333 lines
42 KiB
XML
333 lines
42 KiB
XML
<document id="3C28CC5851C7B32E9E445C73B7146820" ID-DOI="10.11646/zootaxa.2364.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5315506" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629660395335" checkinUser="felipe" docAuthor="Passos, Paulo, Fernandes, Ronaldo, Bérnils, Renato S. & De Moura-Leite, Julio C." docDate="2010" docId="0390751B3D6AFFC3FF61FCAFFC8FBD74" docLanguage="en" docName="zt02364p063.pdf" docOrigin="Zootaxa 2364 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2364.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Atractus caete Passos & Fernandes & Bérnils & De Moura-Leite 2010, sp. nov." docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="FFA90D633D6CFFCBFFF6FFADFFE0BA54" masterDocTitle="Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae) 2364" masterLastPageNumber="63" masterPageNumber="1" pageNumber="7" updateTime="1699091306492" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F5E2F9AF894E794B1D1D97555A2DF1F4">Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae) 2364</mods:title>
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<mods:roleTerm id="EACC08FCAB27FC663E5F4D2B08814E76">Author</mods:roleTerm>
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<mods:namePart id="B751B16E37F9FDA55E32D7205C8F5BA2">Passos, Paulo</mods:namePart>
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<mods:roleTerm id="D9AEB8728DDADDDF76D80832B8E6058A">Author</mods:roleTerm>
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<mods:namePart id="1C0AA7496338967350139DABCB3775D6">Fernandes, Ronaldo</mods:namePart>
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<mods:namePart id="897D9121CCDAE943E7FB647B3010C436">Bérnils, Renato S.</mods:namePart>
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</mods:role>
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<mods:namePart id="C525EA4A7E4CA97B1D58527490DB1DD5">De Moura-Leite, Julio C.</mods:namePart>
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<mods:typeOfResource id="D01FE2902AF1B7C4AB00105BF4B0F042">text</mods:typeOfResource>
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<mods:title id="A491E3CF60B2B76F507BAE392EF26B65">Zootaxa</mods:title>
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<mods:part id="E105FB2FBF0BFB6F575B993E03C2081E">
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<mods:date id="FB13A0006F248365115ABE15E1562BEA">2010</mods:date>
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<mods:number id="8A6D9DB5EA02306F4E3A2993ABF8666D">2010-02-19</mods:number>
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<mods:number id="BA456E44D806CF8CBB27C84234D231E7">2364</mods:number>
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<mods:number id="308D29395F1CEFDA70A085CD139ED974">1</mods:number>
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<mods:start id="1F06DA67CFA063CFA93666FDA7AEF45F">1</mods:start>
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<mods:url id="408C9C879CEB40FCC41C7AB4B593A2D4">https://biotaxa.org/Zootaxa/article/view/zootaxa.2364.1.1</mods:url>
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<mods:classification id="7029366CA21594D7FD1034C22AD46CED">journal article</mods:classification>
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<mods:identifier id="0615652C931BA2D04FD8EFAF6345CCF1" type="DOI">10.11646/zootaxa.2364.1.1</mods:identifier>
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<treatment id="0390751B3D6AFFC3FF61FCAFFC8FBD74" LSID="urn:lsid:plazi:treatment:0390751B3D6AFFC3FF61FCAFFC8FBD74" httpUri="http://treatment.plazi.org/id/0390751B3D6AFFC3FF61FCAFFC8FBD74" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
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<subSubSection id="C32397863D6AFFCDFF61FCAFFE47B948" box="[151,423,770,796]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="8B86C40D3D6AFFCDFF61FCAFFE47B948" blockId="6.[151,423,770,831]" box="[151,423,770,796]" pageId="6" pageNumber="7">
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<heading id="D0CE73613D6AFFCDFF61FCAFFE47B948" bold="true" box="[151,423,770,796]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<emphasis id="B94D181F3D6AFFCDFF61FCAFFE47B948" bold="true" box="[151,423,770,796]" pageId="6" pageNumber="7">
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<taxonomicName id="4C39BF8E3D6AFFCDFF61FCAFFEA3B948" authority="Passos & Fernandes & Bérnils & De Moura-Leite, 2010" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[151,323,770,796]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="caete" status="sp. nov.">
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<emphasis id="B94D181F3D6AFFCDFF61FCAFFEA3B948" bold="true" box="[151,323,770,796]" italics="true" pageId="6" pageNumber="7">Atractus caete</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27EA5643D6AFFCDFEBCFCAFFE47B948" box="[330,423,770,796]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32397863D6AFFCDFF61FC88FE8EB96B" box="[151,366,805,831]" pageId="6" pageNumber="7" type="description">
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<paragraph id="8B86C40D3D6AFFCDFF61FC88FE8EB96B" blockId="6.[151,423,770,831]" box="[151,366,805,831]" pageId="6" pageNumber="7">
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<figureCitation id="1302D8883D6AFFCDFF61FC88FF06B96B" box="[151,230,805,831]" captionStart="FIGURE 1" captionStartId="7.[151,255,1978,2002]" captionTargetBox="[230,1366,435,1954]" captionTargetId="figure-153@7.[201,1385,403,1973]" captionTargetPageId="7" captionText="FIGURE 1. Dorsal (A) and lateral (B) views of head, and lateral view of body (C) of the holotype of Atractus caete sp. nov. (MNRJ 16936). Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315510" httpUri="https://zenodo.org/record/5315510/files/figure.png" pageId="6" pageNumber="7">Figs. 1</figureCitation>
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,
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<figureCitation id="1302D8883D6AFFCDFF03FC88FEFBB96B" box="[245,283,805,831]" captionStart="FIGURE 2" captionStartId="9.[151,255,1856,1880]" captionTargetBox="[155,1428,180,1813]" captionTargetId="figure-50@9.[151,1436,166,1832]" captionTargetPageId="9" captionText="FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. francoi sp. nov. (MNRJ 17537), (D) A. guentheri (BMNH 1946.1.1.76), and (E) A. maculatus (BMNH 1946.1.6.46)." figureDoi="http://doi.org/10.5281/zenodo.5315512" httpUri="https://zenodo.org/record/5315512/files/figure.png" pageId="6" pageNumber="7">2A</figureCitation>
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,
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<figureCitation id="1302D8883D6AFFCDFEDEFC88FE8EB96B" box="[296,366,805,831]" captionStart="FIGURE 3" captionStartId="10.[151,255,1944,1968]" captionTargetBox="[160,1430,181,1936]" captionTargetId="figure-82@10.[151,1436,146,1952]" captionTargetPageId="10" captionText="FIGURE 3. General view in life of (A, B) Atractus caete sp. nov. from Chã Preta, state of Alagoas, photo by G. Skuk, (C) A. francoi sp. nov. from Santa Virgínia, state of São Paulo, photo by P. Hartmann, (D) A. guentheri from Ilhéus, state of Bahia, photo by A.J.S. Argôlo, and (E) A. pantostictus from Palmas, state of Tocantis, photo by O.A.V. Marques." figureDoi="http://doi.org/10.5281/zenodo.5315516" httpUri="https://zenodo.org/record/5315516/files/figure.png" pageId="6" pageNumber="7">3A–B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32397863D6AFFCDFF61FCD8FBD7B9EC" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="8B86C40D3D6AFFCDFF61FCD8FBD7B9EC" blockId="6.[151,1437,885,2035]" pageId="6" pageNumber="7">
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<materialsCitation id="3B51CE503D6AFFCDFF61FCD8FBD7B9EC" collectionCode="R" country="Brazil" elevation="360" latitude="-9.316667" location="Melgarejo" longLatPrecision="1300" longitude="-36.466667" municipality="Quebrangulo" pageId="6" pageNumber="7" specimenCode="MNRJ 16936, IVB 2983" specimenCount="1" stateProvince="Alagoas" typeStatus="holotype">
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<emphasis id="B94D181F3D6AFFCDFF61FCD8FEF2B9DB" bold="true" box="[151,274,885,911]" pageId="6" pageNumber="7">
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<typeStatus id="54827AAF3D6AFFCDFF61FCD8FEEDB9DB" box="[151,269,885,911]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Adult female,
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<specimenCode id="DB9F6C763D6AFFCDFE33FCDBFD84B9C4" box="[453,612,886,912]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ 16936</specimenCode>
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(formerly
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<specimenCode id="DB9F6C763D6AFFCDFD12FCDBFCBDB9C4" box="[740,861,886,912]" collectionCode="IVB" country="Czech Republic" httpUri="http://grbio.org/cool/iskk-kcgq" name="Institute of Vertebrate Biology, The Czech Academy of Sciences" pageId="6" pageNumber="7" type="Other type of research institution/biorepository">IVB 2983</specimenCode>
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), collected on 1986 by A.
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<collectionCode id="ED285CC83D6AFFCDFB60FCDBFB4DB9C4" box="[1174,1197,886,912]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="7">R</collectionCode>
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.
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<location id="8EE692D63D6AFFCDFB4FFCDBFAD4B9C4" LSID="urn:lsid:plazi:treatment:0390751B3D6AFFC3FF61FCAFFC8FBD74:8EE692D63D6AFFCDFB4FFCDBFAD4B9C4" box="[1209,1332,886,912]" country="Brazil" latitude="-9.316667" longLatPrecision="1300" longitude="-36.466667" municipality="Quebrangulo" name="Melgarejo" pageId="6" pageNumber="7" stateProvince="Alagoas">Melgarejo</location>
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, municipality of
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<collectingMunicipality id="6BE25E773D6AFFCDFEF7FC33FE7AB9EC" box="[257,410,926,952]" pageId="6" pageNumber="7">Quebrângulo</collectingMunicipality>
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(
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<geoCoordinate id="EE0DA2CA3D6AFFCDFE5CFC33FDEDB9EC" box="[426,525,926,952]" degrees="09" direction="south" minutes="19" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-9.316667">09°19’S</geoCoordinate>
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,
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<geoCoordinate id="EE0DA2CA3D6AFFCDFDECFC33FD65B9EC" box="[538,645,926,952]" degrees="36" direction="west" minutes="28" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-36.466667">36°28’W</geoCoordinate>
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, ca.
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<quantity id="4CC169E83D6AFFCDFD4CFC33FCE7B9EC" box="[698,775,926,952]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="7" unit="m" value="360.0">
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<elevation id="0014233E3D6AFFCDFD4CFC33FCE7B9EC" box="[698,775,926,952]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="7" unit="m" value="360.0">360 m</elevation>
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</quantity>
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), state of
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<collectingRegion id="49FD0AEF3D6AFFCDFC8FFC33FC3CB9EC" box="[889,988,926,952]" country="Brazil" name="Alagoas" pageId="6" pageNumber="7">Alagoas</collectingRegion>
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,
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<collectingCountry id="F32E849D3D6AFFCDFC1EFC33FBD3B9EC" box="[1000,1075,926,952]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32397863D6AFFCDFF30FC68FCCFBD2F" pageId="6" pageNumber="7" type="diagnosis">
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<paragraph id="8B86C40D3D6AFFCDFF30FC68FB9ABE87" blockId="6.[151,1437,885,2035]" pageId="6" pageNumber="7">
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||
<emphasis id="B94D181F3D6AFFCDFF30FC68FEA8B98B" bold="true" box="[198,328,965,991]" pageId="6" pageNumber="7">Diagnosis:</emphasis>
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<taxonomicName id="4C39BF8E3D6AFFCDFEB9FC6BFE17B98B" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[335,503,966,991]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="caete">
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<emphasis id="B94D181F3D6AFFCDFEB9FC6BFE17B98B" box="[335,503,966,991]" italics="true" pageId="6" pageNumber="7">Atractus caete</emphasis>
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</taxonomicName>
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is distinguished from all congeners by the following combination of characters: (1) 17/17/17 smooth dorsal scale rows; (2) two postoculars; (3) long loreal; (4) temporals 1+2; (5) usually six supralabials, third and fourth contacting orbit; (6) seven infralabials, first four contacting chinshields; (7) seven maxillary teeth; (8) three gular scale rows; (9) four preventrals; (10) 160 ventrals in the single female; (11) 16?–19 subcaudals; (12) dorsum uniformly black; (13) venter beige anteriorly, becoming gradually brown from the 15
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<superScript id="7C4C69453D6AFFCDFE8CFB23FE66BEC8" attach="left" box="[378,390,1166,1180]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
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||
ventral in adults, and cream marked with brown laterally in sub-adults; (14) moderate body size, female reaching
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||
<quantity id="4CC169E83D6AFFCDFE2EFB14FDD8BE87" box="[472,568,1209,1235]" metricMagnitude="-1" metricUnit="m" metricValue="3.76" pageId="6" pageNumber="7" unit="mm" value="376.0">376 mm</quantity>
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SVL; (15) small tail size on female (8.5% SVL).
|
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</paragraph>
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<paragraph id="8B86C40D3D6AFFCDFF30FB4DFC12BF97" blockId="6.[151,1437,885,2035]" pageId="6" pageNumber="7">
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||
<emphasis id="B94D181F3D6AFFCDFF30FB4DFE96BEAE" bold="true" box="[198,374,1248,1274]" pageId="6" pageNumber="7">Comparisons:</emphasis>
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Among all congeners,
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||
<taxonomicName id="4C39BF8E3D6AFFCDFD61FB4CFD19BEAE" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[663,761,1249,1274]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="caete">
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||
<emphasis id="B94D181F3D6AFFCDFD61FB4CFD19BEAE" box="[663,761,1249,1274]" italics="true" pageId="6" pageNumber="7">A. caete</emphasis>
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</taxonomicName>
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shares 17 dorsal scale rows, dorsal colour pattern uniformly black, and at least the posterior region of the venter of adults dark brown with posterior margin of ventral scales having pale (beige) pigmentation only with
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<emphasis id="B94D181F3D6AFFCDFCC9FA9CFADFBF1E" box="[831,1343,1329,1354]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFCC9FA9CFC36BF1E" authorityName="Roze" authorityYear="1961" box="[831,982,1329,1354]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="duidensis">A. duidensis</taxonomicName>
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,
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||
<taxonomicName id="4C39BF8E3D6AFFCDFC11FA9CFB96BF1E" authorityName="Amaral" authorityYear="1930" box="[999,1142,1329,1354]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="serranus">A. serranus</taxonomicName>
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||
,
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||
<taxonomicName id="4C39BF8E3D6AFFCDFB7EFA9CFADBBF1E" authorityName="Roze" authorityYear="1958" box="[1160,1339,1329,1354]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="steyermarki">A. steyermarki</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="4C39BF8E3D6AFFCDFA75FA9CFEFCBF26" authorityName="Amaral" authorityYear="1926" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="trihedrurus">
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<emphasis id="B94D181F3D6AFFCDFA75FA9CFEFCBF26" italics="true" pageId="6" pageNumber="7">A. trihedrurus</emphasis>
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</taxonomicName>
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||
.
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||
<taxonomicName id="4C39BF8E3D6AFFCDFED8FAF4FE3BBF26" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[302,475,1369,1394]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="caete">
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<emphasis id="B94D181F3D6AFFCDFED8FAF4FE3BBF26" box="[302,475,1369,1394]" italics="true" pageId="6" pageNumber="7">Atractus caete</emphasis>
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</taxonomicName>
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differs from these four species by having six supralabials and seven maxillary teeth (vs. seven supralabials in
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<emphasis id="B94D181F3D6AFFCDFDF2FA2CFCACBFCE" box="[516,844,1409,1434]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFDF2FA2CFD6EBFCE" authorityName="Amaral" authorityYear="1930" box="[516,654,1409,1434]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="serranus">A. serranus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFD6CFA2CFCA8BFCE" authorityName="Roze" authorityYear="1958" box="[666,840,1409,1434]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="steyermarki">A. steyermarki</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFC73FA2CFBCABFCE" authorityName="Amaral" authorityYear="1926" box="[901,1066,1409,1434]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="trihedrurus">
|
||
<emphasis id="B94D181F3D6AFFCDFC73FA2CFBCABFCE" box="[901,1066,1409,1434]" italics="true" pageId="6" pageNumber="7">A. trihedrurus</emphasis>
|
||
</taxonomicName>
|
||
, five maxillary teeth in
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFAB2FA2CFF14BF96" authorityName="Roze" authorityYear="1958" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="steyermarki">
|
||
<emphasis id="B94D181F3D6AFFCDFAB2FA2CFF14BF96" italics="true" pageId="6" pageNumber="7">A. steyermarki</emphasis>
|
||
</taxonomicName>
|
||
and eight to ten in
|
||
<emphasis id="B94D181F3D6AFFCDFE20FA04FCE2BF96" box="[470,770,1449,1474]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFE20FA04FD87BF96" authorityName="Roze" authorityYear="1961" box="[470,615,1449,1474]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="duidensis">A. duidensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFD85FA04FD1DBF96" authorityName="Amaral" authorityYear="1930" box="[627,765,1449,1474]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="serranus">A. serranus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFCCDFA04FC01BF96" authorityName="Amaral" authorityYear="1926" box="[827,993,1449,1474]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="trihedrurus">
|
||
<emphasis id="B94D181F3D6AFFCDFCCDFA04FC01BF96" box="[827,993,1449,1474]" italics="true" pageId="6" pageNumber="7">A. trihedrurus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B86C40D3D6AFFCDFF30FA7CFCCFBD2F" blockId="6.[151,1437,885,2035]" pageId="6" pageNumber="7">
|
||
Regarding sympatric or parapatric species,
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFD3AFA7CFCCCBFBE" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[716,812,1489,1514]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="caete">
|
||
<emphasis id="B94D181F3D6AFFCDFD3AFA7CFCCCBFBE" box="[716,812,1489,1514]" italics="true" pageId="6" pageNumber="7">A. caete</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFC3BFA7CFB88BFBE" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[973,1128,1489,1514]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="B94D181F3D6AFFCDFC3BFA7CFB88BFBE" box="[973,1128,1489,1514]" italics="true" pageId="6" pageNumber="7">A. maculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFB53FA7CFAFCBFBE" authorityName="Fernandes" authorityYear="1995" box="[1189,1308,1489,1514]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="potschi">
|
||
<emphasis id="B94D181F3D6AFFCDFB53FA7CFAFCBFBE" box="[1189,1308,1489,1514]" italics="true" pageId="6" pageNumber="7">A. potschi</emphasis>
|
||
</taxonomicName>
|
||
by having dorsum uniformly black in juveniles and adults, belly with lateral margins of ventrals marked with brown in juveniles and almost uniform brown in adults, and six supralabials (vs. dorsal ground colour cream with black transversal bands, blotches, or dots in juveniles and adults, venter uniformly creamish white, and seven supralabials in both species).
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFE05F9DCFD7CBCDE" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[499,668,1649,1674]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="caete">
|
||
<emphasis id="B94D181F3D6AFFCDFE05F9DCFD7CBCDE" box="[499,668,1649,1674]" italics="true" pageId="6" pageNumber="7">Atractus caete</emphasis>
|
||
</taxonomicName>
|
||
is somewhat similar to the central Brazilian
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFB5BF9DCFA89BCDE" authorityName="Da Cunha & Do Nascimento" authorityYear="1983" box="[1197,1385,1649,1674]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="albuquerquei">
|
||
<emphasis id="B94D181F3D6AFFCDFB5BF9DCFA89BCDE" box="[1197,1385,1649,1674]" italics="true" pageId="6" pageNumber="7">A. albuquerquei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFF61F934FF15BCE6" authorityName="Da Silva, Rodrigues Silva, Ribeiro, Souza & Do Amaral Souza" authorityYear="2005" box="[151,245,1689,1714]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="edioi">
|
||
<emphasis id="B94D181F3D6AFFCDFF61F934FF15BCE6" box="[151,245,1689,1714]" italics="true" pageId="6" pageNumber="7">A. edioi</emphasis>
|
||
</taxonomicName>
|
||
, in which the supralabials are reduced to five or six (apparently by fusion of the third and fourth scales), the dorsum mostly brown or black, and having three gular scale rows. However, the new species differs from both by having 17 dorsal scale rows, seven infralabials with first four contacting chinshields, single postdiastemal tooth, and lateral maxillary process poorly developed (vs. 15 dorsals, six infralabials with first three contacting chinshields, two postdiastemal teeth in both species, and lateral process moderately developed in
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFF03F8CCFEB1BD2E" authorityName="Da Silva, Rodrigues Silva, Ribeiro, Souza & Do Amaral Souza" authorityYear="2005" box="[245,337,1889,1914]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="edioi">
|
||
<emphasis id="B94D181F3D6AFFCDFF03F8CCFEB1BD2E" box="[245,337,1889,1914]" italics="true" pageId="6" pageNumber="7">A. edioi</emphasis>
|
||
</taxonomicName>
|
||
and well developed in
|
||
<taxonomicName id="4C39BF8E3D6AFFCDFD97F8CCFCFEBD2E" authorityName="Da Cunha & Do Nascimento" authorityYear="1983" box="[609,798,1889,1914]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="albuquerquei">
|
||
<emphasis id="B94D181F3D6AFFCDFD97F8CCFCFEBD2E" box="[609,798,1889,1914]" italics="true" pageId="6" pageNumber="7">A. albuquerquei</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32397863D6AFFC3FF30F825FC53BEA4" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="8B86C40D3D6AFFC3FF30F825FB72B816" blockId="6.[151,1437,885,2035]" lastBlockId="8.[151,1436,152,1824]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
|
||
<emphasis id="B94D181F3D6AFFCDFF30F825FDC8BDF6" bold="true" box="[198,552,1928,1954]" pageId="6" pageNumber="7">
|
||
Description of the
|
||
<typeStatus id="54827AAF3D6AFFCDFE45F825FDC3BDF6" box="[435,547,1928,1954]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Adult female, SVL
|
||
<quantity id="4CC169E83D6AFFCDFCE9F824FC65BDF7" box="[799,901,1929,1955]" metricMagnitude="-1" metricUnit="m" metricValue="3.76" pageId="6" pageNumber="7" unit="mm" value="376.0">376 mm</quantity>
|
||
, CL
|
||
<quantity id="4CC169E83D6AFFCDFC34F824FBF7BDF7" box="[962,1047,1929,1955]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="7" unit="mm" value="32.0">32 mm</quantity>
|
||
(8.5% SVL); body diameter 6.4 mm (1.7 % SVL); head length 13.4 mm (3.6% SVL); head width 6.3 mm (47% head length); interorbital distance 4.5 mm; rostro-orbital distance 4.1 mm (90% interorbital distance); naso-orbital distance 3.2 mm; cervical constriction barely distinct; head slightly arched in lateral view, round in dorsal view; snout truncate in lateral view, round in dorsal view; rostral sub-triangular in frontal view, 2.0 mm wide, 1.1 mm high, barely visible in dorsal view; internasal 0.8 mm long, as long as wide; internasal suture sinistral with respect to prefrontal suture; prefrontal 3.0 mm long, 2.6 mm wide; supraocular sub-trapezoidal, 1.0 mm long, about as long as wide; frontal triangular, 3.1 mm long, 3.8 mm wide; parietal 4.6 mm long, twice as long as wide; nasal divided; nostril located between prenasal and postnasal; postnasal 0.8 mm high, twice higher than long; loreal 2.6 mm long, 0.6 mm high, contacting second and third supralabials; eye diameter 1.2 mm; pupil subelliptical; two postoculars; upper postocular slightly higher (0.5 mm) and longer (0.4 mm) than lower postocular; temporals 1+2; anterior temporal 2.1 mm long, twice as long as high; upper posterior temporal elongate (3.6 mm), three times longer than wide; six supralabials, third and fourth contacting orbit; second supralabial higher than first and lower than third; sixth supralabial higher and longer than remaining supralabials; symphisial triangular, 1.9 mm wide, about three times broader than long; seven infralabials, first four contacting chinshields; first pair of infralabials in contact behind symphisial, preventing symphisial/chinshield contact; chinshields 3.7 mm long, about three times longer than wide; three gular scale rows; four preventrals; 160 ventrals; 19 (left side) and 20 (right side) subcaudals; 17/17/17 smooth dorsal scale rows; dorsals lacking apical pits, supra-anal tubercles, and keels; midbody diameter 5.7 mm; 10 dorsal scale rows at the level of second subcaudal; anal gland four subcaudals long; caudal spine short, robust, and rhomboid (
|
||
<figureCitation id="1302D8883D64FFC3FBCAFD85FB60B816" box="[1084,1152,552,578]" captionStart="FIGURE 1" captionStartId="7.[151,255,1978,2002]" captionTargetBox="[230,1366,435,1954]" captionTargetId="figure-153@7.[201,1385,403,1973]" captionTargetPageId="7" captionText="FIGURE 1. Dorsal (A) and lateral (B) views of head, and lateral view of body (C) of the holotype of Atractus caete sp. nov. (MNRJ 16936). Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5315510" httpUri="https://zenodo.org/record/5315510/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4694853D6BFFCCFF61F817FDF5BDA0" ID-DOI="http://doi.org/10.5281/zenodo.5315510" ID-Zenodo-Dep="5315510" httpUri="https://zenodo.org/record/5315510/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,255,1978,2002]" targetBox="[230,1366,435,1954]" targetPageId="7">
|
||
<paragraph id="8B86C40D3D6BFFCCFF61F817FDF5BDA0" blockId="7.[151,1436,1978,2036]" pageId="7" pageNumber="8">
|
||
<emphasis id="B94D181F3D6BFFCCFF61F817FEF9BD86" bold="true" box="[151,281,1978,2002]" pageId="7" pageNumber="8">FIGURE 1.</emphasis>
|
||
Dorsal (A) and lateral (B) views of head, and lateral view of body (C) of the holotype of
|
||
<taxonomicName id="4C39BF8E3D6BFFCCFB2BF817FA96BD85" authority="Passos & Fernandes & Bérnils & De Moura-Leite, 2010" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[1245,1398,1978,2001]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caete" status="sp. nov.">
|
||
<emphasis id="B94D181F3D6BFFCCFB2BF817FA96BD85" box="[1245,1398,1978,2001]" italics="true" pageId="7" pageNumber="8">Atractus caete</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EA5643D6BFFCCFA8BF817FF25BDA0" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(MNRJ 16936). Scale = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B86C40D3D64FFC3FF30FDE2FC47B8B6" blockId="8.[151,1436,152,1824]" pageId="8" pageNumber="9">
|
||
<emphasis id="B94D181F3D64FFC3FF30FDE2FE6CB83D" bold="true" box="[198,396,591,617]" pageId="8" pageNumber="9">Maxillary arch:</emphasis>
|
||
Arched in dorsal view, with six prediastemal and one postdiastemal teeth; prediastemal teeth large, well spaced, of similar size, curved posteriorly, angular in cross section, robust at base, narrower on the apices; maxillary diastema moderately long; postdiastemal teeth half size of prediastemal teeth; lateral process of maxilla poorly developed, lacking posterior projections.
|
||
</paragraph>
|
||
<paragraph id="8B86C40D3D64FFC3FF30FD42FC34BE73" blockId="8.[151,1436,152,1824]" pageId="8" pageNumber="9">
|
||
<emphasis id="B94D181F3D64FFC3FF30FD42FE0EB95D" bold="true" box="[198,494,751,777]" pageId="8" pageNumber="9">Colour in preservative:</emphasis>
|
||
Dorsum of head dark brown, except for pale brown spots on distal portion of prefrontals; head dark brown, except for irregular pale brown dots covering anterior portion of loreal and fourth to sixth supralabials; head uniformly black; symphisial and first three infralabials spotted with brown, remaining infralabials, chinshields, and preventrals beige; gulars with dark brown dots; venter uniform beige anteriorly, with brown blotches on lateral portions of ventrals from 15
|
||
<superScript id="7C4C69453D64FFC3FC17FC3DFC0DB9CA" attach="left" box="[993,1005,912,926]" fontSize="6" pageId="8" pageNumber="9">th</superScript>
|
||
scale; lateral blotches increase progressively in size, collapsing at the level of 20
|
||
<superScript id="7C4C69453D64FFC3FD47FC16FD5DB99D" attach="left" box="[689,701,955,969]" fontSize="6" pageId="8" pageNumber="9">th</superScript>
|
||
ventral; posterior to this point venter becomes uniform brown, only with posterior ventral margin pale (creamish brown) pigmented; tail brown with beige dots on posterior suture of subcaudals; dorsum of body uniformly dark brown (
|
||
<figureCitation id="1302D8883D64FFC3FC9FFBA0FC27BE73" box="[873,967,1037,1063]" captionStart="FIGURE 2" captionStartId="9.[151,255,1856,1880]" captionTargetBox="[155,1428,180,1813]" captionTargetId="figure-50@9.[151,1436,166,1832]" captionTargetPageId="9" captionText="FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. francoi sp. nov. (MNRJ 17537), (D) A. guentheri (BMNH 1946.1.1.76), and (E) A. maculatus (BMNH 1946.1.6.46)." figureDoi="http://doi.org/10.5281/zenodo.5315512" httpUri="https://zenodo.org/record/5315512/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B86C40D3D64FFC3FF30FB98FC53BEA4" blockId="8.[151,1436,152,1824]" pageId="8" pageNumber="9">
|
||
<emphasis id="B94D181F3D64FFC3FF30FB98FDC6BE1B" bold="true" box="[198,550,1077,1103]" pageId="8" pageNumber="9">Juvenile colouration in life:</emphasis>
|
||
An immature female (MUFAL not catalogued) has a distinctive juvenile colouration. Dorsum of head uniformly black; ventral margin of supralabials grayish white; gular region uniformly creamish white; venter mostly creamish white, with lateral margin of ventrals dark brown; tail mostly creamish white, with dark brown pigmentation restricted to subcaudal sutures; first four dorsal scales rows dark brown and remaining seven rows uniformly black (
|
||
<figureCitation id="1302D8883D64FFC3FCDCFB7BFC8EBEA4" box="[810,878,1238,1264]" captionStart="FIGURE 3" captionStartId="10.[151,255,1944,1968]" captionTargetBox="[160,1430,181,1936]" captionTargetId="figure-82@10.[151,1436,146,1952]" captionTargetPageId="10" captionText="FIGURE 3. General view in life of (A, B) Atractus caete sp. nov. from Chã Preta, state of Alagoas, photo by G. Skuk, (C) A. francoi sp. nov. from Santa Virgínia, state of São Paulo, photo by P. Hartmann, (D) A. guentheri from Ilhéus, state of Bahia, photo by A.J.S. Argôlo, and (E) A. pantostictus from Palmas, state of Tocantis, photo by O.A.V. Marques." figureDoi="http://doi.org/10.5281/zenodo.5315516" httpUri="https://zenodo.org/record/5315516/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
A-B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32397863D64FFC3FF30FB50FB90BFEC" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="8B86C40D3D64FFC3FF30FB50FB90BFEC" blockId="8.[151,1436,152,1824]" pageId="8" pageNumber="9">
|
||
<emphasis id="B94D181F3D64FFC3FF30FB50FEB7BF43" bold="true" box="[198,343,1277,1303]" pageId="8" pageNumber="9">Etymology:</emphasis>
|
||
The specific epithet caetê is a Tupi indigenous name, here employed as a noun in apposition alluding to the Atlantic Rainforest remnants (caá = forest; etê = true) where
|
||
<taxonomicName id="4C39BF8E3D64FFC3FBC3FA8BFB77BF6B" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[1077,1175,1318,1343]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caete">
|
||
<emphasis id="B94D181F3D64FFC3FBC3FA8BFB77BF6B" box="[1077,1175,1318,1343]" italics="true" pageId="8" pageNumber="9">A. caete</emphasis>
|
||
</taxonomicName>
|
||
was found. The Tupi word caetê is also the denomination for the extinct anthropophagous group of Brazilian Indians native to the region of the
|
||
<typeStatus id="54827AAF3D64FFC3FECDFADBFE8DBFC4" box="[315,365,1398,1424]" pageId="8" pageNumber="9">type</typeStatus>
|
||
locality. Portuguese colonizers later exterminated the “Caetés” after the natives captured and devoured Dom Pero Fernandes Sardinha, first bishop of
|
||
<collectingCountry id="F32E849D3D64FFC3FCA8FA33FC45BFEC" box="[862,933,1438,1464]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
|
||
on
|
||
<date id="FF87E2CD3D64FFC3FC27FA33FB8CBFEC" box="[977,1132,1438,1464]" pageId="8" pageNumber="9" value="1556-07-16">July 16 1556</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32397863D64FFC3FF30FA68FC99BC64" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="8B86C40D3D64FFC3FF30FA68FC99BC64" blockId="8.[151,1436,152,1824]" pageId="8" pageNumber="9">
|
||
<emphasis id="B94D181F3D64FFC3FF30FA68FE89BF8B" bold="true" box="[198,361,1477,1503]" pageId="8" pageNumber="9">Distribution:</emphasis>
|
||
Known only from the municipalities of Quebrângulo (
|
||
<geoCoordinate id="EE0DA2CA3D64FFC3FBF7FA6BFB84BFB4" box="[1025,1124,1478,1504]" degrees="09" direction="south" minutes="19" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-9.316667">09°19’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0DA2CA3D64FFC3FB85FA6BFB00BFB4" box="[1139,1248,1478,1504]" degrees="36" direction="west" minutes="28" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-36.466667">36°28’W</geoCoordinate>
|
||
) and Chã Preta (
|
||
<geoCoordinate id="EE0DA2CA3D64FFC3FF56FA43FEE0BC5C" box="[160,256,1518,1544]" degrees="09" direction="south" minutes="15" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-9.25">09º15’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0DA2CA3D64FFC3FEE7FA43FE9DBC5C" box="[273,381,1518,1544]" degrees="36" direction="west" minutes="18" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-36.3">36º18’W</geoCoordinate>
|
||
), state of
|
||
<collectingRegion id="49FD0AEF3D64FFC3FE0CFA43FDBDBC5C" box="[506,605,1518,1544]" country="Brazil" name="Alagoas" pageId="8" pageNumber="9">Alagoas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32E849D3D64FFC3FD9AFA43FD57BC5C" box="[620,695,1518,1544]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C39BF8E3D64FFC3FD30FA43FC92BC53" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[710,882,1518,1543]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caete">
|
||
<emphasis id="B94D181F3D64FFC3FD30FA43FC92BC53" box="[710,882,1518,1543]" italics="true" pageId="8" pageNumber="9">Atractus caete</emphasis>
|
||
</taxonomicName>
|
||
apparently inhabits small remnant patches of Submontane Rainforest between
|
||
<quantity id="4CC169E83D64FFC3FDEDF9BBFD40BC64" box="[539,672,1558,1584]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="8" pageNumber="9" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300–500 m</quantity>
|
||
elevation (
|
||
<figureCitation id="1302D8883D64FFC3FCD4F9BBFC8BBC64" box="[802,875,1558,1584]" captionStart="FIGURE 4" captionStartId="11.[151,255,1194,1218]" captionTargetBox="[161,1425,157,1177]" captionTargetId="figure-242@11.[161,1426,151,1204]" captionTargetPageId="11" captionText="FIGURE 4. Geographical distribution of Atractus caete, A. guentheri, A. maculatus, A. potschi, and A. ronnie." figureDoi="http://doi.org/10.5281/zenodo.5315518" httpUri="https://zenodo.org/record/5315518/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32397863D64FFC3FF30F990FC8FBD74" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="8B86C40D3D64FFC3FF30F990FC8FBD74" blockId="8.[151,1436,152,1824]" pageId="8" pageNumber="9">
|
||
<emphasis id="B94D181F3D64FFC3FF30F990FEA1BC03" bold="true" box="[198,321,1597,1623]" pageId="8" pageNumber="9">Remarks:</emphasis>
|
||
In the course of this study we examined photographs of a second known specimen of
|
||
<taxonomicName id="4C39BF8E3D64FFC3FACBF993FA7CBC03" authorityName="Passos & Fernandes & Bérnils & De Moura-Leite" authorityYear="2010" box="[1341,1436,1598,1623]" class="Reptilia" family="Dipsadidae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caete">
|
||
<emphasis id="B94D181F3D64FFC3FACBF993FA7CBC03" box="[1341,1436,1598,1623]" italics="true" pageId="8" pageNumber="9">A. caete</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1302D8883D64FFC3FF69F9CBFF10BCD4" box="[159,240,1638,1664]" captionStart="FIGURE 3" captionStartId="10.[151,255,1944,1968]" captionTargetBox="[160,1430,181,1936]" captionTargetId="figure-82@10.[151,1436,146,1952]" captionTargetPageId="10" captionText="FIGURE 3. General view in life of (A, B) Atractus caete sp. nov. from Chã Preta, state of Alagoas, photo by G. Skuk, (C) A. francoi sp. nov. from Santa Virgínia, state of São Paulo, photo by P. Hartmann, (D) A. guentheri from Ilhéus, state of Bahia, photo by A.J.S. Argôlo, and (E) A. pantostictus from Palmas, state of Tocantis, photo by O.A.V. Marques." figureDoi="http://doi.org/10.5281/zenodo.5315516" httpUri="https://zenodo.org/record/5315516/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
|
||
A-B). That individual (MUFAL not catalogued) was collected
|
||
<date id="FF87E2CD3D64FFC3FC20F9CBFB74BCD4" box="[982,1172,1638,1664]" pageId="8" pageNumber="9" value="2008-11">
|
||
November
|
||
<quantity id="4CC169E83D64FFC3FBACF9CBFB74BCD4" box="[1114,1172,1638,1664]" metricMagnitude="1" metricUnit="m" metricValue="5.10032" pageId="8" pageNumber="9" unit="in" value="2008.0">2008</quantity>
|
||
</date>
|
||
in the municipality of Chã Preta (
|
||
<geoCoordinate id="EE0DA2CA3D64FFC3FED7F923FE61BCFC" box="[289,385,1678,1704]" degrees="09" direction="south" minutes="15" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-9.25">09º15’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0DA2CA3D64FFC3FE67F923FE1ABCFC" box="[401,506,1678,1704]" degrees="36" direction="west" minutes="18" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-36.3">36º18’W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="4CC169E83D64FFC3FDC3F923FD65BCFC" box="[565,645,1678,1704]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="500.0">500 m</quantity>
|
||
), state of
|
||
<collectingRegion id="49FD0AEF3D64FFC3FCF7F923FC84BCFC" box="[769,868,1678,1704]" country="Brazil" name="Alagoas" pageId="8" pageNumber="9">Alagoas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32E849D3D64FFC3FC85F923FC5EBCFC" box="[883,958,1678,1704]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
|
||
. Because we had access only to photographs of that specimen we decided to not designate it as
|
||
<typeStatus id="54827AAF3D64FFC3FCB3F91BFC4ABC84" box="[837,938,1718,1744]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
of the new species. On the basis of available photographs it is possible to determine that the specimen (probably an immature female) has seven/six (left/right, respectively) supralabials and about 16 subcaudals.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |