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<document id="041C91B6F32D4B6D00CFF973245ACEC1" ID-CLB-Dataset="46146" ID-DOI="10.5281/zenodo.278958" ID-GBIF-Dataset="56fb15f0-b55b-4ce9-a9ad-d8a2f5a95029" ID-ISSN="1175-5326" ID-Zenodo-Dep="278958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460401525037" checkinUser="plazi" docAuthor="Williams, Kevin A., Brothers, Denis J. &amp; Pitts, James P." docDate="2011" docId="03BF6217D83C215306B6483AFC11FD70" docLanguage="en" docName="zt03064p068.pdf" docOrigin="Zootaxa 3064" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tobantilla kolasma Williams, Brothers &amp; Pitts, sp. nov." docType="treatment" docVersion="11" lastPageNumber="57" masterDocId="FF861A6FD833214306214C5EFF9CFFBC" masterDocTitle="New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae)" masterLastPageNumber="68" masterPageNumber="41" pageNumber="56" updateTime="1698299261469" updateUser="ExternalLinkService">
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<mods:title id="EF4602E3ECD0B58167AA0AAAF8196DCC">New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae)</mods:title>
</mods:titleInfo>
<mods:name id="2C19963004A6414DE99DDDCAEE6ED635" type="personal">
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<mods:namePart id="B10F5CB4F83BC90D28659F3D1ECC323A">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="68D5F32FB7EEF7685FE7CD340D79EA3D">Brothers, Denis J.</mods:namePart>
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<mods:namePart id="AA95379E5008F4FA0305774179F281B8">Pitts, James P.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D4AD3030300418AD1C9FF00A4E3DA506">text</mods:typeOfResource>
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<mods:title id="AB831225B5712A24FAB76D7332A05F45">Zootaxa</mods:title>
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<mods:part id="1D5E827074EF92ADD44EC711FEAE4E75">
<mods:date id="40383A452A48DF403CB2B1ACC2D7CF62">2011</mods:date>
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<treatment id="03BF6217D83C215306B6483AFC11FD70" ID-DOI="http://doi.org/10.5281/zenodo.6191533" ID-GBIF-Taxon="119419954" ID-Zenodo-Dep="6191533" LSID="urn:lsid:zoobank.org:act:74AF9D81-E898-49D9-A032-D8F3ABA0E2A7" httpUri="http://treatment.plazi.org/id/03BF6217D83C215306B6483AFC11FD70" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56">
<subSubSection id="C30C808AD83C214C06B6483AFEDEFB22" pageId="15" pageNumber="56" type="nomenclature">
<paragraph id="8BA9D301D83C214C06B6483AFCA2FBC1" blockId="15.[151,830,1123,1182]" box="[151,830,1123,1150]" pageId="15" pageNumber="56">
<heading id="D0E1646DD83C214C06B6483AFCA2FBC1" bold="true" box="[151,830,1123,1150]" fontSize="11" level="1" pageId="15" pageNumber="56" reason="1">
<emphasis id="B9620F13D83C214C06B6483AFCA2FBC1" bold="true" box="[151,830,1123,1150]" pageId="15" pageNumber="56">
<taxonomicName id="4C16A882D83C214C06B6483AFD48FBC1" authority="Williams, Brothers &amp; Pitts" authorityName="Williams, Brothers &amp; Pitts" box="[151,724,1123,1150]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="kolasma" status="sp. nov.">
<emphasis id="B9620F13D83C214C06B6483AFEE2FBC2" bold="true" box="[151,382,1124,1150]" italics="true" pageId="15" pageNumber="56">Tobantilla kolasma</emphasis>
Williams, Brothers &amp; Pitts
</taxonomicName>
,
<taxonomicNameLabel id="A251B268D83C214C04C1483DFCA2FBC1" box="[736,830,1123,1149]" pageId="15" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA9D301D83C214C06B648D8FEDEFB22" blockId="15.[151,830,1123,1182]" box="[151,322,1158,1182]" pageId="15" pageNumber="56">
(
<figureCitation id="132DCF84D83C214C06BE48D8FF7BFB22" box="[159,231,1158,1182]" captionStart="FIGURES 1 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="15" pageNumber="56">Figs 4</figureCitation>
,
<figureCitation id="132DCF84D83C214C06D248D8FE8DFB22" box="[243,273,1158,1182]" captionStart="FIGURES 7 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="15" pageNumber="56">10</figureCitation>
,
<figureCitation id="132DCF84D83C214C073C48D8FEA6FB22" box="[285,314,1158,1182]" captionStart="FIGURES 19 38" captionStartId="10.[151,264,1672,1695]" captionTargetBox="[176,1419,215,1644]" captionTargetId="figure@10.[157,1429,193,1651]" captionTargetPageId="10" captionText="FIGURES 19 38. Tobantilla species. 19 34. Male genitalia: dorsal view (left), ventral view (middle), lateral view of penis valve (above), and lateral view with penis valve removed (right). 19 22. T. andrikos sp. nov.; 23 26. T. drosos sp. nov.; 27 30. T. ephemeros sp. nov.; 31 34. T. xouthos sp. nov. 35 36. Male clypeus. 28. T. andrikos sp. nov.; 29. T. ephemeros sp. nov. 37, 38. Female pygidium (T 6). 37. T. frigidula (Cresson); 38. T. kolasma sp. nov." httpUri="https://zenodo.org/record/278962/files/figure.png" pageId="15" pageNumber="56">38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD83C214C06B64890FB9BFA92" pageId="15" pageNumber="56" type="diagnosis">
<paragraph id="8BA9D301D83C214C06B64890FB9BFA92" blockId="15.[151,1437,1230,2010]" pageId="15" pageNumber="56">
<emphasis id="B9620F13D83C214C06B64890FE8DFB5B" bold="true" box="[151,273,1230,1255]" pageId="15" pageNumber="56">Diagnosis.</emphasis>
FEMALE. The head and mesosoma are pale brown (
<figureCitation id="132DCF84D83C214C05524890FC25FB5A" box="[883,953,1230,1254]" captionStart="FIGURES 1 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="15" pageNumber="56">Fig. 4</figureCitation>
); the scutellar scale is contiguous with the carinae that separate the dorsal and posterior propodeal faces (
<figureCitation id="132DCF84D83C214C057648ACFC01FAB6" box="[855,925,1266,1290]" captionStart="FIGURES 1 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="15" pageNumber="56">Fig. 4</figureCitation>
); the posterior propodeal face is clothed with dense black setae (
<figureCitation id="132DCF84D83C214C07484948FE20FA92" box="[361,444,1302,1326]" captionStart="FIGURES 7 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="15" pageNumber="56">Fig. 10</figureCitation>
); and the pygidial plate is defined by lateral carinae.
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD83C215306E44964FE04FE3B" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56" type="description">
<paragraph id="8BA9D301D83C215306E44964FC6FFED8" blockId="15.[151,1437,1230,2010]" lastBlockId="16.[151,1436,151,716]" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56">
<emphasis id="B9620F13D83C214C06E44964FECAFAEE" bold="true" box="[197,342,1338,1363]" pageId="15" pageNumber="56">Description.</emphasis>
FEMALE. Body length
<quantity id="4CEE7EE4D83C214C044A4964FD69FAEE" box="[619,757,1338,1362]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="15" pageNumber="56" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
.
<emphasis id="B9620F13D83C214C04DE4965FC1DFAEF" box="[767,897,1339,1363]" italics="true" pageId="15" pageNumber="56">Coloration.</emphasis>
Head, mesosoma, legs, antennae and mandibles pale brown. Metasoma brown except T24 dark brown to black; T2 having two pairs of yellow circular spots, anterior pair of spots longitudinally ovate, slightly larger than posterior circular pair. Tibial spurs white. Head and mesosoma clothed with pale golden or silvery setae, except medial portion of mesosoma having interspersed black and golden setae, area directly anterior to scutellar scale clothed with black setae, posterior face of propodeum densely clothed with thick black setae, and dorsum of head and mesosoma having a few scattered erect brown setae. T23 clothed with black setae, except lateral margins and integumental spots clothed with silver setae. T56 and S16 clothed with silvery white setae.
<emphasis id="B9620F13D83C214C04A04A68FD21F9F2" box="[641,701,1590,1614]" italics="true" pageId="15" pageNumber="56">Head</emphasis>
. Rounded posteriorly, with lowly produced rectangular glabrous tubercles on posterolateral margin. Head width 1.1 × pronotal width. Eye almost circular. Front, vertex and gena reticulate. Genal carina lowly produced, extending anteriorly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe with weak dorsal carina. Antennal tubercle moderately punctate basally. Scape simply punctate. Flagellomere 1 1.5 × pedicel length; flagellomere 2 1.3 × pedicel length.
<emphasis id="B9620F13D83C214C06B64AB2FE93F8BF" box="[151,271,1772,1795]" italics="true" pageId="15" pageNumber="56">Mesosoma</emphasis>
. Mesosomal length 1.1 × width; pronotum slightly wider than mesothorax. Mesosomal dorsum reticulate. Humeral carina low, extending to epaulet. Lateral face of pronotum and mesopleuron micropunctate, dull, mesopleural ridge broadly and shallowly reticulate; clothed with sparse silver setae. Metapleuron dorsal to endophragmal pit and dorsal two-thirds of lateral face of propodeum shagreened; ventral regions of metapleuron and lateral face of propodeum shagreened. In dorsal view, mesosoma gradually narrowed posterior to pronotum, lateral margin of mesothorax weakly sinuate anterior to propodeal spiracle. Scutellar scale broad and highly raised, contiguous with carina separating dorsal and posterior propodeal faces, lacking transverse carinae anteriorly. Dorsal propodeal face weakly convergent posterior to spiracle, posterolateral angle rounded, posterior face virtually flat, vertical, reticulate, and clothed with dense dark brown setae.
<emphasis id="B9620F13D823215305644CE3FC26FF68" box="[837,954,189,212]" italics="true" pageId="16" pageNumber="57">Metasoma</emphasis>
. T1 narrow and petiolate, somewhat cylindrical, 0.4 × as wide as T2. T2 1.1 × longer than wide, with maximum width situated medially. Disc of T2 with small longitudinally ovate contiguous punctures, except yellow spots sparsely punctate; S2 sparsely punctate. T35 densely punctate, S25 moderately punctate. Pygidium well defined by lateral carinae, posterior margin truncate, having 8 widely separated striae, spaces between striae irregularly granulate.
</paragraph>
<paragraph id="8BA9D301D823215306E44D2EFE04FE3B" blockId="16.[151,1436,151,716]" box="[197,408,367,392]" pageId="16" pageNumber="57">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306E44DCAFB29FE73" pageId="16" pageNumber="57" type="materials_examined">
<paragraph id="8BA9D301D823215306E44DCAFB29FE73" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44DCAFEE8FE17" bold="true" box="[197,372,403,428]" pageId="16" pageNumber="57">
<typeStatus id="54AD6DA3D823215306E44DCAFE9CFE10" box="[197,256,404,428]" pageId="16" pageNumber="57">Type</typeStatus>
material.
</emphasis>
<emphasis id="B9620F13D8232153075C4DCAFE7FFE10" box="[381,483,404,428]" italics="true" pageId="16" pageNumber="57">
<typeStatus id="54AD6DA3D8232153075C4DCAFE7FFE10" box="[381,483,404,428]" pageId="16" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
, Ƥ,
<collectingCountry id="F3019391D823215304394DCAFD5FFE10" box="[536,707,404,428]" name="Venezuela" pageId="16" pageNumber="57">VENEZUELA</collectingCountry>
: Guarico, Hato Masaguaral (
<quantity id="4CEE7EE4D8232153022D4DCAFBCAFE10" box="[1036,1110,404,428]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="57" unit="km" value="44.0">44 km</quantity>
S Calabozo),
<date id="FFA8F5C1D823215302D84DCAFA0BFE10" box="[1273,1431,404,428]" pageId="16" pageNumber="57" value="1985-05-11" valueMax="1985-05-19" valueMin="1985-05-11">1119.V.1985</date>
, A.S. Menke &amp; J. Carpenter (
<collectionCode id="ED074BC4D823215307F64DE9FDB2FE73" LSID="urn:lsid:biocol.org:col:34871" box="[471,558,439,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="57">USNM</collectionCode>
).
<emphasis id="B9620F13D823215304604DE7FD2EFE6C" box="[577,690,441,464]" italics="true" pageId="16" pageNumber="57">
<typeStatus id="54AD6DA3D823215304604DE7FD2EFE6C" box="[577,690,441,464]" pageId="16" pageNumber="57" type="paratype">Paratypes</typeStatus>
</emphasis>
. 2Ƥ, same data as
<typeStatus id="54AD6DA3D823215305A74DE9FC74FE73" box="[902,1000,439,463]" pageId="16" pageNumber="57" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED074BC4D823215305D74DE9FBD7FE73" LSID="urn:lsid:biocol.org:col:34871" box="[1014,1099,439,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="57">USNM</collectionCode>
,
<collectionCode id="ED074BC4D823215302764DE9FB36FE73" box="[1111,1194,439,463]" pageId="16" pageNumber="57">EMUS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306E44D85FE1DFDAB" pageId="16" pageNumber="57" type="distribution">
<paragraph id="8BA9D301D823215306E44D85FC0FFE48" blockId="16.[151,1436,151,716]" box="[197,915,475,501]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44D85FEC2FE4F" bold="true" box="[197,350,475,500]" pageId="16" pageNumber="57">Distribution.</emphasis>
This species is known only from the
<typeStatus id="54AD6DA3D823215305234D83FCAEFE49" box="[770,818,477,501]" pageId="16" pageNumber="57">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BA9D301D823215306E44E5EFE1DFDAB" blockId="16.[151,1436,151,716]" box="[197,385,511,536]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44E5EFE98FDA4" bold="true" box="[197,260,512,536]" pageId="16" pageNumber="57">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306E44E7AFE23FD38" pageId="16" pageNumber="57" type="etymology">
<paragraph id="8BA9D301D823215306E44E7AFE23FD38" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44E7AFED7FD80" bold="true" box="[197,331,548,572]" pageId="16" pageNumber="57">Etymology.</emphasis>
From the Greek
<taxonomicName id="4C16A882D8232153042D4E7AFDF5FD80" box="[524,617,548,572]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="kolasma">
<emphasis id="B9620F13D8232153042D4E7AFDF5FD80" box="[524,617,548,572]" italics="true" pageId="16" pageNumber="57">kolasma</emphasis>
</taxonomicName>
“punishment”, a noun in apposition. This species, along with
<taxonomicName id="4C16A882D823215303044E7BFA1CFD80" box="[1317,1408,548,572]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="krima">
<emphasis id="B9620F13D823215303044E7BFA1CFD80" box="[1317,1408,548,572]" italics="true" pageId="16" pageNumber="57">T. krima</emphasis>
</taxonomicName>
is named after the Russian novel Преступление и наказание “Crime and Punishment” written by Fyodor Mikhailovich Dostoevsky.
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306E44ECEFC11FD70" pageId="16" pageNumber="57" type="discussion">
<paragraph id="8BA9D301D823215306E44ECEFC11FD70" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44ECEFEA5FD1B" bold="true" box="[197,313,656,680]" pageId="16" pageNumber="57">Remarks.</emphasis>
The scutellar scale of this species is similar to that of
<taxonomicName id="4C16A882D823215305804ECFFB83FD14" box="[929,1055,656,680]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="frigidula">
<emphasis id="B9620F13D823215305804ECFFB83FD14" box="[929,1055,656,680]" italics="true" pageId="16" pageNumber="57">T. frigidula</emphasis>
</taxonomicName>
. This species also has a narrower pygidial plate than
<taxonomicName id="4C16A882D8232153074F4EEBFE56FD70" box="[366,458,692,716]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="krima">
<emphasis id="B9620F13D8232153074F4EEBFE56FD70" box="[366,458,692,716]" italics="true" pageId="16" pageNumber="57">T. krima</emphasis>
</taxonomicName>
and the Argentinean
<taxonomicName id="4C16A882D8232153049D4EEAFCB2FD70" box="[700,814,692,716]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B9620F13D8232153049D4EEAFCB2FD70" box="[700,814,692,716]" italics="true" pageId="16" pageNumber="57">Tobantilla</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>