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<document id="D947B563F4EF27A01FE2ABF5A3FF7A7F" ID-DOI="10.26107/RBZ-2023-0047" ID-ISSN="2345-7600" ID-Zenodo-Dep="10271307" ID-ZooBank="821BC4EC-5AF9-4727-84A3-C44839DFBE28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701819361795" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, Shane T. &amp; Castro, Peter" docDate="2023" docId="038EC92D79627E4EFCF86B6521AEF958" docLanguage="en" docName="RafflesBZool.71.606-631.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Sphenomerides trapezioides" docType="treatment" docVersion="2" lastPageNumber="622" masterDocId="FFB7B15579727E5EFFBA6E582531FFA7" masterDocTitle="Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="631" masterPageNumber="606" pageNumber="622" updateTime="1701819819889" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9BCF18CAEB8D063629C57C1E33B1F7EC">Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="DFFEA834FC6BAD217A734EB1D91125BF">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="B04BB54197F0C59FD3D2CFBA2AA7A47E">Ahyong, Shane T.</mods:namePart>
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<taxonomicName id="4C2703B879627E4EFCF86B652144FAF2" ID-CoL="4YVM2" baseAuthorityName="Wood-Mason in Wood-Mason &amp; Alcock" baseAuthorityYear="1891" box="[834,1141,1341,1365]" class="Malacostraca" family="Trapeziidae" genus="Sphenomerides" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="622" phylum="Arthropoda" rank="species" species="trapezioides">
<emphasis id="B953A42979627E4EFCF86B652144FAF2" bold="true" box="[834,1141,1341,1365]" italics="true" pageId="16" pageNumber="622">Sphenomerides trapezioides</emphasis>
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(Wood-Mason in
<bibRefCitation id="EFB605CA79627E4EFAF36B6521DCFAD2" author="Wood-Mason J &amp; Alcock A" pageId="16" pageNumber="622" pagination="258 - 272" refId="ref22116" refString="Wood-Mason J &amp; Alcock A (1891) Note on the results of the last seasons deep-sea dredging: Natural history notes from H. M. Indian Marine Survey Steamer Investigator, no. 21. Annals and Magazine of Natural History, Series 6, 7: 186 - 202, 258 - 272." type="journal article" year="1891">Wood-Mason &amp; Alcock, 1891</bibRefCitation>
)
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<paragraph id="8B98783B79627E4EFC2A6B252079FA32" blockId="16.[834,1430,1341,1429]" box="[912,1352,1405,1429]" pageId="16" pageNumber="622">
(
<figureCitation id="131C64BE79627E4EFC226B2526C6FA32" box="[920,1015,1405,1429]" captionStart="Fig" captionStartId="8.[113,148,1170,1192]" captionTargetBox="[113,1451,189,1144]" captionTargetId="figure-13@8.[113,1451,189,1144]" captionTargetPageId="8" captionText="Fig. 5. Frontal view of cephalothorax showing epistome, orbits, antennules, and antenna. A, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; D, Calocarcinus africanus, male (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; E, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; F, Cherusius triunguiculatus, female (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271321" httpUri="https://zenodo.org/record/10271321/files/figure.png" pageId="16" pageNumber="622">Figs. 5C</figureCitation>
,
<figureCitation id="131C64BE79627E4EFBBE6B252116FA32" box="[1028,1063,1405,1429]" captionStart="Fig" captionStartId="9.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1545]" captionTargetId="figure-7@9.[136,1474,189,1545]" captionTargetPageId="9" captionText="Fig. 6. Right maxilliped 3 (A–F); right major chela (G, H). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus, male (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271323" httpUri="https://zenodo.org/record/10271323/files/figure.png" pageId="16" pageNumber="622">6D</figureCitation>
,
<figureCitation id="131C64BE79627E4EFB8E6B262165FA32" box="[1076,1108,1406,1429]" captionStart="Fig" captionStartId="11.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1546]" captionTargetId="figure-8@11.[136,1474,189,1546]" captionTargetPageId="11" captionText="Fig.7. Male anterior thoracic sternum and pleon.A,Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; D, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271327" httpUri="https://zenodo.org/record/10271327/files/figure.png" pageId="16" pageNumber="622">7E</figureCitation>
,
<figureCitation id="131C64BE79627E4EFBDB6B2521ACFA32" box="[1121,1181,1405,1429]" captionStart="Fig" captionStartId="13.[136,171,1937,1959]" captionTargetBox="[144,1464,189,1923]" captionTargetId="figure-8@13.[144,1464,189,1923]" captionTargetPageId="13" captionText="Fig. 8. Posterior part of male thoracic sternum and pleon (A–F); sternopleonal cavity showing tubercle of pleonal-locking mechanism on sternite 5 (G–L). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, I, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, L. Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; K, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia." figureDoi="http://doi.org/10.5281/zenodo.10271329" httpUri="https://zenodo.org/record/10271329/files/figure.png" pageId="16" pageNumber="622">8D, J</figureCitation>
,
<figureCitation id="131C64BE79627E4EFB116B2521ECFA32" box="[1195,1245,1405,1429]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="16" pageNumber="622">9I, J</figureCitation>
,
<figureCitation id="131C64BE79627E4EFB506B252026FA32" box="[1258,1303,1405,1429]" captionStart="Fig" captionStartId="17.[136,171,1911,1933]" captionTargetBox="[136,1474,227,1896]" captionTargetId="figure-115@17.[136,1474,227,1896]" captionTargetPageId="17" captionText="Fig. 10. Vulvae. A, Tetralia nigrolineata (9.8 × 8.6 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (8.3 × 6.7 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (12.2 × 10.1 mm) (ZRC 2007.0079), Singapore; D, Quadrella maculosa (5.9 × 5.0 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.5 × 5.7 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (12.0 × 10.1 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (14.5 × 10.0 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; J, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion." figureDoi="http://doi.org/10.5281/zenodo.10271335" httpUri="https://zenodo.org/record/10271335/files/figure.png" pageId="16" pageNumber="622">10E</figureCitation>
,
<figureCitation id="131C64BE79627E4EFA9F6B252071FA32" box="[1317,1344,1405,1429]" captionStart="Fig" captionStartId="21.[136,171,1975,1997]" captionTargetBox="[136,1474,249,1960]" captionTargetId="figure-7@21.[136,1474,249,1960]" captionTargetPageId="21" captionText="Fig. 12.Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar.A, Dorsal habitus; B, frontal view of cephalothorax; C, dorsal view of carapace; D, male pleonal somites 3–6 (ventral view) showing fusion between somites 3–5; E, left P4 dactylus and propodus; F, comb-like setal rows on left P4 dactylus." figureDoi="http://doi.org/10.5281/zenodo.10271339" httpUri="https://zenodo.org/record/10271339/files/figure.png" pageId="16" pageNumber="622">12</figureCitation>
)
</paragraph>
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<treatmentCitation id="0A865E2A79627E4EFC976BE4209AFA75" author="Wood-Mason J &amp; Alcock A" box="[813,1451,1468,1490]" page="263" pageId="16" pageNumber="622" year="1891">
<taxonomicName id="4C2703B879627E4EFC976BE4209AFA75" ID-CoL="4YVMY" authority="Wood-Mason &amp; Alcock, 1891: 263" authorityName="Wood-Mason &amp; Alcock" authorityPageNumber="263" authorityYear="1891" box="[813,1451,1468,1490]" class="Malacostraca" family="Trapeziidae" genus="Sphenomerus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="622" phylum="Arthropoda" rank="species" species="trapezioides">
<emphasis id="B953A42979627E4EFC976BE42107FA76" box="[813,1078,1468,1489]" italics="true" pageId="16" pageNumber="622">Sphenomerus trapezioides</emphasis>
<bibRefCitation id="EFB605CA79627E4EFB856BE4209AFA75" author="Wood-Mason J &amp; Alcock A" box="[1087,1451,1468,1490]" pageId="16" pageNumber="622" pagination="258 - 272" refId="ref22116" refString="Wood-Mason J &amp; Alcock A (1891) Note on the results of the last seasons deep-sea dredging: Natural history notes from H. M. Indian Marine Survey Steamer Investigator, no. 21. Annals and Magazine of Natural History, Series 6, 7: 186 - 202, 258 - 272." type="journal article" year="1891">Wood-Mason &amp; Alcock, 1891: 263</bibRefCitation>
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</treatmentCitation>
(
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locality: Andaman Sea);
<bibRefCitation id="EFB605CA79627E4EFB3B6B812076FA49" author="Wood-Mason J" box="[1153,1351,1497,1519]" pageId="16" pageNumber="622" refId="ref22173" refString="Wood-Mason J (1892) Illustrations of the Zoology of H. M. Indian Marine Surveying Steamer Investigator, under the command of Commander A. Carpenter, R. N., D. S. O., and of Commander R. F. Hoskyn, R. N. Part I. Crustaceans. Director of the Royal Indian Marine, Calcutta, V pls. [5 unnumbered pages of explanation of pls.]" type="book" year="1892">Wood-Mason, 1892</bibRefCitation>
: pl. 5, fig. 2;
<treatmentCitation id="0A865E2A79627E4EFCD76BAE2114F9AC" author="Alcock A" box="[877,1061,1526,1548]" page="228" pageId="16" pageNumber="622" year="1898">
<bibRefCitation id="EFB605CA79627E4EFCD76BAE2114F9AC" author="Alcock A" box="[877,1061,1526,1548]" pageId="16" pageNumber="622" pagination="67 - 233" refId="ref17961" refString="Alcock A (1898) Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The family Xanthidae. Journal of the Asiatic Society of Bengal, 57 (2): 67 - 233." type="journal article" year="1898">Alcock, 1898: 228</bibRefCitation>
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.
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</paragraph>
<paragraph id="8B98783B79627E4EFC97684A204AF9D9" blockId="16.[813,1452,1468,1663]" pageId="16" pageNumber="622">
<treatmentCitationGroup id="AB375F1579627E4EFC97684A204AF9D9" pageId="16" pageNumber="622">
<taxonomicName id="4C2703B879627E4EFC97684A210AF980" ID-CoL="4YVM2" baseAuthorityName="Wood-Mason in Wood-Mason &amp; Alcock" baseAuthorityYear="1891" box="[813,1083,1554,1575]" class="Malacostraca" family="Trapeziidae" genus="Sphenomerides" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="622" phylum="Arthropoda" rank="species" species="trapezioides">
<emphasis id="B953A42979627E4EFC97684A210AF980" box="[813,1083,1554,1575]" italics="true" pageId="16" pageNumber="622">Sphenomerides trapezioides</emphasis>
</taxonomicName>
—
<treatmentCitation id="0A865E2A79627E4EFBD8684A2025F980" author="Serene R" box="[1122,1300,1554,1575]" page="207" pageId="16" pageNumber="622" year="1973">
<bibRefCitation id="EFB605CA79627E4EFBD8684A2025F980" author="Serene R" box="[1122,1300,1554,1575]" pageId="16" pageNumber="622" pagination="191 - 209" refId="ref21681" refString="Serene R (1973) Observations sur les especes des genres Quadrella Dana 1851 et Sphenomerides Rathbun 1898 (Decapoda- Brachyura). Bulletin de la Societe Zoologique de France, 98: 191 - 209." type="journal article" year="1973">Serène, 1973: 207</bibRefCitation>
</treatmentCitation>
, figs. 6, 27, 28, pl. 5, 1984: 289, fig. 196, pl. 42, fig. D;
<treatmentCitation id="0A865E2A79627E4EFB6C6877209AF9E3" author="Castro P &amp; Ng, PKL &amp; Ahyong, ST" box="[1238,1451,1583,1605]" page="34" pageId="16" pageNumber="622" year="2004">
<bibRefCitation id="EFB605CA79627E4EFB6C6877209AF9E3" author="Castro P &amp; Ng, PKL &amp; Ahyong, ST" box="[1238,1451,1583,1605]" pageId="16" pageNumber="622" pagination="1 - 70" refId="ref18680" refString="Castro P, Ng, PKL &amp; Ahyong, ST (2004) Phylogeny and systematics of the Trapeziidae Miers, 1886 (Crustacea: Brachyura), with the description of a new family. Zootaxa, 643: 1 - 70." type="journal article" year="2004">Castro et al., 2004: 34</bibRefCitation>
</treatmentCitation>
(in key), 60, pl. 4 fig. E;
<treatmentCitation id="0A865E2A79627E4EFBEB6814202AF9C5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1105,1307,1612,1634]" page="185" pageId="16" pageNumber="622" year="2008">
<bibRefCitation id="EFB605CA79627E4EFBEB6814202AF9C5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1105,1307,1612,1634]" pageId="16" pageNumber="622" pagination="1 - 286" refId="ref21055" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 185</bibRefCitation>
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;
<treatmentCitation id="0A865E2A79627E4EFA9268142649F9D8" author="Castro P" page="461" pageId="16" pageNumber="622" year="2013">
<bibRefCitation id="EFB605CA79627E4EFA9268142649F9D8" author="Castro P" pageId="16" pageNumber="622" pagination="437 - 466" refId="ref18529" refString="Castro P (2013) Brachyuran crabs (Crustacea, Brachyura: Crossotonotidae, Ethusidae, Euryplacidae, Goneplacidae, Latreilliidae, Palicidae, Tetraliidae, Trapeziidae) of the MAINBAZA, MIRIKI, and ATIMO VATAE expeditions to the Mozambique Channel and Madagascar. In: Ahyong ST, Chan T-Y, Corbari L &amp; Ng PKL (eds.) Tropical Deep-Sea Benthos, 27. Memoires du Museum national d'Histoire naturelle (Paris), 204: 437 - 466." type="journal article" year="2013">Castro, 2013: 461</bibRefCitation>
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, figs. 2F–H;
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<bibRefCitation id="EFB605CA79627E4EFC4068312017F9D8" author="Silambarasan K &amp; Kar AB &amp; Prasad GVA &amp; Pattnayak SK &amp; Das P &amp; Reddy DB &amp; Ramalingan LP" box="[1018,1318,1641,1663]" pageId="16" pageNumber="622" pagination="541 - 555" refId="ref21806" refString="Silambarasan K, Kar AB, Prasad GVA, Pattnayak SK, Das P, Reddy DB &amp; Ramalingan LP (2023) On some commensal anomuran and brachyuran crabs (Crustacea: Decapoda) off Visakhapatnam, Northwestern Bay of Bengal. Zootaxa, 5325: 541 - 555." type="journal article" year="2023">Silambarasan et al., 2023: 550</bibRefCitation>
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, fig. 4C.
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</paragraph>
</subSubSection>
<subSubSection id="C33D2BB079627E4EFC9768FF215BF918" box="[813,1130,1703,1727]" pageId="16" pageNumber="622" type="diagnosis">
<paragraph id="8B98783B79627E4EFC9768FF215BF918" blockId="16.[813,1130,1703,1727]" box="[813,1130,1703,1727]" pageId="16" pageNumber="622">
<emphasis id="B953A42979627E4EFC9768FF2690F918" bold="true" box="[813,929,1703,1727]" pageId="16" pageNumber="622">Diagnosis.</emphasis>
As for subfamily.
</paragraph>
</subSubSection>
<subSubSection id="C33D2BB079627E4EFC9568BF21AEF958" box="[815,1183,1767,1791]" pageId="16" pageNumber="622" type="discussion">
<paragraph id="8B98783B79627E4EFC9568BF21AEF958" blockId="16.[815,1183,1767,1791]" box="[815,1183,1767,1791]" pageId="16" pageNumber="622">
<emphasis id="B953A42979627E4EFC9568BF26ACF958" bold="true" box="[815,925,1767,1791]" pageId="16" pageNumber="622">Remarks.</emphasis>
See remarks for genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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