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<document id="E9C51638D38134429973AE33D1DEFF93" ID-CLB-Dataset="27854" ID-DOI="10.11646/zootaxa.4527.2.9" ID-GBIF-Dataset="9abfbfd4-e21a-4209-a180-aa66b6bc95bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2612187" ID-ZooBank="CF98E991-A154-48D8-8FF7-0F60FAD574C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553698431019" checkinUser="plazi" docAuthor="Nakahara, Shinichi, Macdonald, John R., Delgado, Francisco &amp; Padrón, Pablo Sebastián" docDate="2018" docId="03D887C1FFDB383E38C5F9CAD287FF3D" docLanguage="en" docName="zootaxa.4527.2.9.pdf" docOrigin="Zootaxa 4527 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Catasticta sibyllae Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón 2018, new species" docType="treatment" docVersion="8" lastPageNumber="287" masterDocId="FFE1FFB9FFD938393852FF85D165FFC5" masterDocTitle="Discovery of a rare and striking new pierid butterfly from Panama (Lepidoptera: Pieridae)" masterLastPageNumber="291" masterPageNumber="281" pageNumber="283" updateTime="1698714458446" updateUser="ExternalLinkService">
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<mods:titleInfo id="1DA479637F7B5533DDF4993E99CA824E">
<mods:title id="71AD125D842A3602818BBB9636D76ADB">Discovery of a rare and striking new pierid butterfly from Panama (Lepidoptera: Pieridae)</mods:title>
</mods:titleInfo>
<mods:name id="3563247EC2481A4BFADDDA877E2EB2D1" type="personal">
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<mods:namePart id="B57A9E3C54996341F3857F05509EC581">Nakahara, Shinichi</mods:namePart>
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<mods:namePart id="1302BB84DCDCC6CD7ACF1982909AB585">Macdonald, John R.</mods:namePart>
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<mods:namePart id="0EAD24C03DA6D778370D3249E288EDAD">Delgado, Francisco</mods:namePart>
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<mods:namePart id="9F6CF3EA44510812410535554EA2B855">Padrón, Pablo Sebastián</mods:namePart>
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<mods:title id="FD5C5CCED29F0D9979EC11E2E2F62EC2">Zootaxa</mods:title>
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<mods:part id="7F8A8EB9347876F4BCD94829EFD68EC1">
<mods:date id="F69982E6FCC1190E8631BE9FAEE5473D">2018</mods:date>
<mods:detail id="08EC7FFC3F3F7DC5604700918A0AF185" type="pubDate">
<mods:number id="0D75406AAB6D266EC7AE61E951C28171">2018-12-05</mods:number>
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<mods:number id="E81B9CF29872DE7474C46804EF4BE555">4527</mods:number>
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<mods:classification id="0938C1143CF9EC9DB5D3E62E65C71AA0">journal article</mods:classification>
<mods:identifier id="6353A67B7B14D64902B0B6730F7AC108" type="CLB-Dataset">27854</mods:identifier>
<mods:identifier id="18CB928FC46247A09563086C246C704F" type="DOI">10.11646/zootaxa.4527.2.9</mods:identifier>
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<treatment id="03D887C1FFDB383E38C5F9CAD287FF3D" ID-DOI="http://doi.org/10.5281/zenodo.5950024" ID-GBIF-Taxon="154508065" ID-Zenodo-Dep="5950024" LSID="urn:lsid:plazi:treatment:03D887C1FFDB383E38C5F9CAD287FF3D" httpUri="http://treatment.plazi.org/id/03D887C1FFDB383E38C5F9CAD287FF3D" lastPageId="7" lastPageNumber="287" pageId="2" pageNumber="283">
<subSubSection id="C36B655CFFDB383B38C5F9CAD2DCF9AF" box="[151,953,1615,1642]" pageId="2" pageNumber="283" type="nomenclature">
<paragraph id="8BCE36D7FFDB383B38C5F9CAD2DCF9AF" blockId="2.[151,953,1615,1675]" box="[151,953,1615,1642]" pageId="2" pageNumber="283">
<heading id="D08681BBFFDB383B38C5F9CAD2DCF9AF" bold="true" box="[151,953,1615,1642]" fontSize="11" level="1" pageId="2" pageNumber="283" reason="1">
<emphasis id="B905EAC5FFDB383B38C5F9CAD2DCF9AF" bold="true" box="[151,953,1615,1642]" pageId="2" pageNumber="283">
<taxonomicName id="4C714D54FFDB383B38C5F9CAD279F9AF" authority="Nakahara, Padron &amp; MacDonald" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[151,796,1615,1642]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDB383B38C5F9CAD014F9AC" bold="true" box="[151,369,1615,1641]" italics="true" pageId="2" pageNumber="283">Catasticta sibyllae</emphasis>
Nakahara, Padrón &amp; MacDonald
</taxonomicName>
,
<taxonomicNameLabel id="A23657BEFFDB383B3B7BF9D5D2DCF9AF" box="[809,953,1616,1642]" pageId="2" pageNumber="283" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B655CFFDB383B38C5F9F7D06EF94E" box="[151,267,1650,1675]" pageId="2" pageNumber="283" type="description">
<paragraph id="8BCE36D7FFDB383B38C5F9F7D06EF94E" blockId="2.[151,953,1615,1675]" box="[151,267,1650,1675]" pageId="2" pageNumber="283">
(
<figureCitation id="134A2A52FFDB383B38CDF9F7D066F94E" box="[159,259,1650,1675]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="4.[151,250,1114,1136]" captionStartId-1="5.[151,250,1821,1843]" captionStartId-2="6.[151,250,1543,1565]" captionStartId-3="7.[151,250,1515,1537]" captionStartId-4="8.[151,250,1969,1991]" captionTargetBox-0="[204,1394,193,1064]" captionTargetBox-1="[177,1432,215,1766]" captionTargetBox-3="[242,1344,301,1501]" captionTargetBox-4="[180,1404,199,1945]" captionTargetId-0="figure@4.[204,1394,193,1064]" captionTargetId-1="figure@5.[151,1435,193,1800]" captionTargetId-3="figure@7.[242,1344,301,1501]" captionTargetId-4="figure@8.[180,1406,193,1954]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2. Catasticta sibyllae n. sp. holotype male (dorsal on left, ventral on right); paratype male (dorsal on left, ventral on right)." captionText-1="FIGURE 3. Catasticta sibyllae n. sp. male genitalia, dissection based on the holotype (genitalic vial: SN-17-177): a) lateral view of genitalia in situ at posterior end of abdomen; b) genitalia in lateral view; c) lateral view of uncus, with valva removed." captionText-2="FIGURE 4. Catasticta sibyllae n. sp. wing venation based on the holotype." captionText-3="FIGURE 5. Catasticta sibyllae n. sp. morphology, all images based on the holotype: a) head in lateral view; b) anntennal club in dorsal view; d) abdomen in ventral view." captionText-4="FIGURE 6. Distribution records for Catasticta sibyllae n. sp., Atlantic slope habitat image of Fortuna forest, where the paratype male was found (photo credit: Thomas J. Riley)." httpUri-0="https://zenodo.org/record/2612191/files/figure.png" httpUri-1="https://zenodo.org/record/2612193/files/figure.png" httpUri-3="https://zenodo.org/record/2612195/files/figure.png" httpUri-4="https://zenodo.org/record/2612197/files/figure.png" pageId="2" pageNumber="283">Figs 2–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36B655CFFDB383B38C5F93FD527F86E" pageId="2" pageNumber="283" type="diagnosis">
<paragraph id="8BCE36D7FFDB383B38C5F93FD527F86E" blockId="2.[151,1436,1722,1963]" pageId="2" pageNumber="283">
<emphasis id="B905EAC5FFDB383B38C5F93FD074F916" bold="true" box="[151,273,1722,1747]" pageId="2" pageNumber="283">Diagnosis.</emphasis>
<taxonomicName id="4C714D54FFDB383B394AF93ED08FF916" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[280,490,1722,1747]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDB383B394AF93ED08FF916" box="[280,490,1722,1747]" italics="true" pageId="2" pageNumber="283">Catasticta sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDB383B39A0F93FD354F916" bold="true" box="[498,561,1722,1747]" pageId="2" pageNumber="283">
<taxonomicNameLabel id="A23657BEFFDB383B39A0F93FD354F916" box="[498,561,1722,1747]" pageId="2" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is easily distinguished from all
<taxonomicName id="4C714D54FFDB383B3BC9F93ED56BF917" authorityName="Butler" authorityYear="1870" box="[923,1038,1723,1746]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAC5FFDB383B3BC9F93ED56BF917" box="[923,1038,1723,1746]" italics="true" pageId="2" pageNumber="283">Catasticta</emphasis>
</taxonomicName>
species by the median area of both surfaces of forewing and hindwing being entirely black with no markings.
<taxonomicName id="4C714D54FFDB383B3C56F95AD5BCF932" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[1028,1241,1758,1783]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDB383B3C56F95AD5BCF932" box="[1028,1241,1758,1783]" italics="true" pageId="2" pageNumber="283">Catasticta sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDB383B3CB1F95BD440F932" bold="true" box="[1251,1317,1758,1783]" pageId="2" pageNumber="283">
<taxonomicNameLabel id="A23657BEFFDB383B3CB1F95BD440F932" box="[1251,1317,1758,1783]" pageId="2" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is readily distinguished from its sister species,
<taxonomicName id="4C714D54FFDB383B3A1AF886D3F0F8DE" authorityName="Baumann &amp; Reissinger" authorityYear="1969" box="[584,661,1794,1819]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="lisa">
<emphasis id="B905EAC5FFDB383B3A1AF886D3F0F8DE" box="[584,661,1794,1819]" italics="true" pageId="2" pageNumber="283">C. lisa</emphasis>
</taxonomicName>
, by the lack of broad white median band on both sides of dorsal forewing and hindwing.
<taxonomicName id="4C714D54FFDB383B39E7F8A2D3EEF8FA" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[437,651,1830,1855]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDB383B39E7F8A2D3EEF8FA" box="[437,651,1830,1855]" italics="true" pageId="2" pageNumber="283">Catasticta sibyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657BEFFDB383B3AC7F8A3D3BDF8FA" box="[661,728,1830,1855]" pageId="2" pageNumber="283" rank="species">
<emphasis id="B905EAC5FFDB383B3AC7F8A3D3BDF8FA" bold="true" box="[661,728,1830,1855]" pageId="2" pageNumber="283">n. sp.</emphasis>
</taxonomicNameLabel>
is further distinguished from
<taxonomicName id="4C714D54FFDB383B3C6BF8A2D5E0F8FA" authorityName="Baumann &amp; Reissinger" authorityYear="1969" box="[1081,1157,1830,1855]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="lisa">
<emphasis id="B905EAC5FFDB383B3C6BF8A2D5E0F8FA" box="[1081,1157,1830,1855]" italics="true" pageId="2" pageNumber="283">C. lisa</emphasis>
</taxonomicName>
by its rather prominent submarginal and marginal spots on both sides of dorsal forewing and hindwing. Also, these spots on the ventral surface appear more whitish in
<taxonomicName id="4C714D54FFDB383B39A5F8EAD315F842" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[503,624,1902,1927]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDB383B39A5F8EAD315F842" box="[503,624,1902,1927]" italics="true" pageId="2" pageNumber="283">C. sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDB383B3A24F8EBD3D1F842" bold="true" box="[630,692,1902,1927]" pageId="2" pageNumber="283">
<taxonomicNameLabel id="A23657BEFFDB383B3A24F8EBD3D1F842" box="[630,692,1902,1927]" pageId="2" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, whereas more yellowish in
<taxonomicName id="4C714D54FFDB383B3BA5F8EAD525F842" authorityName="Baumann &amp; Reissinger" authorityYear="1969" box="[1015,1088,1902,1927]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="lisa">
<emphasis id="B905EAC5FFDB383B3BA5F8EAD525F842" box="[1015,1088,1902,1927]" italics="true" pageId="2" pageNumber="283">C. lisa</emphasis>
</taxonomicName>
. The genitalia appear not to be informative in this group, thus diagnostic genitalic characters are not provided here.
</paragraph>
</subSubSection>
<caption id="DF0E665FFFDA383A38C5FDCDD40AFD9B" box="[151,1391,584,606]" httpUri="https://zenodo.org/record/2612189/files/figure.png" pageId="3" pageNumber="284" startId="3.[151,250,584,606]" targetBox="[293,1369,197,505]" targetPageId="3">
<paragraph id="8BCE36D7FFDA383A38C5FDCDD40AFD9B" blockId="3.[151,1391,584,606]" box="[151,1391,584,606]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A38C5FDCDD076FD98" bold="true" box="[151,275,584,606]" pageId="3" pageNumber="284">FIGURE 1.</emphasis>
BI phylogenetic tree showing relationship of
<taxonomicName id="4C714D54FFDA383A3AB1FDCCD2C4FD9B" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[739,929,584,606]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDA383A3AB1FDCCD2C4FD9B" box="[739,929,584,606]" italics="true" pageId="3" pageNumber="284">Catasticta sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDA383A3BF5FDCCD285FD9B" bold="true" box="[935,992,585,606]" pageId="3" pageNumber="284">
<taxonomicNameLabel id="A23657BEFFDA383A3BF5FDCCD285FD9B" box="[935,992,585,606]" pageId="3" pageNumber="284" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(node support is posterior probability).
</paragraph>
</caption>
<subSubSection id="C36B655CFFDA383D3895FD03D4F8F8FD" lastPageId="4" lastPageNumber="285" pageId="3" pageNumber="284" type="description">
<paragraph id="8BCE36D7FFDA383A3895FD03D57BFD5B" blockId="3.[151,1437,646,2033]" box="[199,1054,646,671]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FD03D0D9FD5A" bold="true" box="[199,444,646,671]" pageId="3" pageNumber="284">Description. MALE:</emphasis>
Forewing length,
<typeStatus id="54CA8875FFDA383A3AD8FD03D395FD5A" box="[650,752,646,671]" pageId="3" pageNumber="284" type="holotype">holotype</typeStatus>
:
<quantity id="4C899B32FFDA383A3AA9FD02D228FD5B" box="[763,845,647,671]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="284" unit="mm" value="30.0">30 mm</quantity>
;
<typeStatus id="54CA8875FFDA383A3B0AFD02D2D8FD5A" box="[856,957,647,671]" pageId="3" pageNumber="284" type="paratype">paratype</typeStatus>
:
<quantity id="4C899B32FFDA383A3B95FD02D57CFD5B" box="[967,1049,647,671]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="284" unit="mm" value="33.0">33 mm</quantity>
.
</paragraph>
<paragraph id="8BCE36D7FFDA383A3895FD2FD51BFCB2" blockId="3.[151,1437,646,2033]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FD2FD0FFFD06" bold="true" box="[199,410,682,707]" pageId="3" pageNumber="284">
Head (
<figureCitation id="134A2A52FFDA383A3947FD2FD0EFFD06" box="[277,394,682,707]" captionStart="FIGURE 5" captionStartId="7.[151,250,1515,1537]" captionTargetBox="[242,1344,301,1501]" captionTargetId="figure@7.[242,1344,301,1501]" captionTargetPageId="7" captionText="FIGURE 5. Catasticta sibyllae n. sp. morphology, all images based on the holotype: a) head in lateral view; b) anntennal club in dorsal view; d) abdomen in ventral view." httpUri="https://zenodo.org/record/2612195/files/figure.png" pageId="3" pageNumber="284">Figs 5a, b</figureCitation>
):
</emphasis>
Eyes naked, with white scales at base; frons covered with brown scales, adorned with black long hair-like scales and white long hair-like scales, area around antennae base similarly with black long hair-like scales and white long hair-like scales; labial palpi roughly twice as long as eye depth, ventrally with white long hair-like scales and brown long hair-like scales, laterally with black scales with ventral and dorsal margin with white scales; antennae approximately half of forewing length, brown, ventrally scattered with white scales, with ca. 35 anntenomeres (n=1, HT), distal 9–10 anntenomeres composing rather prominent club.
</paragraph>
<paragraph id="8BCE36D7FFDA383A3895FC07D36BFC7A" blockId="3.[151,1437,646,2033]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FC07D04CFC5E" bold="true" box="[199,297,898,923]" pageId="3" pageNumber="284">Thorax:</emphasis>
Dorsally brown with light coloured long hair-like scales; ventrally brown with light coloured long hair-like scales and brown scales.
</paragraph>
<paragraph id="8BCE36D7FFDA383A3895FC4FD5ACFBEE" blockId="3.[151,1437,646,2033]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FC4FD06DFC26" bold="true" box="[199,264,970,995]" pageId="3" pageNumber="284">Legs:</emphasis>
Foreleg black, foretarsus longer than tibia, foretarsus and tibia scattered with whitish scales ventrally, in addition to multiple spines, dorsally stripe of white scales present, tibial spurs absent, femur slightly longer then tibia, ventrally brown, dorsally stripe of white scales present; midleg and hindleg not examined.
</paragraph>
<paragraph id="8BCE36D7FFDA383A3895FBB3D2F8FB5C" blockId="3.[151,1437,646,2033]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FBB3D080FB8A" bold="true" box="[199,485,1078,1103]" pageId="3" pageNumber="284">
Wing venation (
<figureCitation id="134A2A52FFDA383A39D9FBB3D0B3FB8A" box="[395,470,1078,1103]" captionStart="FIGURE 4" captionStartId="6.[151,250,1543,1565]" captionText="FIGURE 4. Catasticta sibyllae n. sp. wing venation based on the holotype." pageId="3" pageNumber="284">Fig. 4</figureCitation>
):
</emphasis>
Base of forewing subcostal vein not swollen; four forewing radius veins present; origin of M
<subScript id="17F53492FFDA383A394EFBECD041FBB2" attach="left" box="[284,292,1129,1143]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
not associated with discal cell; base of cubitus not swollen; forewing recurrent vein absent; middle portion of m
<subScript id="17F53492FFDA383A3971FB0AD04EFB58" attach="right" box="[291,299,1167,1181]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-m
<subScript id="17F53492FFDA383A3918FB0AD037FB58" attach="left" box="[330,338,1167,1181]" fontSize="6" pageId="3" pageNumber="284">3</subScript>
portion reduced; hindwing humeral vein developed.
</paragraph>
<paragraph id="8BCE36D7FFDA383A3895FB22D3CAFAE9" blockId="3.[151,1437,646,2033]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FB22D0BCFB05" bold="true" box="[199,473,1191,1216]" pageId="3" pageNumber="284">
Wing shape (
<figureCitation id="134A2A52FFDA383A3930FB22D0C9FB05" box="[354,428,1191,1216]" captionStart="FIGURE 2" captionStartId="4.[151,250,1114,1136]" captionTargetBox="[204,1394,193,1064]" captionTargetId="figure@4.[204,1394,193,1064]" captionTargetPageId="4" captionText="FIGURE 2. Catasticta sibyllae n. sp. holotype male (dorsal on left, ventral on right); paratype male (dorsal on left, ventral on right)." httpUri="https://zenodo.org/record/2612191/files/figure.png" pageId="3" pageNumber="284">Figs 2</figureCitation>
,
<figureCitation id="134A2A52FFDA383A39EBFB22D0AFFB05" box="[441,458,1191,1216]" captionStart="FIGURE 4" captionStartId="6.[151,250,1543,1565]" captionText="FIGURE 4. Catasticta sibyllae n. sp. wing venation based on the holotype." pageId="3" pageNumber="284">4</figureCitation>
):
</emphasis>
Forewing subtriangular, appearing somewhat elongated with costal margin convex in angular manner and concave outer margin, inner margin rather straight; hindwing rounded, appearing slightly elongated, costal margin curved, angles inwards at base, outer margin curved, inner margin slightly concave near tornus, anal lobe almost straight, slightly round.
</paragraph>
<paragraph id="8BCE36D7FFDA383A3895FAB2D1A5F834" blockId="3.[151,1437,646,2033]" pageId="3" pageNumber="284">
<emphasis id="B905EAC5FFDA383A3895FAB2D0A8FA95" bold="true" box="[199,461,1335,1360]" pageId="3" pageNumber="284">
Wing pattern (
<figureCitation id="134A2A52FFDA383A3924FAB2D0D8FA95" box="[374,445,1335,1360]" captionStart="FIGURE 2" captionStartId="4.[151,250,1114,1136]" captionTargetBox="[204,1394,193,1064]" captionTargetId="figure@4.[204,1394,193,1064]" captionTargetPageId="4" captionText="FIGURE 2. Catasticta sibyllae n. sp. holotype male (dorsal on left, ventral on right); paratype male (dorsal on left, ventral on right)." httpUri="https://zenodo.org/record/2612191/files/figure.png" pageId="3" pageNumber="284">Fig. 2</figureCitation>
):
</emphasis>
<emphasis id="B905EAC5FFDA383A3986FAB2D3F0FA95" box="[468,661,1335,1360]" italics="true" pageId="3" pageNumber="284">Dorsal Forewing</emphasis>
: ground colour black; white submarginal spots slightly suffused with black scales, present in all cells from R
<subScript id="17F53492FFDA383A3A18FAEFD337FABD" attach="right" box="[586,594,1386,1400]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
+R
<subScript id="17F53492FFDA383A3A26FAEFD319FABD" attach="left" box="[628,636,1386,1400]" fontSize="6" pageId="3" pageNumber="284">3</subScript>
to 2A, spot in cell R
<subScript id="17F53492FFDA383A3B33FAEFD20CFABD" attach="right" box="[865,873,1386,1400]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
+R
<subScript id="17F53492FFDA383A3BD9FAEFD2F6FABD" attach="left" box="[907,915,1386,1400]" fontSize="6" pageId="3" pageNumber="284">3</subScript>
-R
<subScript id="17F53492FFDA383A3BFDFAEFD2D2FABD" attach="left" box="[943,951,1386,1400]" fontSize="6" pageId="3" pageNumber="284">4</subScript>
elongated, appearing more as costal streak, spot in cell R
<subScript id="17F53492FFDA383A397EFA14D051FA5A" attach="left" box="[300,308,1425,1439]" fontSize="6" pageId="3" pageNumber="284">4</subScript>
-R
<subScript id="17F53492FFDA383A3902FA14D03DFA5A" attach="left" box="[336,344,1425,1439]" fontSize="6" pageId="3" pageNumber="284">5</subScript>
small, similar in appearance, spots in cells from R
<subScript id="17F53492FFDA383A3BDDFA14D2F2FA5A" attach="left" box="[911,919,1425,1439]" fontSize="6" pageId="3" pageNumber="284">4</subScript>
to Cu
<subScript id="17F53492FFDA383A3B8EFA14D281FA5A" attach="left" box="[988,996,1425,1439]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
subsquared, except for streak-like spot in cell M
<subScript id="17F53492FFDA383A38AEFA3DD061FA03" attach="left" box="[252,260,1464,1478]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-M
<subScript id="17F53492FFDA383A3974FA3DD04BFA03" attach="left" box="[294,302,1464,1478]" fontSize="6" pageId="3" pageNumber="284">3</subScript>
, spot in cell Cu
<subScript id="17F53492FFDA383A398DFA3DD082FA03" attach="left" box="[479,487,1464,1478]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-2A appearing somewhat doubled; marginal spots slightly suffused with black scales present in all cells from R
<subScript id="17F53492FFDA383A39ECFA5AD0A3FA28" attach="left" box="[446,454,1503,1517]" fontSize="6" pageId="3" pageNumber="284">4</subScript>
to 2A, those in cells from R
<subScript id="17F53492FFDA383A3B58FA5AD277FA28" attach="left" box="[778,786,1503,1517]" fontSize="6" pageId="3" pageNumber="284">4</subScript>
to Cu
<subScript id="17F53492FFDA383A3B0BFA5AD204FA28" attach="left" box="[857,865,1503,1517]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
teardrop-like with black suffusion visible on basal side, spot in cell Cu
<subScript id="17F53492FFDA383A39C8F983D0C7F9D1" attach="left" box="[410,418,1542,1556]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
-Cu
<subScript id="17F53492FFDA383A399EF983D0B1F9D1" attach="left" box="[460,468,1542,1556]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
squared, spot in cell Cu
<subScript id="17F53492FFDA383A3B43F983D27CF9D1" attach="left" box="[785,793,1542,1556]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-2A smallest, distance between submarginal spots in corresponding cells closer towards posterior end as outer margin traverses towards body; fringe blackish;
<emphasis id="B905EAC5FFDA383A3D1CF998D19AF99F" italics="true" pageId="3" pageNumber="284">Dorsal hindwing</emphasis>
: ground colour similar to forewing; white submarginal spots slightly suffused with black scales, present in all cells from Sc+R to 2A, spots in cells Cu
<subScript id="17F53492FFDA383A3AC7F9F1D3F8F947" attach="left" box="[661,669,1652,1666]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
-Cu
<subScript id="17F53492FFDA383A3A95F9F1D3AAF947" attach="left" box="[711,719,1652,1666]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
and Cu
<subScript id="17F53492FFDA383A3B77F9F1D248F947" attach="left" box="[805,813,1652,1666]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-2A smaller, in particular, spot in cell Cu
<subScript id="17F53492FFDA383A3CA6F9F1D599F947" attach="left" box="[1268,1276,1652,1666]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-2A appearing as white hint; marginal spots present in all cells from Sc-R
<subScript id="17F53492FFDA383A3B11F91ED22EF96C" attach="left" box="[835,843,1691,1705]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
to 2A, spot in Sc+R
<subScript id="17F53492FFDA383A3C6FF91ED520F96C" attach="left" box="[1085,1093,1691,1705]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
-Rs elongated and streak-like, spots in cells Rs-M
<subScript id="17F53492FFDA383A392FF947D0E0F915" attach="left" box="[381,389,1730,1744]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
, M
<subScript id="17F53492FFDA383A39E2F947D0DDF915" attach="left" box="[432,440,1730,1744]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
-M
<subScript id="17F53492FFDA383A3988F947D087F915" attach="left" box="[474,482,1730,1744]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
and M
<subScript id="17F53492FFDA383A3A69F947D326F915" attach="left" box="[571,579,1730,1744]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-M
<subScript id="17F53492FFDA383A3A37F947D308F915" attach="left" box="[613,621,1730,1744]" fontSize="6" pageId="3" pageNumber="284">3</subScript>
rather trapezoidal, spots in cells M
<subScript id="17F53492FFDA383A3C46F947D579F915" attach="left" box="[1044,1052,1730,1744]" fontSize="6" pageId="3" pageNumber="284">3</subScript>
-Cu
<subScript id="17F53492FFDA383A3C14F947D52BF915" attach="right" box="[1094,1102,1730,1744]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
, Cu
<subScript id="17F53492FFDA383A3CD3F947D5ECF915" attach="right" box="[1153,1161,1730,1744]" fontSize="6" pageId="3" pageNumber="284">1</subScript>
-Cu
<subScript id="17F53492FFDA383A3CE1F947D5DEF915" attach="left" box="[1203,1211,1730,1744]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
and Cu
<subScript id="17F53492FFDA383A3D4EF947D441F915" attach="left" box="[1308,1316,1730,1744]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-2A rather rectangular; fringe black;
<emphasis id="B905EAC5FFDA383A3996F95FD3E7F937" box="[452,642,1754,1779]" italics="true" pageId="3" pageNumber="284">Ventral forewing</emphasis>
: ground colour and general wing pattern similar to dorsal forewing except as follows: ground colour slightly paler; marginal whitish spots appearing larger; submarginal whitish spots appearing larger and more pointier towards base, submarginal spot in cell Cu
<subScript id="17F53492FFDA383A3BADF8B4D562F8FA" attach="left" box="[1023,1031,1841,1855]" fontSize="6" pageId="3" pageNumber="284">2</subScript>
-2A appearing as two spots;
<emphasis id="B905EAC5FFDA383A3D19F8A7D19AF8A7" italics="true" pageId="3" pageNumber="284">Ventral hindwing</emphasis>
: ground colour and general wing pattern similar to dorsal hindwing except as follows: ground colour slightly paler; yellow costal streak at base of costal cell, just distal of humeral vein; reddish patch just basal of humeral vein above subcostal vein; yellow patch at base of discal cell; reddish patch at base of anal cell; submarginal spots smaller and slightly yellow; marginal spots somewhat creamy, additional spot present in cell 2A- 3A.
</paragraph>
<caption id="DF0E665FFFDD383D38C5FBDFD1B3FB4A" httpUri="https://zenodo.org/record/2612191/files/figure.png" pageId="4" pageNumber="285" startId="4.[151,250,1114,1136]" targetBox="[204,1394,193,1064]" targetPageId="4">
<paragraph id="8BCE36D7FFDD383D38C5FBDFD1B3FB4A" blockId="4.[151,1436,1114,1167]" pageId="4" pageNumber="285">
<emphasis id="B905EAC5FFDD383D38C5FBDFD076FBAA" bold="true" box="[151,275,1114,1136]" pageId="4" pageNumber="285">FIGURE 2.</emphasis>
<taxonomicName id="4C714D54FFDD383D3948FBDED0B2FBB5" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[282,471,1114,1136]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDD383D3948FBDED0B2FBB5" box="[282,471,1114,1136]" italics="true" pageId="4" pageNumber="285">Catasticta sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDD383D398CFBDFD372FBAA" bold="true" box="[478,535,1114,1135]" pageId="4" pageNumber="285">
<taxonomicNameLabel id="A23657BEFFDD383D398CFBDFD372FBAA" box="[478,535,1114,1135]" pageId="4" pageNumber="285" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype male (dorsal on left, ventral on right); paratype male (dorsal on left, ventral on right).
</paragraph>
</caption>
<paragraph id="8BCE36D7FFDD383D3895FB3ED5BDFB3D" blockId="4.[151,1437,1211,2028]" pageId="4" pageNumber="285">
<emphasis id="B905EAC5FFDD383D3895FB3ED0D1FB11" bold="true" box="[199,436,1211,1236]" pageId="4" pageNumber="285">
Abdomen (
<figureCitation id="134A2A52FFDD383D391FFB3ED0C1FB11" box="[333,420,1211,1236]" captionStart="FIGURE 5" captionStartId="7.[151,250,1515,1537]" captionTargetBox="[242,1344,301,1501]" captionTargetId="figure@7.[242,1344,301,1501]" captionTargetPageId="7" captionText="FIGURE 5. Catasticta sibyllae n. sp. morphology, all images based on the holotype: a) head in lateral view; b) anntennal club in dorsal view; d) abdomen in ventral view." httpUri="https://zenodo.org/record/2612195/files/figure.png" pageId="4" pageNumber="285">Fig. 5c</figureCitation>
):
</emphasis>
Dorsally and laterally black, ventrally with brown scales, white scales and white long hair-like scales, extending along ventral margin of valva; eighth sternite appearing as small plate.
</paragraph>
<paragraph id="8BCE36D7FFDD383D3895FA86D06DF941" blockId="4.[151,1437,1211,2028]" pageId="4" pageNumber="285">
<emphasis id="B905EAC5FFDD383D3895FA86D0FEFAD9" bold="true" box="[199,411,1283,1308]" pageId="4" pageNumber="285">
Genitalia (
<figureCitation id="134A2A52FFDD383D3916FA86D0E9FAD9" box="[324,396,1283,1308]" captionStart="FIGURE 3" captionStartId="5.[151,250,1821,1843]" captionTargetBox="[177,1432,215,1766]" captionTargetId="figure@5.[151,1435,193,1800]" captionTargetPageId="5" captionText="FIGURE 3. Catasticta sibyllae n. sp. male genitalia, dissection based on the holotype (genitalic vial: SN-17-177): a) lateral view of genitalia in situ at posterior end of abdomen; b) genitalia in lateral view; c) lateral view of uncus, with valva removed." httpUri="https://zenodo.org/record/2612193/files/figure.png" pageId="4" pageNumber="285">Fig. 3</figureCitation>
):
</emphasis>
Tegumen somewhat trapezoidal in lateral view; uncus shorter than tegumen in lateral view with convexed ventral margin, ending in a blunt point, with tuft of hair-like setae visible at base; ventral surface of anal tube weakly sclerotized (this structure is referred to as “gnathos” in
<bibRefCitation id="EFE04B26FFDD383D3B96FACED460FAA1" author="Radford, J. &amp; Willmott, K. R." box="[964,1285,1355,1380]" pageId="4" pageNumber="285" pagination="86 - 91" refId="ref6000" refString="Radford, J. &amp; Willmott, K. R. (2013) A distinctive new subspecies of Catasticta poujadei from the Cordillera del Condor in Eastern Ecuador (Lepidoptera: Pieridae: Pierinae). Tropical Lepidoptera Research, 23 (2), 86 - 91." type="journal article" year="2013">Radford &amp; Willmott (2013))</bibRefCitation>
; combination of ventral arms of tegumen and dorsal arms of saccus rather straight, middle portion broadens in lateral view; appendices angulares apparently absent; saccus short, broad and straight, similar to uncus in length; juxta appearing as tall plate in posterior view, narrow long stripe in lateral view; valva round, overall appearing somewhat as bean-shaped in lateral view, anterior margin rounded and positioned above uncus, ventral margin curved with hair-like setae present along margin, dorsal margin basal of costa concave, costa pointy, terminating in blunt end; phallus curved, phallobase about one-fourth of phallus in length with trapezoidal winglet ventrally, ductus ejaculatorius visible, manica covering more than half of aedeagus, vesica not examined as phallus not separated.
</paragraph>
<paragraph id="8BCE36D7FFDD383D3895F90AD3E4F96D" blockId="4.[151,1437,1211,2028]" box="[199,641,1679,1704]" pageId="4" pageNumber="285">
<emphasis id="B905EAC5FFDD383D3895F90AD02DF96D" bold="true" box="[199,328,1679,1704]" pageId="4" pageNumber="285">FEMALE:</emphasis>
Unknown or unrecognized.
</paragraph>
<paragraph id="8BCE36D7FFDD383D3895F936D417F935" blockId="4.[151,1437,1211,2028]" pageId="4" pageNumber="285">
<emphasis id="B905EAC5FFDD383D3895F936D0F5F909" bold="true" box="[199,400,1715,1740]" pageId="4" pageNumber="285">
Types.
<materialsCitation id="3B193C8AFFDD383D3948F936D0F5F909" ID-GBIF-Occurrence="2029173481" box="[282,400,1715,1740]" collectingDate="1981-09-13" collectionCode="USNM" collectorName="Gordon B. Small &amp; Genitalic &amp; S. Nakahara" country="Panama" elevation="800" location="Male" pageId="4" pageNumber="285" specimenCode="ENT 00804432" specimenCount="1" stateProvince="Veraguas" typeStatus="holotype">
<typeStatus id="54CA8875FFDD383D3948F936D0EEF909" box="[282,395,1715,1740]" pageId="4" pageNumber="285" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<location id="8EAE600CFFDD383D39C5F936D0B4F909" LSID="urn:lsid:plazi:treatment:03D887C1FFDB383E38C5F9CAD287FF3D:8EAE600CFFDD383D39C5F936D0B4F909" box="[407,465,1715,1740]" country="Panama" name="Male" pageId="4" pageNumber="285" stateProvince="Veraguas">Male</location>
with following labels written verbatim:
<collectingCountry id="F3667647FFDD383D3BC7F936D575F909" box="[917,1040,1715,1740]" name="Panama" pageId="4" pageNumber="285">PANAMÁ</collectingCountry>
:
<collectingRegion id="49B5F835FFDD383D3C48F931D5E4F909" box="[1050,1153,1716,1740]" country="Panama" name="Veraguas" pageId="4" pageNumber="285">Veraguas</collectingRegion>
Prov. Santa Fe
<quantity id="4C899B32FFDD383D3D62F931D415F909" box="[1328,1392,1716,1740]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="285" unit="m" value="800.0">
<elevation id="005CD1E4FFDD383D3D62F931D415F909" box="[1328,1392,1716,1740]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="285" unit="m" value="800.0">800m</elevation>
</quantity>
<date id="FFCF1017FFDD383D3D25F931D193F935" pageId="4" pageNumber="285" value="1981-09-13">
<collectingDate id="EF8BE9FFFFDD383D3D25F931D193F935" pageId="4" pageNumber="285" value="1981-09-13">13- IX-1981</collectingDate>
</date>
<collectorName id="26845301FFDD383D38AEF952D0DFF935" box="[252,442,1751,1776]" pageId="4" pageNumber="285">Gordon B. Small</collectorName>
//
<collectorName id="26845301FFDD383D3980F952D35CF935" box="[466,569,1751,1776]" pageId="4" pageNumber="285">Genitalic</collectorName>
vial SN-17-177
<collectorName id="26845301FFDD383D3AA6F95DD21EF935" box="[756,891,1751,1776]" pageId="4" pageNumber="285">S. Nakahara</collectorName>
// USNM
<specimenCode id="DBD79EACFFDD383D3BB9F95DD5FDF935" box="[1003,1176,1752,1776]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="285" type="Museum">ENT 00804432</specimenCode>
//FOTO// (USNM).
</paragraph>
<paragraph id="8BCE36D7FFDD383D3895F97ED4F8F8FD" blockId="4.[151,1437,1211,2028]" pageId="4" pageNumber="285">
<materialsCitation id="3B193C8AFFDD383D3895F97ED4F8F8FD" ID-GBIF-Occurrence="2029173482" collectingDate="2018-05-17" collectionCode="MEM" collectorName="John R. MacDonald" country="Panama" elevation="910" latitude="8.778016" location="Fortuna Cabins" longLatPrecision="914" longitude="-82.183334" municipality="Male" pageId="4" pageNumber="285" specimenCount="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<emphasis id="B905EAC5FFDD383D3895F97ED058F8D1" bold="true" box="[199,317,1787,1812]" pageId="4" pageNumber="285">
<typeStatus id="54CA8875FFDD383D3895F97ED05DF8D1" box="[199,312,1787,1812]" pageId="4" pageNumber="285" type="paratype">Paratype</typeStatus>
:
</emphasis>
<collectingMunicipality id="6BAAACADFFDD383D3917F97ED01AF8D1" box="[325,383,1787,1812]" pageId="4" pageNumber="285">Male</collectingMunicipality>
with following labels written verbatim:
<collectingCountry id="F3667647FFDD383D3B19F979D2A3F8D1" box="[843,966,1788,1812]" name="Panama" pageId="4" pageNumber="285">PANAMA</collectingCountry>
:
<collectingRegion id="49B5F835FFDD383D3B80F979D5E4F8D6" box="[978,1153,1787,1812]" country="Panama" name="Bocas del Toro" pageId="4" pageNumber="285">Bocas del Toro</collectingRegion>
; vic.
<location id="8EAE600CFFDD383D3CEDF979D415F8D1" LSID="urn:lsid:plazi:treatment:03D887C1FFDB383E38C5F9CAD287FF3D:8EAE600CFFDD383D3CEDF979D415F8D1" box="[1215,1392,1787,1812]" country="Panama" latitude="8.778016" longLatPrecision="914" longitude="-82.183334" municipality="Male" name="Fortuna Cabins" pageId="4" pageNumber="285" stateProvince="Bocas del Toro">Fortuna Cabins</location>
; ca.
<quantity id="4C899B32FFDD383D38C5F8A5D1B2F8FD" box="[151,215,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="285" unit="m" value="910.0">
<elevation id="005CD1E4FFDD383D38C5F8A5D1B2F8FD" box="[151,215,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="285" unit="m" value="910.0">910m</elevation>
</quantity>
<geoCoordinate id="EE455010FFDD383D388FF8A5D0FDF8FD" box="[221,408,1823,1848]" direction="north" orientation="latitude" pageId="4" pageNumber="285" precision="1" value="8.778016">N 08° 46' 40.86&quot;</geoCoordinate>
<geoCoordinate id="EE455010FFDD383D39CCF8A5D379F8FD" box="[414,540,1824,1848]" direction="west" orientation="longitude" pageId="4" pageNumber="285" precision="925" value="-82.183334">W 082° 11'</geoCoordinate>
2502&quot;
<date id="FFCF1017FFDD383D3A3EF8A5D262F8FD" box="[620,775,1824,1848]" pageId="4" pageNumber="285" value="2018-05-17">
<collectingDate id="EF8BE9FFFFDD383D3A3EF8A5D262F8FD" box="[620,775,1824,1848]" pageId="4" pageNumber="285" value="2018-05-17">May 17, 2018</collectingDate>
</date>
<collectorName id="26845301FFDD383D3B5CF89AD288F8FD" box="[782,1005,1823,1848]" pageId="4" pageNumber="285">John R. MacDonald</collectorName>
// DNA voucher LEP-68704// (
<collectionCode id="ED60AE12FFDD383D3D1AF8A5D4F4F8FD" box="[1352,1425,1824,1848]" country="USA" lsid="urn:lsid:biocol.org:col:15861" name="University of Memphis" pageId="4" pageNumber="285" type="Herbarium">MEM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B655CFFDD383D3895F8C6D37AF82E" pageId="4" pageNumber="285" type="etymology">
<paragraph id="8BCE36D7FFDD383D3895F8C6D37AF82E" blockId="4.[151,1437,1211,2028]" pageId="4" pageNumber="285">
<emphasis id="B905EAC5FFDD383D3895F8C6D02BF899" bold="true" box="[199,334,1859,1884]" pageId="4" pageNumber="285">Etymology.</emphasis>
The specific epithet is in recognition of Maria Sibylla Merian, a European naturalist and entomologist who embarked on a voyage to South America in the late seventeenth century and conducted empirical research focused on butterflies. Her research, in the form of numerous paintings of insects, served as a foundation for the scientific study of insects, now known as “entomology”. This specific epithet is regarded as a latinized feminine noun in the genitive case.
</paragraph>
</subSubSection>
<caption id="DF0E665FFFDC383C38C5F898D4F2F897" httpUri="https://zenodo.org/record/2612193/files/figure.png" pageId="5" pageNumber="286" startId="5.[151,250,1821,1843]" targetBox="[177,1432,215,1766]" targetPageId="5">
<paragraph id="8BCE36D7FFDC383C38C5F898D4F2F897" blockId="5.[151,1435,1821,1874]" pageId="5" pageNumber="286">
<emphasis id="B905EAC5FFDC383C38C5F898D071F8F6" bold="true" box="[151,276,1821,1843]" pageId="5" pageNumber="286">FIGURE 3.</emphasis>
<taxonomicName id="4C714D54FFDC383C394EF89BD0BEF8F6" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[284,475,1821,1843]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDC383C394EF89BD0BEF8F6" box="[284,475,1821,1843]" italics="true" pageId="5" pageNumber="286">Catasticta sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDC383C39B1F89BD37BF8F6" bold="true" box="[483,542,1822,1843]" pageId="5" pageNumber="286">
<taxonomicNameLabel id="A23657BEFFDC383C39B1F89BD37BF8F6" box="[483,542,1822,1843]" pageId="5" pageNumber="286" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
male genitalia, dissection based on the holotype (genitalic vial: SN-17-177): a) lateral view of genitalia
<emphasis id="B905EAC5FFDC383C3915F8B8D0E3F897" box="[327,390,1853,1874]" italics="true" pageId="5" pageNumber="286">in situ</emphasis>
at posterior end of abdomen; b) genitalia in lateral view; c) lateral view of uncus, with valva removed.
</paragraph>
</caption>
<caption id="DF0E665FFFDF383F38C5F982D2F8F9D8" box="[151,925,1543,1565]" pageId="6" pageNumber="287">
<paragraph id="8BCE36D7FFDF383F38C5F982D2F8F9D8" blockId="6.[151,925,1543,1565]" box="[151,925,1543,1565]" pageId="6" pageNumber="287">
<emphasis id="B905EAC5FFDF383F38C5F982D077F9D9" bold="true" box="[151,274,1543,1565]" pageId="6" pageNumber="287">FIGURE 4.</emphasis>
<taxonomicName id="4C714D54FFDF383F394DF98DD0B9F9D8" authority="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón, 2018" authorityName="Nakahara &amp; Macdonald &amp; Delgado &amp; Padrón" authorityYear="2018" box="[287,476,1543,1565]" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="sibyllae" status="sp. nov.">
<emphasis id="B905EAC5FFDF383F394DF98DD0B9F9D8" box="[287,476,1543,1565]" italics="true" pageId="6" pageNumber="287">Catasticta sibyllae</emphasis>
</taxonomicName>
<emphasis id="B905EAC5FFDF383F39B0F98DD37EF9D8" bold="true" box="[482,539,1544,1565]" pageId="6" pageNumber="287">
<taxonomicNameLabel id="A23657BEFFDF383F39B0F98DD37EF9D8" box="[482,539,1544,1565]" pageId="6" pageNumber="287" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
wing venation based on the holotype.
</paragraph>
</caption>
<subSubSection id="C36B655CFFDF383F3895F9C1D345F928" pageId="6" pageNumber="287" type="distribution">
<paragraph id="8BCE36D7FFDF383F3895F9C1D345F928" blockId="6.[151,1437,1604,2026]" pageId="6" pageNumber="287">
<emphasis id="B905EAC5FFDF383F3895F9C1D336F998" bold="true" box="[199,595,1604,1629]" pageId="6" pageNumber="287">
Distribution and habitat (
<figureCitation id="134A2A52FFDF383F39AFF9C1D326F998" box="[509,579,1604,1629]" captionStart="FIGURE 6" captionStartId="8.[151,250,1969,1991]" captionTargetBox="[180,1404,199,1945]" captionTargetId="figure@8.[180,1406,193,1954]" captionTargetPageId="8" captionText="FIGURE 6. Distribution records for Catasticta sibyllae n. sp., Atlantic slope habitat image of Fortuna forest, where the paratype male was found (photo credit: Thomas J. Riley)." httpUri="https://zenodo.org/record/2612197/files/figure.png" pageId="6" pageNumber="287">Fig. 6</figureCitation>
).
</emphasis>
This species is known to date from two sites situated in western
<collectingCountry id="F3667647FFDF383F3D6EF9C3D4FDF998" box="[1340,1432,1606,1629]" name="Panama" pageId="6" pageNumber="287">Panama</collectingCountry>
, namely Bosque Reserva Palo Seco,
<collectingRegion id="49B5F835FFDF383F3A66F9EFD384F944" box="[564,737,1641,1666]" country="Panama" name="Bocas del Toro" pageId="6" pageNumber="287">Bocas del Toro</collectingRegion>
(08̊46' 40.86&quot; N 0 82̊11' 25.02&quot; W) and Santa Fé,
<collectingRegion id="49B5F835FFDF383F3D67F9EFD4F9F947" box="[1333,1436,1642,1666]" country="Panama" name="Veraguas" pageId="6" pageNumber="287">Veraguas</collectingRegion>
(precise coordinates unavailable, but the town of Santa Fe lies at
<geoCoordinate id="EE455010FFDF383F3B2CF908D558F960" box="[894,1085,1677,1702]" direction="north" orientation="latitude" pageId="6" pageNumber="287" precision="1" value="8.499997">
08° 29' 59.99&quot;
<geoCoordinate id="EE455010FFDF383F3C7BF90BD558F960" box="[1065,1085,1678,1701]" direction="north" orientation="latitude" pageId="6" pageNumber="287" precision="925" value="81.05">N</geoCoordinate>
</geoCoordinate>
81° 03' 60.00&quot; W). Both sites are somewhat on the border of two vegetation
<typeStatus id="54CA8875FFDF383F3A90F937D265F90F" box="[706,768,1714,1738]" pageId="6" pageNumber="287">types</typeStatus>
, sub-montane rainforest and montane rainforest, near the continental divide (
<bibRefCitation id="EFE04B26FFDF383F393DF950D371F928" author="Blandin, P." box="[367,532,1749,1774]" pageId="6" pageNumber="287" pagination="147 - 154" refId="ref4989" refString="Blandin, P. (2017) La diversite des populations de Morpho helenor (Cramer, 1776) au Panama (Lepidoptera Nymphalidae Satyrinae Morphini). Antenor, 4 (2), 147 - 154." type="journal article" year="2017">Blandin, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B655CFFDF383E3895F97CD287FF3D" lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287" type="biology_ecology">
<paragraph id="8BCE36D7FFDF383E3895F97CD287FF3D" blockId="6.[151,1437,1604,2026]" lastBlockId="7.[151,1436,151,248]" lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287">
<emphasis id="B905EAC5FFDF383F3895F97CD042F8D7" bold="true" box="[199,295,1785,1810]" pageId="6" pageNumber="287">Biology.</emphasis>
JM observed the
<typeStatus id="54CA8875FFDF383F39ADF97FD305F8D7" box="[511,608,1786,1810]" pageId="6" pageNumber="287" type="paratype">paratype</typeStatus>
male nectaring on flowers of
<taxonomicName id="4C714D54FFDF383F3B99F97FD419F8D7" box="[971,1404,1785,1810]" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="287" phylum="Magnoliopsida" rank="species" species="donnell-smithii">
<emphasis id="B905EAC5FFDF383F3B99F97FD58CF8D7" box="[971,1257,1785,1810]" italics="true" pageId="6" pageNumber="287">Sommera donnell-smithii</emphasis>
(Rubiaceae)
</taxonomicName>
at about
<quantity id="4C899B32FFDF383F388EF898D046F8F0" box="[220,291,1821,1846]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="287" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">5–6 m</quantity>
off the ground. Lekking behavior, often seen in this genus, was not observed with this individual. The approximate time of collection was 11:00 am. Other butterflies nectaring or perching on this tree were
<taxonomicName id="4C714D54FFDF383F38C5F8E0D0E9F8BB" baseAuthorityName="Cramer, [" baseAuthorityYear="1777" box="[151,396,1893,1918]" class="Insecta" family="Pieridae" genus="Glutophrissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="drusilla">
<emphasis id="B905EAC5FFDF383F38C5F8E0D0E9F8BB" box="[151,396,1893,1918]" italics="true" pageId="6" pageNumber="287">Glutophrissa drusilla</emphasis>
</taxonomicName>
(Cramer, [1777]) (
<taxonomicName id="4C714D54FFDF383F3A3EF8E0D3ABF8BB" box="[620,718,1893,1918]" class="Insecta" family="Pieridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="family">Pieridae</taxonomicName>
),
<taxonomicName id="4C714D54FFDF383F3AB7F8E3D5A7F8BB" authority="(Bates, 1864)" authorityName="Bates" authorityYear="1864" baseAuthorityName="Bates" baseAuthorityYear="1864" box="[741,1218,1893,1918]" class="Insecta" family="Pieridae" genus="Enantia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="albania">
<emphasis id="B905EAC5FFDF383F3AB7F8E3D2FBF8BB" box="[741,926,1893,1918]" italics="true" pageId="6" pageNumber="287">Enantia albania</emphasis>
(Bates, 1864) (Pieridae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDF383F3C83F8E0D310F867" authority="(Godman &amp; Salvin, 1887)" authorityName="Godman &amp; Salvin" authorityYear="1887" baseAuthorityName="Godman &amp; Salvin" baseAuthorityYear="1887" class="Insecta" family="Lycaenidae" genus="Panthiades" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="ochus">
<emphasis id="B905EAC5FFDF383F3C83F8E0D4F9F8BB" box="[1233,1436,1893,1918]" italics="true" pageId="6" pageNumber="287">Panthiades ochus</emphasis>
(Godman &amp; Salvin, 1887) (Lycaenidae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDF383F3AD5F80CD5A4F867" authority="(Stoll, 1781)" authorityName="Stoll" authorityYear="1781" baseAuthorityName="Stoll" baseAuthorityYear="1781" box="[647,1217,1929,1954]" class="Insecta" family="Lycaenidae" genus="Cupathecla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="cupentus">
<emphasis id="B905EAC5FFDF383F3AD5F80CD21FF864" box="[647,890,1929,1954]" italics="true" pageId="6" pageNumber="287">Cupathecla cupentus</emphasis>
(Stoll, 1781) (Lycaenidae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDF383F3C81F80FD322F803" authority="Hewitson, 1870" authorityName="Hewitson" authorityYear="1870" class="Insecta" family="Lycaenidae" genus="Necyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="subSpecies" species="duellona" subSpecies="beltiana">
<emphasis id="B905EAC5FFDF383F3C81F80FD196F803" italics="true" pageId="6" pageNumber="287">Necyria duellona beltiana</emphasis>
Hewitson, 1870 (Riodinidae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDF383F3A06F828D56BF803" authority="leucophthalma" box="[596,1038,1965,1990]" class="Insecta" family="Nymphalidae" genus="Adelpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="subSpecies" species="leucophthalma" subSpecies="leucophthalma">
<emphasis id="B905EAC5FFDF383F3A06F828D56BF803" box="[596,1038,1965,1990]" italics="true" pageId="6" pageNumber="287">Adelpha leucophthalma leucophthalma</emphasis>
</taxonomicName>
(Latreille, [1809]) (
<taxonomicName id="4C714D54FFDF383F3CA1F828D4F5F803" box="[1267,1424,1965,1990]" class="Insecta" family="Nymphalidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
),
<taxonomicName id="4C714D54FFDF383F38C5F857D03BF82C" authorityName="Boisduval" authorityYear="1836" box="[151,350,2002,2025]" class="Insecta" family="Nymphalidae" genus="Marpesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="species" species="merops">
<emphasis id="B905EAC5FFDF383F38C5F857D03BF82C" box="[151,350,2002,2025]" italics="true" pageId="6" pageNumber="287">Marpesia merops</emphasis>
</taxonomicName>
(Doyère, [1840]) (
<taxonomicName id="4C714D54FFDF383F3A6FF854D3BEF82F" box="[573,731,2001,2026]" class="Insecta" family="Nymphalidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
). Other species observed in the vicinity included:
<taxonomicName id="4C714D54FFDF383E3D7BF857D229FF75" authority="Eitschberger &amp; Racheli, 1998" authorityName="Eitschberger &amp; Racheli" authorityYear="1998" class="Insecta" family="Pieridae" genus="Catasticta" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="7" lastPageNumber="288" order="Lepidoptera" pageId="6" pageNumber="287" phylum="Arthropoda" rank="subSpecies" species="sisamnus" subSpecies="smalli">
<emphasis id="B905EAC5FFDF383E3D7BF857D034FF75" italics="true" lastPageId="7" lastPageNumber="288" pageId="6" pageNumber="287">Catasticta sisamnus smalli</emphasis>
<bibRefCitation id="EFE04B26FFDE383E3930FF12D3AFFF75" author="Eitschberger, U. &amp; Racheli, T." box="[354,714,151,176]" pageId="7" pageNumber="288" refId="ref5286" refString="Eitschberger, U. &amp; Racheli, T. (1998) Catasticta studie" type="book" year="1998">Eitschberger &amp; Racheli, 1998</bibRefCitation>
(Pieridae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDE383E3B0DFF12D06DFF10" authority="(Butler, 1873)" authorityName="Butler" authorityYear="1873" baseAuthorityName="Butler" baseAuthorityYear="1873" class="Insecta" family="Pieridae" genus="Archonias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="subSpecies" species="brassolis" subSpecies="approximata">
<emphasis id="B905EAC5FFDE383E3B0DFF12D580FF75" box="[863,1253,151,176]" italics="true" pageId="7" pageNumber="288">Archonias brassolis approximata</emphasis>
(Butler, 1873) (Pieridae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDE383E394BFF39D22EFF10" authority="(Hewitson, 1870)" authorityName="Hewitson" authorityYear="1870" baseAuthorityName="Hewitson" baseAuthorityYear="1870" box="[281,843,188,213]" class="Insecta" family="Lycaenidae" genus="Thestius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="epopea">
<emphasis id="B905EAC5FFDE383E394BFF39D0B1FF11" box="[281,468,188,213]" italics="true" pageId="7" pageNumber="288">Thestius epopea</emphasis>
(Hewitson, 1870) (Lycaenidae)
</taxonomicName>
,
<taxonomicName id="4C714D54FFDE383E3B09FF39D021FF3D" authority="LeMoult &amp; Real, 1962" authorityName="LeMoult &amp; Real" authorityYear="1962" class="Insecta" family="Nymphalidae" genus="Morpho" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="subSpecies" species="helenor" subSpecies="veragua">
<emphasis id="B905EAC5FFDE383E3B09FF39D5E5FF11" box="[859,1152,188,213]" italics="true" pageId="7" pageNumber="288">Morpho helenor veragua</emphasis>
LeMoult &amp; Réal, 1962 (Nymphalidae)
</taxonomicName>
, and
<taxonomicName id="4C714D54FFDE383E392CFF65D3B1FF3D" authority="Doubleday" authorityName="Doubleday" box="[382,724,223,248]" class="Insecta" family="Nymphalidae" genus="Oressinoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="typhla">
<emphasis id="B905EAC5FFDE383E392CFF65D334FF3D" box="[382,593,223,248]" italics="true" pageId="7" pageNumber="288">Oressinoma typhla</emphasis>
Doubleday
</taxonomicName>
, [1849] (
<taxonomicName id="4C714D54FFDE383E3B6AFF5AD2B3FF3D" box="[824,982,223,248]" class="Insecta" family="Nymphalidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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