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<document id="B2546ECF2C7942A229C9F3402230DE7C" ID-DOI="10.11646/zootaxa.5575.4.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="14762844" ID-ZooBank="170DB620-EF80-4D0B-968E-23790471262E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738152554261" checkinUser="plazi" docAuthor="Yang, Yi &amp; Hu, Jia-Yao" docDate="2025" docId="03D887A6FF95FFC889C98A8FFB7FFE6E" docLanguage="en" docName="zootaxa.5575.4.8.pdf" docOrigin="Zootaxa 5575 (4)" docSource="https://doi.org/10.11646/zootaxa.5575.4.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nazeris zhaotongus Yang &amp; Hu 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="581" masterDocId="FFE1FFDEFF96FFCC895E8A18FF9EFFC3" masterDocTitle="Four new species of Nazeris Fauvel from Wumengshan Nature Reserve, Yunnan, China (Coleoptera, Staphylinidae, Paederinae)" masterLastPageNumber="587" masterPageNumber="577" pageNumber="580" updateTime="1738153034235" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D9031E64CD613A8309442EAC0D8FBA4E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0CDEC746D219FD749AD0C93BFAF312CB">
<mods:title id="53F1FD2CE36C99D83EA3F787F96D01B6">Four new species of Nazeris Fauvel from Wumengshan Nature Reserve, Yunnan, China (Coleoptera, Staphylinidae, Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="C642FC3FD887A4F9DA7C64E4255B9955" type="personal">
<mods:role id="DF09D443A172D049B21D5F8A3234D04A">
<mods:roleTerm id="AB07AA377B187DA398A2973A22701DA8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F65C8C5E026044A3EE62B04024FF3815">Yang, Yi</mods:namePart>
<mods:nameIdentifier id="BF96ECA8F8DABB04AEC728BD185B6448" type="ORCID">0000-0003-1132-9103</mods:nameIdentifier>
<mods:affiliation id="240E76DC6552EF1A07F30497725335B0">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai, 200234, China.</mods:affiliation>
<mods:nameIdentifier id="4FD39FCA1355958BDBC470A8D55C1552" type="email">931009245@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2EBF4607DE6ABD3C96A14C73EC33A6F4" type="personal">
<mods:role id="68640F9DFE8F0EE888EBB416F5B7B371">
<mods:roleTerm id="BE7553FF14C3612DD4CF12F29B070F07">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F8442464200D44EA9353C866B107756">Hu, Jia-Yao</mods:namePart>
<mods:nameIdentifier id="6BE1559761E8634B0BCBC4138902A5A6" type="ORCID">0000-0002-9861-9551</mods:nameIdentifier>
<mods:affiliation id="191BC96375515DEBFC83CCF047DDC4DE">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai, 200234, China.</mods:affiliation>
<mods:nameIdentifier id="EB7FDC0D626696D62EB04442D2B80161" type="email">hujiayao@shnu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0FF8EE57006B0AE422C177E89F48B761">text</mods:typeOfResource>
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<mods:titleInfo id="A7B790286B72152AA7D75E75FFA4EC09">
<mods:title id="C8F83D42AAC78211AB4244A96A276273">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="564FF0E1FE12F2C486002AB4F538EA6C">
<mods:date id="8A6C43FACD2C7ED6DEF8D878D4419C60">2025</mods:date>
<mods:detail id="8D884277A7871F8A12B335C1ED6A95D5" type="pubDate">
<mods:number id="026454EAD416731D62693399526B458C">2025-01-28</mods:number>
</mods:detail>
<mods:detail id="B1EFA8E0CBE70D1159DC423569330DB0" type="volume">
<mods:number id="35BD41706578141B54981438358AB70A">5575</mods:number>
</mods:detail>
<mods:detail id="83925E37C2502056C163B8D5D57D848E" type="issue">
<mods:number id="1C739C9FC96DB23AACBA2B378696BE92">4</mods:number>
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<mods:start id="4BB81D465B25C034F6973E357B9097C1">577</mods:start>
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<mods:location id="49E73E7546B8497A9897D83717C524A7">
<mods:url id="36F52478E984995642816501215CD5AE">https://doi.org/10.11646/zootaxa.5575.4.8</mods:url>
</mods:location>
<mods:classification id="3897B336A105287E8C0EC07FBF108BAB">journal article</mods:classification>
<mods:identifier id="52BE864F3458B1D2561B5B829ED47ED7" type="DOI">10.11646/zootaxa.5575.4.8</mods:identifier>
<mods:identifier id="01748D4285D28ED85D5078030DF2A97E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DA61A95482186D68C1966549190FB194" type="Zenodo-Dep">14762844</mods:identifier>
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<treatment id="03D887A6FF95FFC889C98A8FFB7FFE6E" ID-DOI="http://doi.org/10.5281/zenodo.14762852" ID-Zenodo-Dep="14762852" LSID="urn:lsid:plazi:treatment:03D887A6FF95FFC889C98A8FFB7FFE6E" httpUri="http://treatment.plazi.org/id/03D887A6FF95FFC889C98A8FFB7FFE6E" lastPageId="4" lastPageNumber="581" pageId="3" pageNumber="580">
<subSubSection id="C36B653BFF95FFCF89C98A8FFDE0FF72" box="[151,638,151,178]" pageId="3" pageNumber="580" type="nomenclature">
<paragraph id="8BCE36B0FF95FFCF89C98A8FFDE0FF72" blockId="3.[151,638,151,213]" box="[151,638,151,178]" pageId="3" pageNumber="580">
<heading id="D08681DCFF95FFCF89C98A8FFDE0FF72" bold="true" box="[151,638,151,178]" fontSize="11" level="1" pageId="3" pageNumber="580" reason="1">
<emphasis id="B905EAA2FF95FFCF89C98A8FFDE0FF72" bold="true" box="[151,638,151,178]" pageId="3" pageNumber="580">
<taxonomicName id="4C714D33FF95FFCF89C98A8FFD87FF72" authority="Yang &amp; Hu" authorityName="Yang &amp; Hu" authorityYear="2025" box="[151,537,151,178]" class="Insecta" family="Staphylinidae" genus="Nazeris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="zhaotongus" status="sp. nov.">
<emphasis id="B905EAA2FF95FFCF89C98A8FFE1BFF71" bold="true" box="[151,389,151,178]" italics="true" pageId="3" pageNumber="580">Nazeris zhaotongus</emphasis>
Yang &amp; Hu
</taxonomicName>
<taxonomicNameLabel id="A23657D9FF95FFCF8B7F8A8FFDE0FF72" box="[545,638,151,177]" pageId="3" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B653BFF95FFCF89C98AA3FEDCFF16" box="[151,322,187,213]" pageId="3" pageNumber="580" type="description">
<paragraph id="8BCE36B0FF95FFCF89C98AA3FEDCFF16" blockId="3.[151,638,151,213]" box="[151,322,187,213]" pageId="3" pageNumber="580">
<figureCitation id="134A2A35FF95FFCF89C98AA3FF6FFF16" box="[151,241,187,213]" captionStart="FIGURE 1" captionStartId="1.[152,255,1934,1958]" captionTargetBox="[157,1424,1254,1899]" captionTargetId="figure-307@1.[151,1436,1242,1908]" captionTargetPageId="1" captionText="FIGURE 1. Habitus A Nazeris wumengensis sp. nov.; B N. zhaotongus sp. nov.; C N. guohaoi sp. nov.; D N. serratimarginatus sp. nov. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14762848" httpUri="https://zenodo.org/record/14762848/files/figure.png" pageId="3" pageNumber="580">Figs 1B</figureCitation>
,
<figureCitation id="134A2A35FF95FFCF89A38AA3FEDCFF16" box="[253,322,187,213]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">3A–G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B653BFF95FFCF89C98B1BFEABFEA6" pageId="3" pageNumber="580" type="materials_examined">
<paragraph id="8BCE36B0FF95FFCF89C98B1BFEABFEA6" blockId="3.[151,1436,259,357]" pageId="3" pageNumber="580">
<emphasis id="B905EAA2FF95FFCF89C98B1BFEDBFEDE" bold="true" box="[151,325,259,285]" pageId="3" pageNumber="580">Type material.</emphasis>
<materialsCitation id="3B193CEDFF95FFCF88158B1BFEABFEA6" collectingDate="2023-07-15" collectionCode="R, SNUC" collectorName="Wei Guohao" country="China" county="China" elevation="1952" latitude="28.20714" location="Wumeng Mt." longLatPrecision="1" longitude="103.91373" municipality="Zhaotong" pageId="3" pageNumber="580" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B905EAA2FF95FFCF88158B1BFDB1FEDE" bold="true" box="[331,559,259,285]" pageId="3" pageNumber="580">
<typeStatus id="54CA8812FF95FFCF88158B1BFE22FEDE" box="[331,444,259,285]" pageId="3" pageNumber="580" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3667620FF95FFCF88968B1BFDB4FEDE" box="[456,554,259,285]" name="China" pageId="3" pageNumber="580">CHINA</collectingCountry>
:
</emphasis>
male: “
<collectingCounty id="62AF4E3CFF95FFCF8BDB8B1BFD50FEDE" box="[645,718,259,285]" pageId="3" pageNumber="580">China</collectingCounty>
:
<collectingRegion id="49B5F852FF95FFCF8B868B1CFCAFFEDD" box="[728,817,260,286]" country="China" name="Yunnan" pageId="3" pageNumber="580">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BAAACCAFF95FFCF8A658B1BFC37FEDE" box="[827,937,259,285]" pageId="3" pageNumber="580">Zhaotong</collectingMunicipality>
, Sanjiangkou
<taxonomicNameLabel id="A23657D9FF95FFCF8D198B1CFB1AFEDD" box="[1095,1156,260,286]" pageId="3" pageNumber="580" rank="subSpecies">
N.
<collectionCode id="ED60AE75FF95FFCF8D378B1CFB1FFEDD" box="[1129,1153,260,286]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="580">R</collectionCode>
.
</taxonomicNameLabel>
,
<location id="8EAE606BFF95FFCF8DCE8B1CFAB9FEDD" LSID="urn:lsid:plazi:treatment:03D887A6FF95FFC889C98A8FFB7FFE6E:8EAE606BFF95FFCF8DCE8B1CFAB9FEDD" box="[1168,1319,260,286]" country="China" county="China" latitude="28.20714" longLatPrecision="1" longitude="103.91373" municipality="Zhaotong" name="Wumeng Mt." pageId="3" pageNumber="580" stateProvince="Yunnan">Wumeng Mt.</location>
,
<location id="8EAE606BFF95FFCF8C6C8B1BFA06FEDE" LSID="urn:lsid:plazi:treatment:03D887A6FF95FFC889C98A8FFB7FFE6E:8EAE606BFF95FFCF8C6C8B1BFA06FEDE" box="[1330,1432,259,285]" country="China" county="China" latitude="28.20714" longLatPrecision="1" longitude="103.91373" municipality="Zhaotong" name="Laziping" pageId="3" pageNumber="580" stateProvince="Yunnan">Laziping</location>
,
<geoCoordinate id="EE455077FF95FFCF89C98B3FFEA2FE81" box="[151,316,295,322]" degrees="28" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="580" precision="1" seconds="25.7" value="28.20714">28°12’25.7’’N</geoCoordinate>
,
<geoCoordinate id="EE455077FF95FFCF88178B3FFE66FE81" box="[329,504,295,322]" degrees="103" direction="east" minutes="54" orientation="longitude" pageId="3" pageNumber="580" precision="1" seconds="49.4" value="103.91373">103°54’49.4’’E</geoCoordinate>
,
<quantity id="4C899B55FF95FFCF8B5B8B3FFDCBFE81" box="[517,597,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.952" pageId="3" pageNumber="580" unit="m" value="1952.0">
<elevation id="005CD183FF95FFCF8B5B8B3FFDCBFE81" box="[517,597,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.952" pageId="3" pageNumber="580" unit="m" value="1952.0">1952m</elevation>
</quantity>
,
<date id="FFCF1070FF95FFCF8B3C8B3FFD7AFE81" box="[610,740,295,322]" pageId="3" pageNumber="580" value="2023-07-15">
<collectingDate id="EF8BE998FF95FFCF8B3C8B3FFD7AFE81" box="[610,740,295,322]" pageId="3" pageNumber="580" value="2023-07-15">15.vii.2023</collectingDate>
</date>
,
<collectorName id="26845366FF95FFCF8BAE8B3FFCE0FE82" box="[752,894,295,321]" pageId="3" pageNumber="580">Wei Guohao</collectorName>
leg. [
<emphasis id="B905EAA2FF95FFCF8A9E8B30FF2DFEA0" bold="true" pageId="3" pageNumber="580">云南省Dzdz市三江口保护区乌ae山OiŦ 坪</emphasis>
]” (
<collectionCode id="ED60AE75FF95FFCF89868B53FEB7FEA6" box="[216,297,331,357]" pageId="3" pageNumber="580">SNUC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF0E6638FF95FFCF89C68F2AFCA1FAAD" ID-DOI="http://doi.org/10.5281/zenodo.14762854" ID-Zenodo-Dep="14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580" startId="3.[152,255,1330,1354]" targetBox="[157,1434,393,1297]" targetPageId="3" targetType="figure">
<paragraph id="8BCE36B0FF95FFCF89C68F2AFCA1FAAD" blockId="3.[151,1436,1330,1390]" pageId="3" pageNumber="580">
<emphasis id="B905EAA2FF95FFCF89C68F2AFE89FA89" bold="true" box="[152,279,1330,1354]" pageId="3" pageNumber="580">FIGURE 3.</emphasis>
<taxonomicName id="4C714D33FF95FFCF88438F2BFE7DFA89" authorityName="Yang &amp; Hu" authorityYear="2025" box="[285,483,1330,1354]" class="Insecta" family="Staphylinidae" genus="Nazeris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="zhaotongus" status="sp. nov.">
<emphasis id="B905EAA2FF95FFCF88438F2BFE7DFA89" box="[285,483,1330,1354]" italics="true" pageId="3" pageNumber="580">Nazeris zhaotongus</emphasis>
</taxonomicName>
<emphasis id="B905EAA2FF95FFCF88B78F2AFDCFFA89" bold="true" box="[489,593,1330,1354]" pageId="3" pageNumber="580">
<taxonomicNameLabel id="A23657D9FF95FFCF88B78F2AFDA7FA89" box="[489,569,1330,1354]" pageId="3" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
Forebody;
<emphasis id="B905EAA2FF95FFCF8B9A8F2AFD4BFA89" bold="true" box="[708,725,1330,1354]" pageId="3" pageNumber="580">B</emphasis>
male sternite VII;
<emphasis id="B905EAA2FF95FFCF8ACF8F2AFC3DFA89" bold="true" box="[913,931,1330,1354]" pageId="3" pageNumber="580">C</emphasis>
male sternite VIII;
<emphasis id="B905EAA2FF95FFCF8D368F2AFB09FA89" bold="true" box="[1128,1175,1330,1354]" pageId="3" pageNumber="580">D, E</emphasis>
aedeagus in ventral view;
<emphasis id="B905EAA2FF95FFCF89C98F4EFF5BFAAD" bold="true" box="[151,197,1366,1390]" pageId="3" pageNumber="580">F, G</emphasis>
aedeagus in lateral view. Scale bars:
<emphasis id="B905EAA2FF95FFCF8B618F4EFDCFFAAD" bold="true" box="[575,593,1366,1390]" pageId="3" pageNumber="580">A</emphasis>
1.0 mm;
<emphasis id="B905EAA2FF95FFCF8BEC8F4EFD7AFAAD" bold="true" box="[690,740,1366,1390]" pageId="3" pageNumber="580">B–G</emphasis>
0.5 mm.
</paragraph>
</caption>
<subSubSection id="C36B653BFF95FFCF89998FD8FB2CF9E0" pageId="3" pageNumber="580" type="diagnosis">
<paragraph id="8BCE36B0FF95FFCF89998FD8FB2CF9E0" blockId="3.[151,1437,1472,2039]" pageId="3" pageNumber="580">
<emphasis id="B905EAA2FF95FFCF89998FD8FEDCFA19" bold="true" box="[199,322,1472,1498]" pageId="3" pageNumber="580">Diagnosis.</emphasis>
Body reddish brown; head with umbilicate punctation; abdomen with fine microsculpture on all tergite; aedeagal ventral process widened near middle in ventral view, with small triangular excision at apex; dorso-lateral apophyses wide, with round apex in ventral view, not reaching apex of ventral process.
</paragraph>
</subSubSection>
<subSubSection id="C36B653BFF95FFC889998C34FDE9FF16" lastPageId="4" lastPageNumber="581" pageId="3" pageNumber="580" type="description">
<paragraph id="8BCE36B0FF95FFCF89998C34FCF5F984" blockId="3.[151,1437,1472,2039]" box="[199,875,1580,1607]" pageId="3" pageNumber="580">
<emphasis id="B905EAA2FF95FFCF89998C34FEC7F985" bold="true" box="[199,345,1580,1606]" pageId="3" pageNumber="580">Description.</emphasis>
Body length
<quantity id="4C899B55FF95FFCF88AE8C34FDD7F984" box="[496,585,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="580" unit="mm" value="5.8">5.8 mm</quantity>
; forebody length
<quantity id="4C899B55FF95FFCF8A508C35FCF9F984" box="[782,871,1581,1607]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="580" unit="mm" value="2.9">2.9 mm</quantity>
.
</paragraph>
<paragraph id="8BCE36B0FF95FFCF89998C48FC27F9A9" blockId="3.[151,1437,1472,2039]" box="[199,953,1616,1643]" pageId="3" pageNumber="580">
Body (
<figureCitation id="134A2A35FF95FFCF884D8C48FEF6F9A8" box="[275,360,1616,1643]" captionStart="FIGURE 1" captionStartId="1.[152,255,1934,1958]" captionTargetBox="[157,1424,1254,1899]" captionTargetId="figure-307@1.[151,1436,1242,1908]" captionTargetPageId="1" captionText="FIGURE 1. Habitus A Nazeris wumengensis sp. nov.; B N. zhaotongus sp. nov.; C N. guohaoi sp. nov.; D N. serratimarginatus sp. nov. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14762848" httpUri="https://zenodo.org/record/14762848/files/figure.png" pageId="3" pageNumber="580">Fig. 1B</figureCitation>
) reddish brown; antennae and legs yellowish brown.
</paragraph>
<paragraph id="8BCE36B0FF95FFCF89998C6CFA03F971" blockId="3.[151,1437,1472,2039]" pageId="3" pageNumber="580">
Head (
<figureCitation id="134A2A35FF95FFCF884C8C6CFEF5F94C" box="[274,363,1652,1679]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">Fig. 3A</figureCitation>
) 1.08 times as long as wide; punctation moderately dense and coarse, distinctly umbilicate, but not confluent, interstices lacking microsculpture; postocular portion approximately 1.9 times as long as eye length.
</paragraph>
<paragraph id="8BCE36B0FF95FFCF89998CA5FD87F8DD" blockId="3.[151,1437,1472,2039]" pageId="3" pageNumber="580">
Pronotum (
<figureCitation id="134A2A35FF95FFCF88198CA4FE3EF914" box="[327,416,1724,1751]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">Fig. 3A</figureCitation>
) 1.17 times as long as wide, as long as and 0.92 times as broad as head; punctation non-umbilicate, moderately dense, less coarse than that of head; midline with long and narrow impunctate elevation; interstices lacking microsculpture.
</paragraph>
<paragraph id="8BCE36B0FF95FFCF89998D30FBCFF8A5" blockId="3.[151,1437,1472,2039]" pageId="3" pageNumber="580">
Elytra (
<figureCitation id="134A2A35FF95FFCF88448D30FEECF880" box="[282,370,1832,1859]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">Fig. 3A</figureCitation>
) 0.72 times as long as wide, 0.57 times as long and 0.92 times as broad as pronotum; punctation more dense and less coarse than that of pronotum; interstices lacking microsculpture.
</paragraph>
<paragraph id="8BCE36B0FF95FFCF89998D68FBFBF86D" blockId="3.[151,1437,1472,2039]" pageId="3" pageNumber="580">Abdomen with punctation dense and rather coarse on tergites III–IV, dense and less coarse on tergite V, moderately dense and fine on tergites VI–VIII; with fine microsculpture on all tergites.</paragraph>
<paragraph id="8BCE36B0FF95FFC889998DA0FDE9FF16" blockId="3.[151,1437,1472,2039]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="581" pageId="3" pageNumber="580">
Male. Sternite VII (
<figureCitation id="134A2A35FF95FFCF88F98DA0FE60F810" box="[423,510,1976,2003]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">Fig. 3B</figureCitation>
) with posterior margin slightly protruding in middle. Sternite VIII (
<figureCitation id="134A2A35FF95FFCF8C5C8DA0FAC7F810" box="[1282,1369,1976,2003]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">Fig. 3C</figureCitation>
) with triangular posterior excision. Aedeagus (
<figureCitation id="134A2A35FF95FFCF8B3E8DC4FD41F834" box="[608,735,2012,2039]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="3" pageNumber="580">Figs 3D–G</figureCitation>
) moderately sclerotized; ventral process widened near middle in ventral view, with small triangular excision at apex; dorso-lateral apophyses wide, with round apex in ventral view, not reaching apex of ventral process.
</paragraph>
</subSubSection>
<subSubSection id="C36B653BFF92FFC889998AC6FAEDFE4A" pageId="4" pageNumber="581" type="distribution">
<paragraph id="8BCE36B0FF92FFC889998AC6FC09FEDE" blockId="4.[151,1437,151,429]" pageId="4" pageNumber="581">
<emphasis id="B905EAA2FF92FFC889998AC6FDB3FF3A" bold="true" box="[199,557,222,249]" pageId="4" pageNumber="581">Distribution and habitat data.</emphasis>
The species is known only from Wumengshan
<taxonomicNameLabel id="A23657D9FF92FFC88D1F8AC7FBE3FF3A" box="[1089,1149,223,249]" pageId="4" pageNumber="581" rank="subSpecies">N. R.</taxonomicNameLabel>
in northeast
<collectingRegion id="49B5F852FF92FFC88C508AC7FAF9FF3A" box="[1294,1383,223,249]" country="China" name="Yunnan" pageId="4" pageNumber="581">Yunnan</collectingRegion>
. The specimen was collected by sifting leaf litter at an altitude of
<quantity id="4C899B55FF92FFC88A658B1AFC0DFEDE" box="[827,915,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.952" pageId="4" pageNumber="581" unit="m" value="1952.0">1952 m</quantity>
.
</paragraph>
<paragraph id="8BCE36B0FF92FFC889998B3EFAEDFE4A" blockId="4.[151,1437,151,429]" pageId="4" pageNumber="581">
<emphasis id="B905EAA2FF92FFC889998B3EFE2EFE82" bold="true" box="[199,432,294,321]" pageId="4" pageNumber="581">Comparative notes.</emphasis>
The new species is distinguished from all the known species of
<taxonomicName id="4C714D33FF92FFC88D218B3FFB4BFE83" authorityName="Fauvel" authorityYear="1873" box="[1151,1237,295,320]" class="Insecta" family="Staphylinidae" genus="Nazeris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAA2FF92FFC88D218B3FFB4BFE83" box="[1151,1237,295,320]" italics="true" pageId="4" pageNumber="581">Nazeris</emphasis>
</taxonomicName>
from
<collectingRegion id="49B5F852FF92FFC88C498B3FFAF3FE82" box="[1303,1389,295,321]" country="China" name="Yunnan" pageId="4" pageNumber="581">Yunnan</collectingRegion>
and adjacent area by the abdomen with fine microsculpture on all tergites; by the sternite VII with posterior margin slightly protrude in the middle (
<figureCitation id="134A2A35FF92FFC888A58B77FDCDFE4A" box="[507,595,367,393]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="4" pageNumber="581">Fig. 3B</figureCitation>
); and by the wide dorso-lateral apophyses in ventral view (
<figureCitation id="134A2A35FF92FFC88DB68B77FAF8FE4A" box="[1256,1382,367,393]" captionStart="FIGURE 3" captionStartId="3.[152,255,1330,1354]" captionTargetBox="[157,1434,393,1297]" captionTargetId="figure-55@3.[151,1436,381,1306]" captionTargetPageId="3" captionText="FIGURE 3. Nazeris zhaotongus sp. nov. A Forebody; B male sternite VII; C male sternite VIII; D, E aedeagus in ventral view; F, G aedeagus in lateral view. Scale bars: A 1.0 mm; B–G 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14762854" httpUri="https://zenodo.org/record/14762854/files/figure.png" pageId="4" pageNumber="581">Figs 3D, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B653BFF92FFC889998B8BFB7FFE6E" box="[199,1249,403,429]" pageId="4" pageNumber="581" type="etymology">
<paragraph id="8BCE36B0FF92FFC889998B8BFB7FFE6E" blockId="4.[151,1437,151,429]" box="[199,1249,403,429]" pageId="4" pageNumber="581">
<emphasis id="B905EAA2FF92FFC889998B8BFED0FE6E" bold="true" box="[199,334,403,429]" pageId="4" pageNumber="581">Etymology.</emphasis>
The specific epithet is derived from the name of the
<typeStatus id="54CA8812FF92FFC88AC18B8BFC51FE6E" box="[927,975,403,429]" pageId="4" pageNumber="581">type</typeStatus>
locality: Zhaotong City.
</paragraph>
</subSubSection>
</treatment>
</document>
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