F0560DFD6FFCE4" box="[499,649,790,815]" italics="true" pageId="3" pageNumber="502">Boana prasina</emphasis>
</taxonomicName>
, which was published for the first time by
<bibRefCitation id="EFE04B205D05FFA965CE562EFD22FC84" author="DORIGO T. A. &amp; ROCHA C. F. D." box="[461,708,821,848]" pageId="3" pageNumber="502" pagination="1 - 11" refId="ref5358" refString="DORIGO T. A., VRCIBRADIC, D. &amp; ROCHA C. F. D. 2018. - The amphibians of the state of Rio de Janeiro, Brazil: an updated and commented list. Papeis Avulsos de Zoologia 58: 1 - 11. https: // doi. org / 10.11606 / 1807 - 0205 / 2018.58.05" type="journal article" year="2018">
Dorigo
<emphasis id="B905EAC35D05FFA96621562DFDB0FC84" box="[546,598,821,847]" italics="true" pageId="3" pageNumber="502">et al.</emphasis>
(2018: 3)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BCE36D15D05FFA96498564EFDF6FB05" blockId="3.[130,777,279,1804]" pageId="3" pageNumber="502">
<bibRefCitation id="EFE04B205D05FFA96498564EFEF8FCBB" author="SHEA G. M." box="[155,286,853,880]" pageId="3" pageNumber="502" pagination="338 - 340" refId="ref6907" refString="SHEA G. M. 2001. - Hyla lesueurii Bory de Saint-Vincent, 1828: an overlooked and problematic frog species name. Journal of Herpetology 35 (2): 338 - 340." type="journal article" year="2001">Shea (2001)</bibRefCitation>
rediscovered the publication of Bory de
<collectingCountry id="F36676415D05FFA966D3564EFF3CFC44" name="Saint Vincent and the Grenadines" pageId="3" pageNumber="502">Saint Vincent</collectingCountry>
and subsequently Caramaschi &amp; Niemeyer (2010) stated that the nomen
<taxonomicName id="4C714D525D05FFA96576568EFE00FC64" authorityName="Bory de Saint-Vincent" authorityYear="1828" box="[373,486,917,943]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="quoyi">
<emphasis id="B905EAC35D05FFA96576568EFE00FC64" box="[373,486,917,943]" italics="true" pageId="3" pageNumber="502">Hyla quoyi</emphasis>
</taxonomicName>
in this work applied to the same species as
<taxonomicName id="4C714D525D05FFA9653156AEFE5FFC05" authorityName="Burmeister" authorityYear="1856" box="[306,441,949,975]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA9653156AEFE5FFC05" box="[306,441,949,975]" italics="true" pageId="3" pageNumber="502">Hyla prasina</emphasis>
</taxonomicName>
, but they argued that the nomen
<taxonomicName id="4C714D525D05FFA964BF56CEFEE1FC25" authorityName="Burmeister" authorityYear="1856" box="[188,263,981,1006]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA964BF56CEFEE1FC25" box="[188,263,981,1006]" italics="true" pageId="3" pageNumber="502">prasina</emphasis>
</taxonomicName>
should be maintained for this species by virtue of Article 23.9. However they failed to provide 25 references to the use of this nomen as expressly required by this Article.
<bibRefCitation id="EFE04B205D05FFA96487512FFED3FB84" author="DUBOIS A." box="[132,309,1076,1103]" pageId="3" pageNumber="502" pagination="1 - 48" refId="ref5788" refString="DUBOIS A. 2017 b. - The nomenclatural status of Hysaplesia, Hylaplesia, Dendrobates and related nomina (Amphibia, Anura), with general comments on zoological nomenclature and its governance, as well as on taxonomic databases and websites. Bionomina 11: 1 - 48. https: // doi. org / 10.11646 / bionomina. 11.1.1" type="journal article" year="2017">Dubois (2017b)</bibRefCitation>
had noticed that the conditions of Article 23.9 had not been complied with for invalidation of the nomen
<taxonomicName id="4C714D525D05FFA964B9516FFECCFB45" authorityName="Bory de Saint-Vincent" authorityYear="1828" box="[186,298,1140,1166]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="quoyi">
<emphasis id="B905EAC35D05FFA964B9516FFECCFB45" box="[186,298,1140,1166]" italics="true" pageId="3" pageNumber="502">Hyla quoyi</emphasis>
</taxonomicName>
so that, without further detail, he stated that the nomen
<taxonomicName id="4C714D525D05FFA96507518FFE91FB65" authorityName="Bory de Saint-Vincent" authorityYear="1828" box="[260,375,1172,1198]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="quoyi">
<emphasis id="B905EAC35D05FFA96507518FFE91FB65" box="[260,375,1172,1198]" italics="true" pageId="3" pageNumber="502">Hyla quoyi</emphasis>
</taxonomicName>
was the valid nomen of this species under the combination
<taxonomicName id="4C714D525D05FFA9658451AEFDEFFB05" baseAuthorityName="Bory de Saint-Vincent" baseAuthorityYear="1828" box="[391,521,1205,1230]" class="Amphibia" family="Hylidae" genus="Boana" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="quoyi">
<emphasis id="B905EAC35D05FFA9658451AEFDEFFB05" box="[391,521,1205,1230]" italics="true" pageId="3" pageNumber="502">Boana quoyi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCE36D15D05FFA9649851CFFE18FA26" blockId="3.[130,777,279,1804]" pageId="3" pageNumber="502">
<bibRefCitation id="EFE04B205D05FFA9649851CFFE40FB25" author="OHLER A. &amp; DUBOIS A." box="[155,422,1236,1262]" pageId="3" pageNumber="502" pagination="109 - 121" refId="ref6602" refString="OHLER A. &amp; DUBOIS A. 2018. - Article 23.9 of the Code cannot be used to reject the nomen Hyla quoyi Bory de Saint-Vincent, 1828 as a nomen oblitum. Zoosystema 40 (6): 109 - 121. https: // doi. org / 10.5252 / zoosystema 2018 v 40 a 6." type="journal article" year="2018">Ohler &amp; Dubois (2018)</bibRefCitation>
came back to this problem and stressed that indeed Caramaschi &amp; Niemeyer (2010) had not validly used Article 23.9. They carried out an internet search which uncovered only 11 recent references (in the immediately preceding 50 years) to the use of the name
<taxonomicName id="4C714D525D05FFA9667E5048FCE6FAA6" authorityName="Burmeister" authorityYear="1856" box="[637,768,1363,1389]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA9667E5048FCE6FAA6" box="[637,768,1363,1389]" italics="true" pageId="3" pageNumber="502">Hyla prasina</emphasis>
</taxonomicName>
, and therefore confirmed Dubois (2017b) conclusion. Before publication, their paper had been sent out to four referees, none of whom questioned the statement that this nomen was not very well known of zoologists.
</paragraph>
<paragraph id="8BCE36D15D05FFA9649850E8FCE0F947" blockId="3.[130,777,279,1804]" pageId="3" pageNumber="502">
Shortly after,
<bibRefCitation id="EFE04B205D05FFA9652B50E8FDC7F9C6" author="KOLENC F. &amp; BALDO D." box="[296,545,1523,1549]" pageId="3" pageNumber="502" pagination="589 - 594" refId="ref6375" refString="KOLENC F. &amp; BALDO D. 2018. - Hyla prasina Burmeister, 1856 meets the requirements of Article 23.9. 1 of the Code for a reversal of precedence over Hyla quoyi Bory de Saint-Vincent, 1828 (Anura, Hylidae). Zootaxa 4442 (4): 589 - 594. https: // doi. org / 10.11646 / zootaxa. 4442.4.8" type="journal article" year="2018">Kolenc &amp; Baldo (2018)</bibRefCitation>
and
<bibRefCitation id="EFE04B205D05FFA9665650E8FF2AF9E6" author="COSTA H. C. &amp; SANTANA D. J." pageId="3" pageNumber="502" pagination="493 - 499" refId="ref5197" refString="COSTA H. C. &amp; SANTANA D. J. 2018. - The dangers of inadequate literature search to nomenclatural stability: the case of the nomina Hyla quoyi Bory de Saint-Vincent, 1828 and Hyla prasina Burmeister, 1856 (Amphibia: Anura). Zoosystema 40 (22): 493 - 499. https: // doi. org / 10.5252 / zoosystema 2018 v 40 a 22. http: // zoosystema. com / 40 / 22" type="journal article" year="2018">Costa &amp; Santana (2018)</bibRefCitation>
independently showed that Ohler &amp; Dubois (2018) internet search had provided incomplete results, and they gave lists of respectively 86 and 93 recent references to the use of this nomen as valid, thus complying with the criterion of Article 23.9.
</paragraph>
<paragraph id="8BCE36D15D05FFA964985388FD4DF8C7" blockId="3.[130,777,279,1804]" pageId="3" pageNumber="502">
We hereby agree that this number of references fulfills the requirements of Article 23.9 and should have led us to use this Article to validate the nomen
<taxonomicName id="4C714D525D05FFA965E953C9FD8BF927" authorityName="Burmeister" authorityYear="1856" box="[490,621,1746,1772]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="prasina">
<emphasis id="B905EAC35D05FFA965E953C9FD8BF927" box="[490,621,1746,1772]" italics="true" pageId="3" pageNumber="502">Hyla prasina</emphasis>
</taxonomicName>
. However this case raises several problems that deserve discussion.
</paragraph>
</subSubSection>
</treatment>
</document>