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<document id="0FF9ACBBB761904817342A5F706553BB" ID-CLB-Dataset="225933" ID-DOI="10.11646/zootaxa.5228.4.3" ID-GBIF-Dataset="bb50dc3b-5fb9-4739-9573-84f8e625c342" ID-ISSN="1175-5326" ID-Zenodo-Dep="7539938" ID-ZooBank="F6AD0451-C428-43CD-A7CD-FFA10D06577A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673854054836" checkinUser="plazi" docAuthor="Le, Son X., Schileyko, Arkady A. &amp; Nguyen, Anh D." docDate="2023" docId="03914E40E94FFF81FF3DBC07FE7CF8A9" docLanguage="en" docName="zootaxa.5228.4.3.pdf" docOrigin="Zootaxa 5228 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scolopocryptops Newport 1844" docType="key" docVersion="5" lastPageNumber="437" masterDocId="FFA83638E955FF9BFFAABF19FFE4FFAF" masterDocTitle="A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species" masterLastPageNumber="447" masterPageNumber="441" pageNumber="437" updateTime="1699473965273" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1EC066932F66686E70703019920CF62E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="93873CFC4DE7E03356D518482ACC4EE8">
<mods:title id="093E7B91919ADA92584D870786A050F9">A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species</mods:title>
</mods:titleInfo>
<mods:name id="93595856F9EDFA0038D25579C1783265" type="personal">
<mods:role id="18B0BE1E79BED3B2FD1031695D89C48C">
<mods:roleTerm id="971C5851D4F8FF5F6C46AB7CBB359F36">Author</mods:roleTerm>
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<mods:namePart id="1D85C7B2E19405ABB15C3B2A2242D91A">Le, Son X.</mods:namePart>
<mods:nameIdentifier id="80608109E4E68AB11D47ACF69054D7E5" type="ORCID">0000-0003-1648-6773</mods:nameIdentifier>
<mods:affiliation id="8AEF57B64E3695396F1DC49B4CE3390D">Institute of Tropical Ecology, Vietnam-Russia Tropical Centre, 63 Nguyen Van Huyen Rd, Cau Giay District, Hanoi, Vietnam. lesonenv 86 @ gmail. com; https: // orcid. org / 0000 - 0003 - 1648 - 6773 &amp; VNU University of Sciences, Vietnam National University, 334 Nguyen Trai, Thanh Xuan District, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="B5B71F9D5FFE7EA5707F9A375709F3F8" type="email">lesonenv86@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="38F422FB7165EAA527385E9F1DF49D3F" type="personal">
<mods:role id="A80A61F451BD2134572DC05A98DD221F">
<mods:roleTerm id="2135B30C001F486892162000BFB86A59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8304E4CE597247249C811FF69A1A0E12">Schileyko, Arkady A.</mods:namePart>
<mods:nameIdentifier id="074AF24A97778CBB821CE0C195FAB475" type="ORCID">0000-0002-6139-5240</mods:nameIdentifier>
<mods:affiliation id="19A735367B9994F64D6168679C43BF2A">Zoological Museum of the Moscow Lomonosov State University, Bolshaya Nikitskaja Str. 2, Moscow, 125009, Russia. schileyko 1965 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6139 - 5240</mods:affiliation>
<mods:nameIdentifier id="3DC941E1C8B1C329F8FD489A656B9DE6" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="6005C23A3EF21A45DB9994ABE48584A2" type="personal">
<mods:role id="F8F33695A5A36B0480F69D773B72463E">
<mods:roleTerm id="031DFC038E6ECCA49D5FB6D08F37C49C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFFF5959665C97DB8AF17F153F93BE83">Nguyen, Anh D.</mods:namePart>
<mods:nameIdentifier id="EFD89D3A18CEA6E58A4ABEFA795BC2C9" type="ORCID">0000-0001-9273-0040</mods:nameIdentifier>
<mods:affiliation id="CE0E88C5010650DED285FCB0D747B799">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Cau Giay District, Hanoi, Vietnam. ducanh. iebr @ gmail. com; https: // orcid. org / 0000 - 0001 - 9273 - 0040 &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Cau Giay District, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="DA2E5BC1271071D656A64AF8C91CB3D4" type="email">ducanh.iebr@gmail.com</mods:nameIdentifier>
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<mods:title id="4EB21022443EC1B68F198BAAB4396B42">Zootaxa</mods:title>
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<mods:part id="89FF02DD67C27D3E0C027394A84C4F22">
<mods:date id="FA5EBB1124CF555386C368CBFCA2942C">2023</mods:date>
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<mods:number id="85E2723C28BCFAAAD06645D47EE984B5">2023-01-16</mods:number>
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<mods:number id="9E917D1BFF3F4C99D77B8916825783D6">5228</mods:number>
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<mods:identifier id="71E3A02DC7C1CF098C587610672C1D40" type="DOI">10.11646/zootaxa.5228.4.3</mods:identifier>
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<treatment id="03914E40E94FFF81FF3DBC07FE7CF8A9" ID-DOI="http://doi.org/10.5281/zenodo.7541473" ID-GBIF-Taxon="208976655" ID-Zenodo-Dep="7541473" LSID="urn:lsid:plazi:treatment:03914E40E94FFF81FF3DBC07FE7CF8A9" httpUri="http://treatment.plazi.org/id/03914E40E94FFF81FF3DBC07FE7CF8A9" lastPageNumber="437" pageId="26" pageNumber="437">
<subSubSection id="C322ACDDE94FFF81FF3DBC07FC67FC97" box="[151,899,798,825]" pageId="26" pageNumber="437" type="nomenclature">
<paragraph id="8B87FF56E94FFF81FF3DBC07FC67FC97" blockId="26.[151,899,798,825]" box="[151,899,798,825]" pageId="26" pageNumber="437">
<heading id="D0CF483AE94FFF81FF3DBC07FC67FC97" bold="true" box="[151,899,798,825]" fontSize="11" level="1" pageId="26" pageNumber="437" reason="1">
<emphasis id="B94C2344E94FFF81FF3DBC07FC67FC97" bold="true" box="[151,899,798,825]" pageId="26" pageNumber="437">
An identification key to
<taxonomicName id="4C3884D5E94FFF81FE6FBC07FD6EFC96" ID-CoL="63T5J" ID-ENA="118498" authority="Newport, 1844" authorityName="Newport" authorityYear="1844" box="[453,650,798,825]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B94C2344E94FFF81FE6FBC07FD6EFC96" bold="true" box="[453,650,798,825]" italics="true" pageId="26" pageNumber="437">Scolopocryptops</emphasis>
</taxonomicName>
species of
<collectingCountry id="F32FBFC6E94FFF81FCA5BC06FC67FC97" box="[783,899,798,825]" name="China" pageId="26" pageNumber="437">East Asia</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322ACDDE94FFF81FF3DBC7DFE7CF8A9" pageId="26" pageNumber="437" type="key">
<key id="FFA9A867E94FFF81FF3DBC7DFE7CF8A9" pageId="26" pageNumber="437">
<keyStep id="30CCE7F3E94FFF81FF3DBC7DFA78FC39" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBC7DFA79FCD6" blockId="26.[151,1437,867,1799]" box="[151,1437,867,890]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBC7DFA79FCD6" box="[151,1437,867,890]" pageId="26" pageNumber="437">
1. LBS 7 with well-developed spiracles (
<figureCitation id="1303E3D3E94FFF81FDFBBC7DFD49FCD6" box="[593,685,868,890]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="26" pageNumber="437">Figs 14D</figureCitation>
,
<figureCitation id="1303E3D3E94FFF81FD1DBC7AFD05FCD6" box="[695,737,867,889]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="26" pageNumber="437">15E</figureCitation>
)....................................................... 2.
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBC98FA78FC39" blockId="26.[151,1437,867,1799]" box="[151,1436,897,919]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBC98FA78FC39" box="[151,1436,897,919]" pageId="26" pageNumber="437">- LBS 7 lacking spiracles............................................................................... 3.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBC87FA78FC7D" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBC87FA78FC1B" blockId="26.[151,1437,867,1799]" box="[151,1436,926,948]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBC87FA78FC1B" box="[151,1436,926,948]" pageId="26" pageNumber="437">
2. Ultimate tergite with median suture......................................
<taxonomicName id="4C3884D5E94FFF81FBA5BC86FAF9FC1C" box="[1039,1309,926,948]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="broelemanni">
<emphasis id="B94C2344E94FFF81FBA5BC86FAF9FC1C" box="[1039,1309,926,948]" italics="true" pageId="26" pageNumber="437">S. broelemanni broelemanni</emphasis>
</taxonomicName>
(East
<collectingCountry id="F32FBFC6E94FFF81FAF1BC87FA72FC1B" box="[1371,1430,926,948]" name="China" pageId="26" pageNumber="437">China</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBCA5FA78FC7D" blockId="26.[151,1437,867,1799]" box="[151,1436,956,978]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBCA5FA78FC7D" box="[151,1436,956,978]" pageId="26" pageNumber="437">
- Ultimate tergite lacking median suture.....................................
<taxonomicName id="4C3884D5E94FFF81FBBBBCA5FB13FC7E" authorityName="Chagas" authorityYear="2008" box="[1041,1271,956,977]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
<emphasis id="B94C2344E94FFF81FBBBBCA5FB13FC7E" box="[1041,1271,956,977]" italics="true" pageId="26" pageNumber="437">S. broelemanni esulcata</emphasis>
</taxonomicName>
(South
<collectingCountry id="F32FBFC6E94FFF81FAE9BCA5FA72FC7D" box="[1347,1430,956,978]" name="Vietnam" pageId="26" pageNumber="437">Vietnam</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBCC0FA79FBA4" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBCC0FA79FC41" blockId="26.[151,1437,867,1799]" box="[151,1437,985,1007]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBCC0FA79FC41" box="[151,1437,985,1007]" pageId="26" pageNumber="437">3. Cephalic plate with complete lateral margination........................................................... 7.</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBCEEFA79FBA4" blockId="26.[151,1437,867,1799]" box="[151,1437,1014,1036]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBCEEFA79FBA4" box="[151,1437,1014,1036]" pageId="26" pageNumber="437">- Cephalic plate lacks lateral margination or this margination is much shortened.................................... 4.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBB0DFA78FBE8" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBB0DFA78FB85" blockId="26.[151,1437,867,1799]" box="[151,1436,1044,1066]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBB0DFA78FB85" box="[151,1436,1044,1066]" pageId="26" pageNumber="437">
4. Length of the ultimate leg up to 40% of body length....................................
<taxonomicName id="4C3884D5E94FFF81FB38BB0DFAEBFB86" authorityName="Qiao, Xiao &amp; Di" authorityYear="2021" box="[1170,1295,1044,1065]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="zhijinensis">
<emphasis id="B94C2344E94FFF81FB38BB0DFAEBFB86" box="[1170,1295,1044,1065]" italics="true" pageId="26" pageNumber="437">S. zhijinensis</emphasis>
</taxonomicName>
(South
<collectingCountry id="F32FBFC6E94FFF81FAF1BB0DFA72FB85" box="[1371,1430,1044,1066]" name="China" pageId="26" pageNumber="437">China</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBB28FA78FBE8" blockId="26.[151,1437,867,1799]" box="[151,1436,1073,1095]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBB28FA78FBE8" box="[151,1436,1073,1095]" pageId="26" pageNumber="437">- Length of the ultimate leg less than a 30% of body length.................................................... 5.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBB57FA79FB30" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBB57FC6AFB2D" blockId="26.[151,1437,867,1799]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBB57FC6AFB2D" pageId="26" pageNumber="437">
5. Ultimate tergite lacks lateral margination......................................................
<taxonomicName id="4C3884D5E94FFF81FAAEBB56FA78FBCC" authorityName=": Qiao, Xiao &amp; Di" authorityYear="2021" box="[1284,1436,1102,1124]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="melanostoma">
<emphasis id="B94C2344E94FFF81FAAEBB56FA78FBCC" box="[1284,1436,1102,1124]" italics="true" pageId="26" pageNumber="437">S. melanostoma</emphasis>
</taxonomicName>
(
<collectingCountry id="F32FBFC6E94FFF81FF42BB75FED6FB2D" box="[232,306,1132,1154]" name="Taiwan" pageId="26" pageNumber="437">Taiwan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FE96BB75FE4FFB2D" box="[316,427,1132,1154]" name="Philippines" pageId="26" pageNumber="437">Philippines</collectingCountry>
, E
<collectingCountry id="F32FBFC6E94FFF81FE63BB75FDCDFB2D" box="[457,553,1132,1154]" name="Indonesia" pageId="26" pageNumber="437">Indonesia</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FD98BB75FD0AFB2D" box="[562,750,1132,1154]" name="Papua New Guinea" pageId="26" pageNumber="437">Papua New Guinea</collectingCountry>
, South
<collectingCountry id="F32FBFC6E94FFF81FC9FBB75FC6CFB2D" box="[821,904,1132,1154]" name="Vietnam" pageId="26" pageNumber="437">Vietnam</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBB90FA79FB30" blockId="26.[151,1437,867,1799]" box="[151,1437,1161,1183]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBB90FA79FB30" box="[151,1437,1161,1183]" pageId="26" pageNumber="437">- Ultimate tergite with definite lateral margination........................................................... 6.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBBBFFA79FB59" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBBBFFA78FB13" blockId="26.[151,1437,867,1799]" box="[151,1436,1190,1212]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBBBFFA78FB13" box="[151,1436,1190,1212]" pageId="26" pageNumber="437">
6. Cephalic plate lacks lateral margination; 3 basal antennal articles with sparse minute setae.......
<taxonomicName id="4C3884D5E94FFF81FB00BBBEFB1AFB13" ID-CoL="4VSML" baseAuthorityName="Takakuwa" baseAuthorityYear="1939" box="[1194,1278,1191,1212]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="B94C2344E94FFF81FB00BBBEFB1AFB13" box="[1194,1278,1191,1212]" italics="true" pageId="26" pageNumber="437">S. curtus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32FBFC6E94FFF81FAA0BBBFFAA0FB14" box="[1290,1348,1190,1211]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
, Taiwan)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBBDDFA79FB59" blockId="26.[151,1437,867,1799]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBBDDFA79FB59" pageId="26" pageNumber="437">
- Cephalic plate with very short, but definite tracks of lateral margination in posterior half; 1 basal antennal article with sparse minute setae...........................................................................
<taxonomicName id="4C3884D5E94FFF81FB44BBF8FA79FB59" authority="(Japan)" authorityName="Takakuwa" authorityYear="1937" baseAuthorityName="Japan" box="[1262,1437,1249,1270]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B94C2344E94FFF81FB44BBF8FAABFB59" box="[1262,1359,1249,1270]" italics="true" pageId="26" pageNumber="437">S. elegans</emphasis>
(
<collectingCountry id="F32FBFC6E94FFF81FAF6BBF8FA71FB59" box="[1372,1429,1249,1270]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBBE6FA78FAE1" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBBE6FA79FA9D" blockId="26.[151,1437,867,1799]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBBE6FA79FA9D" pageId="26" pageNumber="437">
7. Length of the ultimate leg more than 40% of body length; tarsus 1 and 2 of leg 22 each with well-developed spur..............................................................................................
<taxonomicName id="4C3884D5E94FFF81FB4EBA05FA79FA9D" authority="(China)" authorityName="Xiao, Chen &amp; Di" authorityYear="2021" baseAuthorityName="China" box="[1252,1437,1308,1330]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B94C2344E94FFF81FB4EBA05FAA9FA9E" box="[1252,1357,1308,1329]" italics="true" pageId="26" pageNumber="437">S. longipes</emphasis>
(
<collectingCountry id="F32FBFC6E94FFF81FAF3BA05FA71FA9D" box="[1369,1429,1308,1330]" name="China" pageId="26" pageNumber="437">China</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBA20FA78FAE1" blockId="26.[151,1437,867,1799]" box="[151,1436,1337,1359]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBA20FA78FAE1" box="[151,1436,1337,1359]" pageId="26" pageNumber="437">- Length of the ultimate leg less than a 30% of body length; leg 22 with 1 tarsal spur at most.......................... 8.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBA4FFA79FA26" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBA4FFA79FAC4" blockId="26.[151,1437,867,1799]" box="[151,1437,1366,1388]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBA4FFA79FAC4" box="[151,1437,1366,1388]" pageId="26" pageNumber="437">
8. Tergites 8–18(20) with complete paramedian sutures (
<figureCitation id="1303E3D3E94FFF81FD42BA4FFCD0FAC4" box="[744,820,1366,1388]" captionStart="FIGURES 2" captionStartId="5.[152,270,1727,1751]" captionTargetBox="[161,1427,185,1702]" captionTargetId="figure-18@5.[159,1429,181,1703]" captionTargetPageId="5" captionText="FIGURES 2A–F. Scolopocryptops rubiginosus C.L. Koch, 1878; SVR.PO.020 A head and LBS 1, dorsally; B head and forcipular segment, ventrally; C LBS 6–10, dorsally; D LBS 6–10, ventrally; Rc 6738 (ZMMU) E head and forcipular segment, ventrally; Rc 6686 (ZMMU) F LBS 7, 8 (right side), laterally." figureDoi="http://doi.org/10.5281/zenodo.7539942" httpUri="https://zenodo.org/record/7539942/files/figure.png" pageId="26" pageNumber="437">Fig. 2C</figureCitation>
)................................................ 9.
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBA6DFA79FA26" blockId="26.[151,1437,867,1799]" box="[151,1437,1395,1418]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBA6DFA79FA26" box="[151,1437,1395,1418]" pageId="26" pageNumber="437">
- Tergites without complete paramedian sutures (
<figureCitation id="1303E3D3E94FFF81FD02BA6DFD00FA26" box="[680,740,1395,1418]" captionStart="FIGURES 5" captionStartId="10.[152,270,1856,1880]" captionTargetBox="[152,1433,188,1831]" captionTargetId="figure-33@10.[151,1436,181,1832]" captionTargetPageId="10" captionText="FIGURES 5A–F. Scolopocryptops spinicaudus Wood, 1862; SVR.PO.202 A head + LBS 1–3, dorsally; B head, forcipular segment and LBS 1–3, ventrally; C LBS 8–12, ventrally; D LBS 8–12, dorsally; Rc 7165 (ZMMU) E articles 1–6 of right antenna, laterally; Rc 6614 (ZMMU) F LBS 16–23, dorso-laterally." figureDoi="http://doi.org/10.5281/zenodo.7539950" httpUri="https://zenodo.org/record/7539950/files/figure.png" pageId="26" pageNumber="437">Figs 5</figureCitation>
DF, 8C)................................................ 10.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBA88FA78FA6C" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBA88FA78FA08" blockId="26.[151,1437,867,1799]" box="[151,1436,1425,1447]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBA88FA78FA08" box="[151,1436,1425,1447]" pageId="26" pageNumber="437">
9. Tergites colored uniformly...................................
<taxonomicName id="4C3884D5E94FFF81FC21BA88FBF0FA09" ID-CoL="4VSN8" authorityName="C. L. Koch" authorityYear="1878" box="[907,1044,1425,1446]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="rubiginosus">
<emphasis id="B94C2344E94FFF81FC21BA88FBF0FA09" box="[907,1044,1425,1446]" italics="true" pageId="26" pageNumber="437">S. rubiginosus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32FBFC6E94FFF81FB8BBA88FB8FFA08" box="[1057,1131,1425,1447]" name="Taiwan" pageId="26" pageNumber="437">Taiwan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FBDFBA88FB55FA08" box="[1141,1201,1425,1447]" name="China" pageId="26" pageNumber="437">China</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FB10BA88FB13FA09" box="[1210,1271,1425,1446]" name="South Korea" pageId="26" pageNumber="437">Korea</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FAABBA88FADEFA09" box="[1281,1338,1425,1446]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FAE9BA88FA72FA08" box="[1347,1430,1425,1447]" name="Vietnam" pageId="26" pageNumber="437">Vietnam</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DBAB6FA78FA6C" blockId="26.[151,1437,867,1799]" box="[151,1436,1454,1476]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBAB6FA78FA6C" box="[151,1436,1454,1476]" pageId="26" pageNumber="437">
- Tergites with some accumulations of dark pigment............................
<taxonomicName id="4C3884D5E94FFF81FBB5BAB6FB2CFA6C" ID-CoL="4VSN2" authorityName="Song, Song &amp; Zhou" authorityYear="2004" box="[1055,1224,1454,1476]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="nigrimaculatus">
<emphasis id="B94C2344E94FFF81FBB5BAB6FB2CFA6C" box="[1055,1224,1454,1476]" italics="true" pageId="26" pageNumber="437">S. nigrimaculatus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32FBFC6E94FFF81FB7FBAB7FAF6FA6B" box="[1237,1298,1454,1476]" name="China" pageId="26" pageNumber="437">China</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FAB6BAB7FA72FA6C" box="[1308,1430,1454,1476]" name="South Korea" pageId="26" pageNumber="437">South Korea</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DBAD5FA79F9B4" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DBAD5FA78FA50" blockId="26.[151,1437,867,1799]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DBAD5FA78FA50" pageId="26" pageNumber="437">
10. Tergites 6–20 with very characteristic deep “drop-like” longitudinal median depression bordered by paramedian keels (Figs 112CD, 12B).................................................................
<taxonomicName id="4C3884D5E94FFF81FBD0BAF0FB1DFA51" authority="Le &amp; Schileyko &amp; Nguyen, 2023" authorityName="Le &amp; Schileyko &amp; Nguyen" authorityYear="2023" box="[1146,1273,1513,1534]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="hoanglieni" status="sp. nov.">
<emphasis id="B94C2344E94FFF81FBD0BAF0FB1DFA51" box="[1146,1273,1513,1534]" italics="true" pageId="26" pageNumber="437">S. hoanglieni</emphasis>
</taxonomicName>
<emphasis id="B94C2344E94FFF81FB55BAF0FAD1FA51" bold="true" box="[1279,1333,1513,1534]" pageId="26" pageNumber="437">
<taxonomicNameLabel id="A27F9E3FE94FFF81FB55BAF0FAD1FA51" box="[1279,1333,1513,1534]" pageId="26" pageNumber="437" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F32FBFC6E94FFF81FAEBBAF0FA71FA50" box="[1345,1429,1513,1535]" name="Vietnam" pageId="26" pageNumber="437">Vietnam</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DB91EFA79F9B4" blockId="26.[151,1437,867,1799]" box="[151,1437,1542,1564]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DB91EFA79F9B4" box="[151,1437,1542,1564]" pageId="26" pageNumber="437">
- Tergites lacking deep “drop-like” longitudinal depression (
<figureCitation id="1303E3D3E94FFF81FCBFB91FFCB5F9B3" box="[789,849,1542,1564]" captionStart="FIGURES 5" captionStartId="10.[152,270,1856,1880]" captionTargetBox="[152,1433,188,1831]" captionTargetId="figure-33@10.[151,1436,181,1832]" captionTargetPageId="10" captionText="FIGURES 5A–F. Scolopocryptops spinicaudus Wood, 1862; SVR.PO.202 A head + LBS 1–3, dorsally; B head, forcipular segment and LBS 1–3, ventrally; C LBS 8–12, ventrally; D LBS 8–12, dorsally; Rc 7165 (ZMMU) E articles 1–6 of right antenna, laterally; Rc 6614 (ZMMU) F LBS 16–23, dorso-laterally." figureDoi="http://doi.org/10.5281/zenodo.7539950" httpUri="https://zenodo.org/record/7539950/files/figure.png" pageId="26" pageNumber="437">Figs 5</figureCitation>
DF, 8C)....................................... 11.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DB93DFA79F9F9" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DB93DFA79F996" blockId="26.[151,1437,867,1799]" box="[151,1437,1572,1594]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DB93DFA79F996" box="[151,1437,1572,1594]" pageId="26" pageNumber="437">
11. Tergites 6–20 with four longitudinal keels...............................................
<taxonomicName id="4C3884D5E94FFF81FB1AB93DFA79F996" authority="(Japan)" baseAuthorityName="Japan" baseAuthorityYear="1934" box="[1200,1437,1572,1593]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="B94C2344E94FFF81FB1AB93DFAB4F996" box="[1200,1360,1572,1593]" italics="true" pageId="26" pageNumber="437">S. quadristriatus</emphasis>
(
<collectingCountry id="F32FBFC6E94FFF81FAF6B93DFA71F996" box="[1372,1429,1572,1593]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DB958FA79F9F9" blockId="26.[151,1437,867,1799]" box="[151,1437,1601,1623]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DB958FA79F9F9" box="[151,1437,1601,1623]" pageId="26" pageNumber="437">- Tergites lacking longitudinal keels...................................................................... 12.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CCE7F3E94FFF81FF3DB947FE7CF8A9" pageId="26" pageNumber="437">
<paragraph id="8B87FF56E94FFF81FF3DB947FCBCF901" blockId="26.[151,1437,867,1799]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DB947FCBCF901" pageId="26" pageNumber="437">
12. Two basal antennal articles sparsely setose dorsally. Ultimate sternite tongue-shaped. Tibia and tarsus of ultimate leg (
<figureCitation id="1303E3D3E94FFF81FAE0B947FA71F9DC" box="[1354,1429,1630,1652]" captionStart="FIGURES 9" captionStartId="15.[152,270,1675,1699]" captionTargetBox="[169,1435,211,1651]" captionTargetId="figure-18@15.[151,1435,181,1651]" captionTargetPageId="15" captionText="FIGURES 9A–D. Scolopocryptops capillipedatus (Takakuwa, 1938); SVR.PH.048 A LBS 21–23, dorsally; B LBS 21–23, laterally; C LBS 20–23, ventrally; D left ultimate leg, medially." figureDoi="http://doi.org/10.5281/zenodo.7539960" httpUri="https://zenodo.org/record/7539960/files/figure.png" pageId="26" pageNumber="437">Fig. 9D</figureCitation>
) very densely setose.............................................
<taxonomicName id="4C3884D5E94FFF81FC10B965FBBEF93E" baseAuthorityName="Takakuwa" baseAuthorityYear="1938" box="[954,1114,1660,1681]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="capillipedatus">
<emphasis id="B94C2344E94FFF81FC10B965FBBEF93E" box="[954,1114,1660,1681]" italics="true" pageId="26" pageNumber="437">S. capillipedatus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32FBFC6E94FFF81FBCDB965FB41F93E" box="[1127,1189,1660,1681]" name="South Korea" pageId="26" pageNumber="437">Korea</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FB05B965FB1CF93D" box="[1199,1272,1660,1682]" name="Taiwan" pageId="26" pageNumber="437">Taiwan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FAABB965FADEF93E" box="[1281,1338,1660,1681]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FAE9B965FA72F93D" box="[1347,1430,1660,1682]" name="Vietnam" pageId="26" pageNumber="437">Vietnam</collectingCountry>
) + “
<taxonomicName id="4C3884D5E94FFF81FEAAB980FEBBF900" authorityName="Shinohara" authorityYear="1984" box="[256,351,1689,1711]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="ogawai">
<emphasis id="B94C2344E94FFF81FEAAB980FEBBF900" box="[256,351,1689,1711]" italics="true" pageId="26" pageNumber="437">S. ogawai</emphasis>
</taxonomicName>
” (
<collectingCountry id="F32FBFC6E94FFF81FEDCB980FE4BF901" box="[374,431,1689,1710]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
) + ”
<taxonomicName id="4C3884D5E94FFF81FE70B980FD99F901" authorityName="Shinohara" authorityYear="1984" box="[474,637,1689,1710]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="mushashiensis">
<emphasis id="B94C2344E94FFF81FE70B980FD99F901" box="[474,637,1689,1710]" italics="true" pageId="26" pageNumber="437">S. mushashiensis</emphasis>
</taxonomicName>
” (
<collectingCountry id="F32FBFC6E94FFF81FD3EB980FD2AF901" box="[660,718,1689,1710]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FD72B980FCB6F901" box="[728,850,1689,1711]" name="South Korea" pageId="26" pageNumber="437">South Korea</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8B87FF56E94FFF81FF3DB9AEFE7CF8A9" blockId="26.[151,1437,867,1799]" pageId="26" pageNumber="437">
<keyLead id="30C95C63E94FFF81FF3DB9AEFE7CF8A9" pageId="26" pageNumber="437">
- Two basal antennal articles virtually lacking setae dorsally. Ultimate sternite trapeziform. Distal articles of ultimate leg not setose (
<figureCitation id="1303E3D3E94FFF81FE83B9CDFE82F945" box="[297,358,1748,1770]" captionStart="FIGURES 6" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[242,1346,184,1917]" captionTargetId="figure-18@11.[241,1346,181,1919]" captionTargetPageId="11" captionText="FIGURES 6A–F. Scolopocryptops spinicaudus Wood, 1862; SVR.PO.202 A LBS 23 and right ultimate leg, laterally; B LBS 21–23 and ultimate legs, ventrally; C right ultimate leg, medially; Rc 7497 (ZMMU) D LBS 7 and 8, laterally; Rc 7225 (ZMMU) E left ultimate leg, dorsally; Rc 7497 (ZMMU) F LBS 13–16, dorsally." figureDoi="http://doi.org/10.5281/zenodo.7539954" httpUri="https://zenodo.org/record/7539954/files/figure.png" pageId="26" pageNumber="437">Figs 6</figureCitation>
AC), rarely with numerous setae (
<figureCitation id="1303E3D3E94FFF81FD06B9CDFD12F945" box="[684,758,1748,1770]" captionStart="FIGURES 6" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[242,1346,184,1917]" captionTargetId="figure-18@11.[241,1346,181,1919]" captionTargetPageId="11" captionText="FIGURES 6A–F. Scolopocryptops spinicaudus Wood, 1862; SVR.PO.202 A LBS 23 and right ultimate leg, laterally; B LBS 21–23 and ultimate legs, ventrally; C right ultimate leg, medially; Rc 7497 (ZMMU) D LBS 7 and 8, laterally; Rc 7225 (ZMMU) E left ultimate leg, dorsally; Rc 7497 (ZMMU) F LBS 13–16, dorsally." figureDoi="http://doi.org/10.5281/zenodo.7539954" httpUri="https://zenodo.org/record/7539954/files/figure.png" pageId="26" pageNumber="437">Fig. 6E</figureCitation>
)........
<taxonomicName id="4C3884D5E94FFF81FCF5B9CDFC0EF946" ID-CoL="4VSNB" authorityName="Wood" authorityYear="1862" box="[863,1002,1748,1769]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="spinicaudus">
<emphasis id="B94C2344E94FFF81FCF5B9CDFC0EF946" box="[863,1002,1748,1769]" italics="true" pageId="26" pageNumber="437">S. spinicaudus</emphasis>
</taxonomicName>
(
<collectingCountry id="F32FBFC6E94FFF81FC5CB9CDFBD0F946" box="[1014,1076,1748,1769]" name="South Korea" pageId="26" pageNumber="437">Korea</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FB94B9CDFB9EF945" box="[1086,1146,1748,1770]" name="China" pageId="26" pageNumber="437">China</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FB29B9CDFB28F945" box="[1155,1228,1748,1770]" name="Taiwan" pageId="26" pageNumber="437">Taiwan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FB7FB9CDFAEAF946" box="[1237,1294,1748,1769]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
,
<collectingCountry id="F32FBFC6E94FFF81FABDB9CDFA8EF945" box="[1303,1386,1748,1770]" name="Vietnam" pageId="26" pageNumber="437">Vietnam</collectingCountry>
) +
<taxonomicName id="4C3884D5E94FFF81FA20B9CDFE7CF8A9" ID-CoL="4VSN3" authority="(Japan)" authorityName="Shinohara" authorityYear="1990" baseAuthorityName="Japan" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="nipponicus">
<emphasis id="B94C2344E94FFF81FA20B9CDFEAFF8A8" italics="true" pageId="26" pageNumber="437">S. nipponicus</emphasis>
(
<collectingCountry id="F32FBFC6E94FFF81FEF2B9E8FE75F8A9" box="[344,401,1777,1798]" name="Japan" pageId="26" pageNumber="437">Japan</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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