234 lines
24 KiB
XML
234 lines
24 KiB
XML
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<mods:title id="146CAD803BA40406B982805A6CFF6645">New synonyms and new species of European aquatic dance flies (Diptera, Empididae)</mods:title>
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<mods:namePart id="4BEFCCA94880EA667BD04C5B2723A0E2">Wagner, Rüdiger</mods:namePart>
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<mods:nameIdentifier id="B685F2E78CCD561D7CF7D7CB832773C4" type="ORCID">0000-0002-2024-1827</mods:nameIdentifier>
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<mods:affiliation id="9039BB069733B4318AF8A4DF2381A67B">University of Kassel, FB 10, Biology-Zoology, Heinrich-Plett-Strasse 40, D- 34132 Kassel, Germany; Parkstrasse 65, D- 36110 Schlitz, Germany. ruediger 2. wagner @ t-online. de; http: // orcid. org / 0000 - 0002 - 2024 - 1827</mods:affiliation>
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<mods:nameIdentifier id="63814BA5FF657C003DBE20EBEC1F4A0D" type="email">ruediger2.wagner@t-online.de</mods:nameIdentifier>
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<mods:namePart id="6BF89505E1AFFDE3AD96AFD2D1C0FD8A">Plant, Adrian</mods:namePart>
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<mods:affiliation id="C1ED4C244B3CB8BAECF21D28F9E29A66">Division of Research Facilitation and Dissemination, Mahasarakham University, Mahasarakham, 44150, Thailand eanddrs @ gmail. com</mods:affiliation>
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<mods:namePart id="AB21BABA616B06E24CF5B7C4D924BD7A">Ivković, Marija</mods:namePart>
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<mods:affiliation id="584BD416C95312C774EFD9A5C2196308">Department of Biology, Faculty of Science, Rooseveltov trg 6, 10000 Zagreb, Croatia</mods:affiliation>
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<mods:namePart id="070213D623DAC0173AE89B0EAEF26771">Gattolliat, Jean-Luc</mods:namePart>
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<mods:affiliation id="0B153C20A52E547F11DF7DF1ECF84630">Musée cantonal de zoologie, Palais de Rumine, Pl. de la Riponne 6, CH- 1014 Lausanne, Switzerland</mods:affiliation>
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<mods:title id="D210E21B1D9923313E89551F06E33CBC">Zootaxa</mods:title>
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<treatment id="03D687F2EB6DFFE1FF53FCFEAAAD5DEC" ID-DOI="http://doi.org/10.5281/zenodo.7471840" ID-GBIF-Taxon="207927119" ID-Zenodo-Dep="7471840" LSID="urn:lsid:plazi:treatment:03D687F2EB6DFFE1FF53FCFEAAAD5DEC" httpUri="http://treatment.plazi.org/id/03D687F2EB6DFFE1FF53FCFEAAAD5DEC" lastPageNumber="438" pageId="11" pageNumber="438">
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<heading id="D0888188EB6DFFE1FF53FCFEA9B25980" bold="true" box="[151,647,895,922]" fontSize="11" level="1" pageId="11" pageNumber="438" reason="1">
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<taxonomicName id="4C7F4D67EB6DFFE1FF53FCFEA9175980" ID-CoL="BDJJS" authority="Wagner" authorityName="Wagner & Plant & Ivković & Gattolliat" authorityYear="2022" box="[151,546,895,922]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="remschakae" status="sp. nov.">
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<emphasis id="B90BEAF6EB6DFFE1FF53FCFEAA8D5980" bold="true" box="[151,440,895,922]" italics="true" pageId="11" pageNumber="438">Phaeobalia remschakae</emphasis>
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Wagner
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</taxonomicName>
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<taxonomicNameLabel id="A238578DEB6DFFE1FDEDFC01A9B25980" box="[553,647,896,922]" pageId="11" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C365656FEB6DFFE1FF53FC25AA2759A4" box="[151,274,932,958]" pageId="11" pageNumber="438" type="description">
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<paragraph id="8BC036E4EB6DFFE1FF53FC25AA2759A4" blockId="11.[151,647,895,958]" box="[151,274,932,958]" pageId="11" pageNumber="438">
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(
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<figureCitation id="13442A61EB6DFFE1FF5BFC25AA3F59A4" box="[159,266,932,958]" captionStart="FIGURE 9" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[270,1271,182,1920]" captionTargetId="figure-22@12.[270,1318,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Phaeobalia species. a, P. pokornyi Mik, slide with lectotype; b, male terminalia, lateral view; c, P. remschakae Wagner sp. nov., slide with holotype; d, male terminalia, lateral view; e, epandrium with clasping cercus, inner view. (photos RW)." figureDoi="http://doi.org/10.5281/zenodo.7471844" httpUri="https://zenodo.org/record/7471844/files/figure.png" pageId="11" pageNumber="438">Fig. 9c–e</figureCitation>
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)
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<subSubSection id="C365656FEB6DFFE1FF53FC6DA9EC5EC4" pageId="11" pageNumber="438" type="materials_examined">
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<emphasis id="B90BEAF6EB6DFFE1FF53FC6DAA705E1C" bold="true" box="[151,325,1004,1030]" pageId="11" pageNumber="438">Type material.</emphasis>
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<materialsCitation id="3B173CB9EB6DFFE1FE89FC6DAEA85E54" ID-GBIF-Occurrence="4003032322" collectingDate="2019-07-24" collectionCode="WGS" collectorName="Haseke & Remschak" elevation="2" latitude="46.97996" location="Rieseltraufe am Langtalweg" longLatPrecision="1" longitude="12.76495" pageId="11" pageNumber="438" specimenCode="WGS 84" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54C48846EB6DFFE1FE89FC6DAAC65E1C" box="[333,499,1004,1030]" pageId="11" pageNumber="438" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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♁ (slide mounted): red label:
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<taxonomicName id="4C7F4D67EB6DFFE1FC8CFC6DAFE35E1C" authority=", RW, 2021" authorityName=", RW" authorityYear="2021" box="[840,1238,1004,1031]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="remschakae">
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<emphasis id="B90BEAF6EB6DFFE1FC8CFC6DAF635E1C" box="[840,1110,1004,1030]" italics="true" pageId="11" pageNumber="438">Phaeobalia remschakae</emphasis>
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||
, RW, 2021
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</taxonomicName>
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,
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<typeStatus id="54C48846EB6DFFE1FB26FC6DAE615E1C" box="[1250,1364,1004,1030]" pageId="11" pageNumber="438" type="holotype">holotypus</typeStatus>
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; small white label below red label:
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<collectingDate id="EF85E9CCEB6DFFE1FE10FB91A95D5E30" box="[468,616,1040,1066]" pageId="11" pageNumber="438" value="2019-07-24">24 July 2019</collectingDate>
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</date>
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, leg.
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||
<collectorName id="268A5332EB6DFFE1FD67FB91A9C05E30" box="[675,757,1040,1066]" pageId="11" pageNumber="438">Haseke</collectorName>
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&
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<collectorName id="268A5332EB6DFFE1FCDCFB91A8A55E30" box="[792,912,1040,1066]" pageId="11" pageNumber="438">Remschak</collectorName>
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; large white label: A[ustria], N[ational] P[ark], Hohe Tauern, [Ĝnita 05]
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<location id="8EA0603FEB6DFFE1FE10FBB5A82E5E54" LSID="urn:lsid:plazi:treatment:03D687F2EB6DFFE1FF53FCFEAAAD5DEC:8EA0603FEB6DFFE1FE10FBB5A82E5E54" box="[468,795,1076,1103]" latitude="46.97996" longLatPrecision="1" longitude="12.76495" name="Rieseltraufe am Langtalweg" pageId="11" pageNumber="438">Rieseltraufe am Langtalweg</location>
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,
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(
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<specimenCode id="DBD99E9FEB6DFFE1FBB1FBB5AFD45E54" box="[1141,1249,1076,1102]" collectionCode="WGS" pageId="11" pageNumber="438">WGS 84</specimenCode>
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),
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a.s.l.
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</elevation>
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</materialsCitation>
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<materialsCitation id="3B173CB9EB6DFFE1FF5CFBD9A9E05EC4" ID-GBIF-Occurrence="4003032317" collectingDate="1982-07-03" collectionCode="NHMW, RW" collectorName="Zwick. The & Natural History Museum" county="Quellbach" location="Schruns" municipality="Silbertal" pageId="11" pageNumber="438" specimenCount="4" stateProvince="Vorarlberg" typeStatus="paratype">
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<emphasis id="B90BEAF6EB6DFFE1FF5CFBD9AA095E68" bold="true" box="[152,316,1112,1138]" pageId="11" pageNumber="438">
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<typeStatus id="54C48846EB6DFFE1FF5CFBD9AA065E68" box="[152,307,1112,1138]" pageId="11" pageNumber="438" type="paratype">PARATYPE</typeStatus>
|
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:
|
||
</emphasis>
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||
♁ (slide mounted): red label
|
||
<taxonomicName id="4C7F4D67EB6DFFE1FD52FBD9A8905E68" authorityName=", RW" authorityYear="2021" box="[662,933,1112,1138]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="remschakae">
|
||
<emphasis id="B90BEAF6EB6DFFE1FD52FBD9A8905E68" box="[662,933,1112,1138]" italics="true" pageId="11" pageNumber="438">Phaeobalia remschakae</emphasis>
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</taxonomicName>
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<typeStatus id="54C48846EB6DFFE1FC6AFBD8AF2E5E69" box="[942,1051,1113,1139]" pageId="11" pageNumber="438" type="paratype">paratypus</typeStatus>
|
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RW 2021; white label: A[ustria],
|
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<collectingRegion id="49BBF806EB6DFFE1FF53FBFDAA245E8C" box="[151,273,1148,1174]" country="Austria" name="Vorarlberg" pageId="11" pageNumber="438">Vorarlberg</collectingRegion>
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,
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||
<collectingCounty id="62A14E68EB6DFFE1FED9FBFDAAA55E8C" box="[285,400,1148,1174]" pageId="11" pageNumber="438">Quellbach</collectingCounty>
|
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bei
|
||
<collectingMunicipality id="6BA4AC9EEB6DFFE1FE05FBFDA9155E8C" box="[449,544,1148,1174]" pageId="11" pageNumber="438">Silbertal</collectingMunicipality>
|
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bei
|
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<location id="8EA0603FEB6DFFE1FD96FBFDA9995E8C" LSID="urn:lsid:plazi:treatment:03D687F2EB6DFFE1FF53FCFEAAAD5DEC:8EA0603FEB6DFFE1FD96FBFDA9995E8C" box="[594,684,1148,1174]" county="Quellbach" municipality="Silbertal" name="Schruns" pageId="11" pageNumber="438" stateProvince="Vorarlberg">Schruns</location>
|
||
[=spring stream near Silbertal, near Schruns], 25.6.–
|
||
<date id="FFC11024EB6DFFE1FAC5FBFDAE525E8D" box="[1281,1383,1148,1175]" pageId="11" pageNumber="438" value="1982-07-03">
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<collectingDate id="EF85E9CCEB6DFFE1FAC5FBFDAE525E8D" box="[1281,1383,1148,1175]" pageId="11" pageNumber="438" value="1982-07-03">3.7.1982</collectingDate>
|
||
</date>
|
||
, leg.
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||
<collectorName id="268A5332EB6DFFE1FF53FB21AA225EA0" box="[151,279,1184,1210]" pageId="11" pageNumber="438">Zwick. The</collectorName>
|
||
<typeStatus id="54C48846EB6DFFE1FED9FB21AA4A5EA0" box="[285,383,1184,1210]" pageId="11" pageNumber="438" type="holotype">holotype</typeStatus>
|
||
will be lodged in the collection of the
|
||
<collectorName id="268A5332EB6DFFE1FCE3FB21AF085EA1" box="[807,1085,1184,1211]" pageId="11" pageNumber="438">Natural History Museum</collectorName>
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<collectingRegion id="49BBF806EB6DFFE1FB87FB21AFA75EA0" box="[1091,1170,1184,1210]" country="Austria" name="Wien" pageId="11" pageNumber="438">Vienna</collectingRegion>
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(
|
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<collectionCode id="ED6EAE21EB6DFFE1FB64FB21AE375EA0" box="[1184,1282,1184,1210]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="438" type="Museum">NHMW</collectionCode>
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), the
|
||
<typeStatus id="54C48846EB6DFFE1FAF8FB20AEA95EA1" box="[1340,1436,1185,1211]" pageId="11" pageNumber="438" type="paratype">paratype</typeStatus>
|
||
remains in the collection of the senior author (
|
||
<collectionCode id="ED6EAE21EB6DFFE1FD58FB45A9FB5EC4" box="[668,718,1220,1246]" pageId="11" pageNumber="438">RW</collectionCode>
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)
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</materialsCitation>
|
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.
|
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C365656FEB6DFFE1FF03FB69AAAF5D04" pageId="11" pageNumber="438" type="description">
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<paragraph id="8BC036E4EB6DFFE1FF03FB69A84D5F50" blockId="11.[151,1437,1004,2038]" pageId="11" pageNumber="438">
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<emphasis id="B90BEAF6EB6DFFE1FF03FB69AA965F18" bold="true" box="[199,419,1256,1282]" pageId="11" pageNumber="438">Description. Male.</emphasis>
|
||
Body brown, head with circular eyes, most setae broken; mouthparts brown. Antenna with short tubular scape as wide as long, pedicel as long as scape but about 1.5 times wider than long; further antennal segments broken. Body length
|
||
<quantity id="4C879B01EB6DFFE1FE37FAB1A97C5F51" box="[499,585,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="438" unit="mm" value="5.8">5.8 mm</quantity>
|
||
(slide mounted specimen).
|
||
</paragraph>
|
||
<paragraph id="8BC036E4EB6DFFE1FF03FAD5AA4A5F89" blockId="11.[151,1437,1004,2038]" pageId="11" pageNumber="438">Thorax broken, 6–7 pairs of dorsocentral setae recognizable on mesonotum. Legs with tarsomere 4 half as long as tarsomeres 3 or 5.</paragraph>
|
||
<paragraph id="8BC036E4EB6DFFE1FF03FA1DAAB05C39" blockId="11.[151,1437,1004,2038]" pageId="11" pageNumber="438">
|
||
Wing with brownish membrane, pterostigma elongate. Sc thinner than R
|
||
<subScript id="17FB34A1EB6DFFE1FC2FFA2FA8C65FA6" attach="left" box="[1003,1011,1454,1468]" fontSize="6" pageId="11" pageNumber="438">1</subScript>
|
||
with short crossvein to C. Brown area between base of R
|
||
<subScript id="17FB34A1EB6DFFE1FEAFFA53AA465FFA" attach="left" box="[363,371,1490,1504]" fontSize="6" pageId="11" pageNumber="438">1</subScript>
|
||
and M
|
||
<subScript id="17FB34A1EB6DFFE1FE02FA53AAEA5FFA" attach="left" box="[454,479,1490,1504]" fontSize="6" pageId="11" pageNumber="438">1+2</subScript>
|
||
to crossvein r-m; further patch at crossvein m-cu and broad band across wing from pterostigma along short imperfect crossvein to M
|
||
<subScript id="17FB34A1EB6DFFE1FD08FA77A9E15C1E" attach="left" box="[716,724,1526,1540]" fontSize="6" pageId="11" pageNumber="438">1</subScript>
|
||
/M
|
||
<subScript id="17FB34A1EB6DFFE1FD31FA77A9C85C1E" attach="left" box="[757,765,1526,1540]" fontSize="6" pageId="11" pageNumber="438">2</subScript>
|
||
fork and to M
|
||
<subScript id="17FB34A1EB6DFFE1FC63FA77A8F55C1E" attach="left" box="[935,960,1526,1540]" fontSize="6" pageId="11" pageNumber="438">1+2</subScript>
|
||
to M
|
||
<subScript id="17FB34A1EB6DFFE1FBC5FA77AF3C5C1E" attach="left" box="[1025,1033,1526,1540]" fontSize="6" pageId="11" pageNumber="438">4</subScript>
|
||
; further small patch on R
|
||
<subScript id="17FB34A1EB6DFFE1FAF7FA77AE0E5C1E" attach="left" box="[1331,1339,1526,1540]" fontSize="6" pageId="11" pageNumber="438">4</subScript>
|
||
/R
|
||
<subScript id="17FB34A1EB6DFFE1FA91FA77AE685C1E" attach="left" box="[1365,1373,1526,1540]" fontSize="6" pageId="11" pageNumber="438">5</subScript>
|
||
fork. Wing length
|
||
<quantity id="4C879B01EB6DFFE1FEECF989AAB45C39" box="[296,385,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="11" pageNumber="438" unit="mm" value="5.2">5.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC036E4EB6DFFE1FF03F9ADAA8A5CE0" blockId="11.[151,1437,1004,2038]" pageId="11" pageNumber="438">
|
||
Abdomen with sternites and tergites brown. Terminalia (
|
||
<figureCitation id="13442A61EB6DFFE1FCFEF9ADA8935C5C" box="[826,934,1580,1606]" captionStart="FIGURE 9" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[270,1271,182,1920]" captionTargetId="figure-22@12.[270,1318,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Phaeobalia species. a, P. pokornyi Mik, slide with lectotype; b, male terminalia, lateral view; c, P. remschakae Wagner sp. nov., slide with holotype; d, male terminalia, lateral view; e, epandrium with clasping cercus, inner view. (photos RW)." figureDoi="http://doi.org/10.5281/zenodo.7471844" httpUri="https://zenodo.org/record/7471844/files/figure.png" pageId="11" pageNumber="438">Fig. 9d, e</figureCitation>
|
||
): with rhomboid hypandrium, corners round, phallus slightly bent forward, apical flagellum approximately 0.4 length of phallus. Epandrium triangular, higher than long, dorsal corner round; surstylus small, not outreaching epandrium edge. Clasping cercus with lateral lobe S-shaped, base to middle broad, increasingly thinner apically, inner surface at base and in middle covered with strong black setae. Inner lobe upright, almost straight, about half as wide as lateral lobe, with long setae along basal edge and on inner surface.
|
||
</paragraph>
|
||
<paragraph id="8BC036E4EB6DFFE1FF03F885AAAF5D04" blockId="11.[151,1437,1004,2038]" box="[199,410,1796,1822]" pageId="11" pageNumber="438">
|
||
<emphasis id="B90BEAF6EB6DFFE1FF03F885AA105D04" bold="true" box="[199,293,1796,1822]" pageId="11" pageNumber="438">Female.</emphasis>
|
||
Unknown
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365656FEB6DFFE1FF03F8A9A8FE5D7C" pageId="11" pageNumber="438" type="etymology">
|
||
<paragraph id="8BC036E4EB6DFFE1FF03F8A9A8FE5D7C" blockId="11.[151,1437,1004,2038]" pageId="11" pageNumber="438">
|
||
<emphasis id="B90BEAF6EB6DFFE1FF03F8A9AA7B5D58" bold="true" box="[199,334,1832,1858]" pageId="11" pageNumber="438">Etymology.</emphasis>
|
||
The species named in honour of Christina Remschak, part of the scientific staff of the Austrian National Park ‘Gesäuse’. She works on the ecology and fauna of springs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365656FEB6DFFE1FF03F8F1AAAD5DEC" pageId="11" pageNumber="438" type="discussion">
|
||
<paragraph id="8BC036E4EB6DFFE1FF03F8F1AAAD5DEC" blockId="11.[151,1437,1004,2038]" pageId="11" pageNumber="438">
|
||
<emphasis id="B90BEAF6EB6DFFE1FF03F8F1AA0E5D90" bold="true" box="[199,315,1904,1930]" pageId="11" pageNumber="438">Remarks.</emphasis>
|
||
The new species belongs to the
|
||
<taxonomicName id="4C7F4D67EB6DFFE1FD65F8F1A8BE5D90" box="[673,907,1904,1930]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="trinotata">
|
||
<emphasis id="B90BEAF6EB6DFFE1FD65F8F1A8BE5D90" box="[673,907,1904,1930]" italics="true" pageId="11" pageNumber="438">Phaeobalia trinotata</emphasis>
|
||
</taxonomicName>
|
||
-group with almost equal sized lateral and inner lobes of the clasping cercus. A curved lateral lobe also occurs in
|
||
<taxonomicName id="4C7F4D67EB6DFFE1FC47F815AFFB5DB4" authority="Vaillant" authorityName="Vaillant" authorityYear="1973" box="[899,1230,1940,1966]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B90BEAF6EB6DFFE1FC47F815AF5A5DB4" box="[899,1135,1940,1966]" italics="true" pageId="11" pageNumber="438">Phaeobalia montana</emphasis>
|
||
Vaillant
|
||
</taxonomicName>
|
||
from the Western Alps, but the shape of the curvature is dissimilar; furthermore, the inner lobe is curved in
|
||
<taxonomicName id="4C7F4D67EB6DFFE1FBB9F838AFCE5DC8" authorityName="Vaillant" authorityYear="1973" box="[1149,1275,1977,2002]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="B90BEAF6EB6DFFE1FBB9F838AFCE5DC8" box="[1149,1275,1977,2002]" italics="true" pageId="11" pageNumber="438">P. montana</emphasis>
|
||
</taxonomicName>
|
||
but straight in
|
||
<taxonomicName id="4C7F4D67EB6DFFE1FF53F85CAA0D5DEC" authority="Wagner & Plant & Ivković & Gattolliat, 2022" authorityName="Wagner & Plant & Ivković & Gattolliat" authorityYear="2022" box="[151,312,2012,2038]" class="Insecta" family="Empididae" genus="Phaeobalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="438" phylum="Arthropoda" rank="species" species="remschakae" status="sp. nov.">
|
||
<emphasis id="B90BEAF6EB6DFFE1FF53F85CAA0D5DEC" box="[151,312,2012,2038]" italics="true" pageId="11" pageNumber="438">P. remschakae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90BEAF6EB6DFFE1FEFAF85DAAAD5DEC" bold="true" box="[318,408,2012,2038]" pageId="11" pageNumber="438">
|
||
<taxonomicNameLabel id="A238578DEB6DFFE1FEFAF85DAAAD5DEC" box="[318,408,2012,2038]" pageId="11" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |