146 lines
18 KiB
XML
146 lines
18 KiB
XML
<document id="97E200CF178EC1744E122E843DE9D8B4" ID-DOI="10.5281/zenodo.5170031" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170031" ID-ZooBank="A5348E25-CC3E-476B-9AD2-0A6C3A37A61A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628280835588" checkinUser="felipe" docAuthor="Gordon, Robert D. & Hanley, Guy A." docDate="2017" docId="038E8B10D84FFFDBBF96E6E62C47F911" docLanguage="en" docName="InsectaMundi.0601.1-158.pdf" docOrigin="Insecta Mundi 2017 (601)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Prodilis joanna Gordon and Hanley 2017, new species" docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="FFB7F368D814FF80BF56E4652816FFCA" masterDocTitle="South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="90" updateTime="1699039770044" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="A94B87F55B539D72658C044AB2FB3A97">
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<mods:title id="3AC0D3B007A8F87BB8CBD2A5F77F4EBA">South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)</mods:title>
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<mods:name id="5B383D10FAD8D398ADE16AFB0CD988A8" type="personal">
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<mods:namePart id="B5D8631646C622932394C147CF3FBF26">Gordon, Robert D.</mods:namePart>
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<mods:name id="E9122DD6D853BA22F79749047298E13C" type="personal">
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<mods:roleTerm id="7850B2F4E6BFD8C9C4682370160E239C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B0914772F97D30AC3C431BAC7F646743">Hanley, Guy A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C20EDBB6CDCB037728F428160F2D0365">text</mods:typeOfResource>
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<mods:titleInfo id="D3C9AE524EAC2467E10DB2CF12B69B00">
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<mods:title id="CD0290813C68691B0D3516DDA44FF494">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="8EC84921A752CAE5C8669954DAE7B8B5">
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<mods:date id="5B1645E75B1871624664169E603E4AB8">2017</mods:date>
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<mods:detail id="783E484332D12F686A65CEF57B1301BB" type="pubDate">
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<mods:number id="B5BB6B2AE9AAF73E626DB7F63B298238">2017-12-29</mods:number>
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</mods:detail>
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<mods:detail id="121D17E42392AEED71F19B3016573CD2" type="volume">
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<mods:number id="2548EC708B46800D39BB6A5F0F9950DF">2017</mods:number>
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</mods:detail>
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<mods:detail id="519DA426745A1697DE2AE429E660AD58" type="issue">
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<mods:number id="C5D2A9B7E913628FA83E5E7F99A12F22">601</mods:number>
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<mods:classification id="7DE8951D9530F6B566F327D9CAA3B543">journal article</mods:classification>
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<mods:identifier id="74283D8F491013C9B7DDA1E80AB57A15" type="DOI">10.5281/zenodo.5170031</mods:identifier>
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<mods:identifier id="2287A667A85D55AB36CB04E1AABC0A45" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="77F891BD03A8FC55B1F71D29E73AA3DF" type="Zenodo-Dep">5170031</mods:identifier>
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<treatment id="038E8B10D84FFFDBBF96E6E62C47F911" ID-DOI="http://doi.org/10.5281/zenodo.5186238" ID-Zenodo-Dep="5186238" LSID="urn:lsid:plazi:treatment:038E8B10D84FFFDBBF96E6E62C47F911" httpUri="http://treatment.plazi.org/id/038E8B10D84FFFDBBF96E6E62C47F911" lastPageNumber="90" pageId="91" pageNumber="90">
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<subSubSection id="C33D698DD84FFFDBBF96E6E62B9AFD51" box="[192,908,643,667]" pageId="91" pageNumber="90" type="nomenclature">
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<paragraph id="8B983A06D84FFFDBBF96E6E62B9AFD51" blockId="91.[192,908,643,667]" box="[192,908,643,667]" pageId="91" pageNumber="90">
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<heading id="D0D08D6AD84FFFDBBF96E6E62B9AFD51" bold="true" box="[192,908,643,667]" fontSize="10" level="2" pageId="91" pageNumber="90" reason="3">
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<emphasis id="B953E614D84FFFDBBF96E6E62B9AFD51" bold="true" box="[192,908,643,667]" pageId="91" pageNumber="90">
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47.
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<taxonomicName id="4C274185D84FFFDBBFB8E6E62ACAFD51" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2017" box="[238,732,643,667]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="90" phylum="Arthropoda" rank="species" species="joanna" status="sp. nov.">
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<emphasis id="B953E614D84FFFDBBFB8E6E629D3FD51" bold="true" box="[238,453,643,667]" italics="true" pageId="91" pageNumber="90">Prodilis joanna</emphasis>
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Gordon and Hanley
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2605B6FD84FFFDBBDBCE6E62B9AFD51" box="[746,908,643,667]" pageId="91" pageNumber="90" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D698DD84FFFDBBF96E6A62936FA31" pageId="91" pageNumber="90" type="description">
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<paragraph id="8B983A06D84FFFDBBF96E6A62BCDFAF1" blockId="91.[192,1441,707,1339]" pageId="91" pageNumber="90">
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<emphasis id="B953E614D84FFFDBBF96E6A629AFFD11" bold="true" box="[192,441,707,731]" pageId="91" pageNumber="90">Description. Male</emphasis>
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<typeStatus id="549C84A4D84FFFDBBE92E6A62A27FD11" box="[452,561,707,731]" pageId="91" pageNumber="90" type="holotype">holotype</typeStatus>
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. Length
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<quantity id="4CDF97E3D84FFFDBBDF7E6A62B14FD11" box="[673,770,707,731]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="91" pageNumber="90" unit="mm" value="1.4">1.4 mm</quantity>
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, width
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<quantity id="4CDF97E3D84FFFDBBC34E6A62BD3FD11" box="[866,965,707,731]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="91" pageNumber="90" unit="mm" value="1.4">1.4 mm</quantity>
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; body nearly round, elytron with side rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny. Color yellowish brown (
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<figureCitation id="131C2683D84FFFDBBE22E76629CDFCD1" box="[372,475,771,795]" captionStart="Figures 575–590" captionStartId="150.[192,287,1644,1666]" captionTargetBox="[217,1405,207,1589]" captionTargetId="figure-20@150.[203,1450,192,1622]" captionTargetPageId="150" captionText="Figures 575–590. Prodilis spp. 575–580) Prodilis rosie 575) Habitus. 576) Basal abdominal ventrites. 577) Frons. 578) Phallobase ventral. 579) Phallobase lateral. 580) Sipho. 581–584) Prodilis joanna 581) Habitus. 582) Basal abdominal ventrites. 583) Frons. 584) Phallobase ventral. 585–590) Prodilis iris 585) Habitus. 586) Basal abdominal ventrite. 587) Frons. 588) Phallobase ventral. 589) Phallobase lateral. 590) Sipho, apex lost." figureDoi="http://doi.org/10.5281/zenodo.5170099" httpUri="https://zenodo.org/record/5170099/files/figure.png" pageId="91" pageNumber="90">Fig. 581</figureCitation>
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); head black except narrow apex of frons yellowish brown (
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<figureCitation id="131C2683D84FFFDBBBFCE7662D04FCD1" box="[1194,1298,771,795]" captionStart="Figures 575–590" captionStartId="150.[192,287,1644,1666]" captionTargetBox="[217,1405,207,1589]" captionTargetId="figure-20@150.[203,1450,192,1622]" captionTargetPageId="150" captionText="Figures 575–590. Prodilis spp. 575–580) Prodilis rosie 575) Habitus. 576) Basal abdominal ventrites. 577) Frons. 578) Phallobase ventral. 579) Phallobase lateral. 580) Sipho. 581–584) Prodilis joanna 581) Habitus. 582) Basal abdominal ventrites. 583) Frons. 584) Phallobase ventral. 585–590) Prodilis iris 585) Habitus. 586) Basal abdominal ventrite. 587) Frons. 588) Phallobase ventral. 589) Phallobase lateral. 590) Sipho, apex lost." figureDoi="http://doi.org/10.5281/zenodo.5170099" httpUri="https://zenodo.org/record/5170099/files/figure.png" pageId="91" pageNumber="90">Fig. 583</figureCitation>
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); pronotum medially dark brown, lateral 1/3 yellowish brown; antenna, mouthparts, epipleuron, legs yellow; abdomen brownish yellow except median portion of basal 3 ventrites brown. Head punctures large, separated by a diameter or less; pronotal punctures much smaller than on head, separated by less than 3 times a diameter; elytral punctures slightly larger than on pronotum, separated by 1 to 3 times a diameter; prosternal, mesosternal punctures large, separated by a diameter or less; metasternal punctures as large as on mesosternum, separated by less than a diameter except more widely spaced medially; abdomen with punctures on ventrites 1–2 large, separated by less than 3 times a diameter, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons not widened from vertex to clypeus, lateral margins parallel, 1.4 times width of eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin wide, equal in width from base to apex. Epipleuron flat, wide in basal ½, as wide as pronotal hypomeron, without feeble depressions for reception of femoral apices. Prosternum longer than wide, longer than short mesosternum, base arcuate medially, lateral carina slender, extended to apex of procoxa. Postcoxal line on ventrite 1 short, rounded, extended slightly beyond middle of ventrite (
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<figureCitation id="131C2683D84FFFDBBBA6E0A62D43FB11" box="[1264,1365,1219,1243]" captionStart="Figures 575–590" captionStartId="150.[192,287,1644,1666]" captionTargetBox="[217,1405,207,1589]" captionTargetId="figure-20@150.[203,1450,192,1622]" captionTargetPageId="150" captionText="Figures 575–590. Prodilis spp. 575–580) Prodilis rosie 575) Habitus. 576) Basal abdominal ventrites. 577) Frons. 578) Phallobase ventral. 579) Phallobase lateral. 580) Sipho. 581–584) Prodilis joanna 581) Habitus. 582) Basal abdominal ventrites. 583) Frons. 584) Phallobase ventral. 585–590) Prodilis iris 585) Habitus. 586) Basal abdominal ventrite. 587) Frons. 588) Phallobase ventral. 589) Phallobase lateral. 590) Sipho, apex lost." figureDoi="http://doi.org/10.5281/zenodo.5170099" httpUri="https://zenodo.org/record/5170099/files/figure.png" pageId="91" pageNumber="90">Fig. 582</figureCitation>
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) Apex of ventrite 5 truncate. Genitalia with long phallobase, basal lobe longer than paramere, slender, lateral margin gently curved to bluntly rounded apex, apex not emarginate; paramere nearly straight, sinuate in lateral view, without marginal serrations (
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<figureCitation id="131C2683D84FFFDBBDB5E1462B5CFAF1" box="[739,842,1315,1339]" captionStart="Figures 575–590" captionStartId="150.[192,287,1644,1666]" captionTargetBox="[217,1405,207,1589]" captionTargetId="figure-20@150.[203,1450,192,1622]" captionTargetPageId="150" captionText="Figures 575–590. Prodilis spp. 575–580) Prodilis rosie 575) Habitus. 576) Basal abdominal ventrites. 577) Frons. 578) Phallobase ventral. 579) Phallobase lateral. 580) Sipho. 581–584) Prodilis joanna 581) Habitus. 582) Basal abdominal ventrites. 583) Frons. 584) Phallobase ventral. 585–590) Prodilis iris 585) Habitus. 586) Basal abdominal ventrite. 587) Frons. 588) Phallobase ventral. 589) Phallobase lateral. 590) Sipho, apex lost." figureDoi="http://doi.org/10.5281/zenodo.5170099" httpUri="https://zenodo.org/record/5170099/files/figure.png" pageId="91" pageNumber="90">Fig. 584</figureCitation>
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); sipho lost.
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</paragraph>
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<paragraph id="8B983A06D84FFFDBBF96E106291EFA51" blockId="91.[192,1440,1379,1435]" pageId="91" pageNumber="90">
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<emphasis id="B953E614D84FFFDBBF96E106293AFAB1" bold="true" box="[192,300,1379,1403]" pageId="91" pageNumber="90">Female.</emphasis>
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Similar to male except female genitalia with spermathecal capsule, short, slightly curved medially.
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</paragraph>
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<paragraph id="8B983A06D84FFFDBBF96E1A62936FA31" blockId="91.[192,1440,1475,1531]" pageId="91" pageNumber="90">
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<emphasis id="B953E614D84FFFDBBF96E1A62954FA11" bold="true" box="[192,322,1475,1499]" pageId="91" pageNumber="90">Variation</emphasis>
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. Width
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<quantity id="4CDF97E3D84FFFDBBEF7E1A62A2FFA11" box="[417,569,1475,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="91" pageNumber="90" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
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. Lateral pronotal margin sometimes entirely black or with reduced pale margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D698DD84FFFDBBF96E2462AE7F9B1" pageId="91" pageNumber="90" type="materials_examined">
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<paragraph id="8B983A06D84FFFDBBF96E2462AE7F9B1" blockId="91.[192,1441,1570,1660]" pageId="91" pageNumber="90">
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<emphasis id="B953E614D84FFFDBBF96E246299AF9F1" bold="true" box="[192,396,1571,1595]" pageId="91" pageNumber="90">Type material.</emphasis>
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<materialsCitation id="3B4F305BD84FFFDBBEC3E2462D8AF991" collectingDate="1999-12-28" collectionCode="USNM" country="Costa Rica" elevation="100" latitude="10.433333" location="Tachigalia" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva." pageId="91" pageNumber="90" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
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<typeStatus id="549C84A4D84FFFDBBEC3E2462A12F9F1" box="[405,516,1571,1595]" pageId="91" pageNumber="90" type="holotype">Holotype</typeStatus>
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male;
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<collectingCountry id="F3307A96D84FFFDBBD0FE2462B10F9F1" box="[601,774,1571,1595]" name="Costa Rica" pageId="91" pageNumber="90">COSTA RICA</collectingCountry>
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:
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<collectingRegion id="49E3F4E4D84FFFDBBC4DE2462B96F9F1" box="[795,896,1571,1595]" country="Costa Rica" name="Heredia" pageId="91" pageNumber="90">Heredia</collectingRegion>
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,
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<collectingMunicipality id="6BFCA07CD84FFFDBBCDBE2412C6DF9F1" box="[909,1147,1571,1595]" pageId="91" pageNumber="90">Est. Biol. La Selva.</collectingMunicipality>
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||
<quantity id="4CDF97E3D84FFFDBBBD5E2462CE3F9F1" box="[1155,1269,1571,1595]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="91" pageNumber="90" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
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<elevation id="000ADD35D84FFFDBBBD5E2462CE3F9F1" box="[1155,1269,1571,1595]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="91" pageNumber="90" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE135CC1D84FFFDBBA55E2462D64F9F1" box="[1283,1394,1570,1595]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="91" pageNumber="90" precision="925" value="10.433333">
|
||
10
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<superScript id="7C52974ED84FFFDBBA76E2472D3EF9FA" attach="left" box="[1312,1320,1570,1584]" fontSize="6" pageId="91" pageNumber="90">o</superScript>
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26' N
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</geoCoordinate>
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<geoCoordinate id="EE135CC1D84FFFDBBA2DE24628E0F991" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="91" pageNumber="90" precision="925" value="-84.01667">
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||
84
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<superScript id="7C52974ED84FFFDBBACEE2472DB6F9FA" attach="none" box="[1432,1440,1570,1584]" fontSize="6" pageId="91" pageNumber="90">o</superScript>
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01W
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</geoCoordinate>
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, Prov. Alas, INBio–OET, FOT/41/01–40,
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<location id="8EF86CDDD84FFFDBBDAEE2262B61F991" LSID="urn:lsid:plazi:treatment:038E8B10D84FFFDBBF96E6E62C47F911:8EF86CDDD84FFFDBBDAEE2262B61F991" box="[760,887,1603,1627]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva." name="Tachigalia" pageId="91" pageNumber="90" stateProvince="Heredia">Tachigalia</location>
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costaricensis,
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<date id="FF991CC6D84FFFDBBB64E2262D37F991" box="[1074,1313,1603,1627]" pageId="91" pageNumber="90" value="1999-12-28">
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||
<collectingDate id="EFDDE52ED84FFFDBBB64E2262D37F991" box="[1074,1313,1603,1627]" pageId="91" pageNumber="90" value="1999-12-28">28 Diciembre 1999</collectingDate>
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</date>
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. (
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<collectionCode id="ED36A2C3D84FFFDBBA6FE2262D82F991" box="[1337,1428,1603,1627]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="91" pageNumber="90" type="Museum">USNM</collectionCode>
|
||
)
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||
</materialsCitation>
|
||
.
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||
<materialsCitation id="3B4F305BD84FFFDBBF96E2012AFBF9B1" box="[192,749,1635,1660]" collectingDate="1999-12-28" collectionCode="USNM" country="Costa Rica" elevation="100" latitude="10.433333" location="Tachigalia" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva." pageId="91" pageNumber="90" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
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<typeStatus id="549C84A4D84FFFDBBF96E201292BF9B6" box="[192,317,1636,1660]" pageId="91" pageNumber="90" type="paratype">Paratypes</typeStatus>
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; 2, same data as holotype. (
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<collectionCode id="ED36A2C3D84FFFDBBDDFE2062AF2F9B1" box="[649,740,1635,1659]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="91" pageNumber="90" type="Museum">USNM</collectionCode>
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||
)
|
||
</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33D698DD84FFFDBBF96E2C62C47F911" pageId="91" pageNumber="90" type="discussion">
|
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<paragraph id="8B983A06D84FFFDBBF96E2C62C47F911" blockId="91.[192,1440,1699,1755]" pageId="91" pageNumber="90">
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||
<emphasis id="B953E614D84FFFDBBF96E2C62952F971" bold="true" box="[192,324,1699,1723]" pageId="91" pageNumber="90">Remarks.</emphasis>
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||
<taxonomicName id="4C274185D84FFFDBBE1CE2C62A11F971" authorityName="Gordon and Hanley" authorityYear="2017" box="[330,519,1699,1723]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="90" phylum="Arthropoda" rank="species" species="joanna">
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<emphasis id="B953E614D84FFFDBBE1CE2C62A11F971" box="[330,519,1699,1723]" italics="true" pageId="91" pageNumber="90">Prodilis joanna</emphasis>
|
||
</taxonomicName>
|
||
is a small, distinctive species characterized by a unique dorsal color pattern, a black male head, widely spaced elytral punctures, and the male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |