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<document id="D12524C82BD33862F9EE650948E04BD6" ID-DOI="10.5281/zenodo.5170031" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170031" ID-ZooBank="A5348E25-CC3E-476B-9AD2-0A6C3A37A61A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628280835588" checkinUser="felipe" docAuthor="Gordon, Robert D. &amp; Hanley, Guy A." docDate="2017" docId="038E8B10D84BFFDFBF96E5C62A3CFA11" docLanguage="en" docName="InsectaMundi.0601.1-158.pdf" docOrigin="Insecta Mundi 2017 (601)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Prodilis madeline Gordon and Hanley 2017, new species" docType="treatment" docVersion="6" lastPageNumber="94" masterDocId="FFB7F368D814FF80BF56E4652816FFCA" masterDocTitle="South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="94" updateTime="1699039770044" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="14C08965F1662A74CE64EC6AB2675A48" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="154453C97CBFA3FB42ACB325A0D310F6">
<mods:title id="9F4A84D11F208762D7CE61B27A441926">South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)</mods:title>
</mods:titleInfo>
<mods:name id="345DD1A4B5E619B695A0ACBF4095764E" type="personal">
<mods:role id="54B499C116656D97529628BEB42532EF">
<mods:roleTerm id="67B109AD489CDE8A4FC2DE6097CD233B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68ECFD5CF4A1FC89A19021169D66086D">Gordon, Robert D.</mods:namePart>
</mods:name>
<mods:name id="AEDBD38147D2AFD8C0C86AF437A40C12" type="personal">
<mods:role id="F75B8955603B8D81AAE04BAAAB3E7EB8">
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</mods:role>
<mods:namePart id="4473A7A9D1720FA945DCF4C45476C763">Hanley, Guy A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2E7F8A6346D7B3E8331AB730935C6414">text</mods:typeOfResource>
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<mods:titleInfo id="C6CDDE5BD0DC6A9A96F1CBC0DBC9E2C4">
<mods:title id="BA2D3C7C67579927997D9CF07252BE5F">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="99C2E44998156D09679490C95CF07102">
<mods:date id="0989685D3FFF8B446BEC54721FEAAD9E">2017</mods:date>
<mods:detail id="2703C484429899F61446FD763133D9C4" type="pubDate">
<mods:number id="0781D1E2333DE3910981CBFE72420277">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="920192AF34E9B6B6F971AA8381769ABE" type="volume">
<mods:number id="27E2E7F83DD3ADFF107363AE4BF95271">2017</mods:number>
</mods:detail>
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<mods:number id="96FDF36A1D5C32B349E640E4BFB1FBAF">601</mods:number>
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<mods:classification id="455903712E4C3BA3A141BA8A62E90C2E">journal article</mods:classification>
<mods:identifier id="DE21A46D090CEC331BC57F6CA5EA2DD1" type="DOI">10.5281/zenodo.5170031</mods:identifier>
<mods:identifier id="C49A8352D239229620CAAD7347870E28" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="F387655BD91D0DD9516E442D92CCB86D" type="ZooBank">A5348E25-CC3E-476B-9AD2-0A6C3A37A61A</mods:identifier>
</mods:mods>
<treatment id="038E8B10D84BFFDFBF96E5C62A3CFA11" ID-DOI="http://doi.org/10.5281/zenodo.5186250" ID-Zenodo-Dep="5186250" LSID="urn:lsid:plazi:treatment:038E8B10D84BFFDFBF96E5C62A3CFA11" httpUri="http://treatment.plazi.org/id/038E8B10D84BFFDFBF96E5C62A3CFA11" lastPageNumber="94" pageId="95" pageNumber="94">
<subSubSection id="C33D698DD84BFFDFBF96E5C62BBCFE71" box="[192,938,419,443]" pageId="95" pageNumber="94" type="nomenclature">
<paragraph id="8B983A06D84BFFDFBF96E5C62BBCFE71" blockId="95.[192,938,419,443]" box="[192,938,419,443]" pageId="95" pageNumber="94">
<heading id="D0D08D6AD84BFFDFBF96E5C62BBCFE71" bold="true" box="[192,938,419,443]" fontSize="10" level="2" pageId="95" pageNumber="94" reason="3">
<emphasis id="B953E614D84BFFDFBF96E5C62BBCFE71" bold="true" box="[192,938,419,443]" pageId="95" pageNumber="94">
53.
<taxonomicName id="4C274185D84BFFDFBFB8E5C62AECFE71" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2017" box="[238,762,419,443]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="94" phylum="Arthropoda" rank="species" species="madeline" status="sp. nov.">
<emphasis id="B953E614D84BFFDFBFB8E5C629F5FE71" bold="true" box="[238,483,419,443]" italics="true" pageId="95" pageNumber="94">Prodilis madeline</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A2605B6FD84BFFDFBC5EE5C62BBCFE71" box="[776,938,419,443]" pageId="95" pageNumber="94" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD84BFFDFBF96E58629C7FB31" pageId="95" pageNumber="94" type="description">
<paragraph id="8B983A06D84BFFDFBF96E5862A3CFBB1" blockId="95.[192,1441,483,1147]" pageId="95" pageNumber="94">
<emphasis id="B953E614D84BFFDFBF96E58629AEFE31" bold="true" box="[192,440,483,507]" pageId="95" pageNumber="94">Description. Male</emphasis>
<typeStatus id="549C84A4D84BFFDFBE94E5862A39FE31" box="[450,559,483,507]" pageId="95" pageNumber="94" type="holotype">holotype</typeStatus>
. Length
<quantity id="4CDF97E3D84BFFDFBDC9E5862AE9FE31" box="[671,767,483,507]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="95" pageNumber="94" unit="mm" value="2.1">2.1 mm</quantity>
, width
<quantity id="4CDF97E3D84BFFDFBC08E5862BD6FE31" box="[862,960,483,507]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="95" pageNumber="94" unit="mm" value="1.8">1.8 mm</quantity>
; body oval, elytron with side rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface shiny except elytron with trace of microsculpture. Color black (
<figureCitation id="131C2683D84BFFDFBD73E6462A9BFDF1" box="[549,653,547,571]" captionStart="Figures 608–625" captionStartId="152.[192,287,1617,1639]" captionTargetBox="[207,1422,202,1588]" captionTargetId="figure-20@152.[192,1440,187,1616]" captionTargetPageId="152" captionText="Figures 608–625. Prodilis spp. 608–613) Prodilis lynda 608) Habitus. 609) Basal abdominal ventrites. 610) Frons. 611) Phallobase ventral. 612) Phallobase lateral. 613) Sipho. 614–619) Prodilis madeline 614) Habitus. 615) Basal abdominal ventrites. 616) Frons. 617) Phallobase ventral 618) Phallobase lateral. 619) Sipho. 620–625) Prodilis mamie 620) Habitus. 621) Basal abdominal ventrites. 622) Frons. 623) Phallobase ventral. 624) Phallobase lateral. 625) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170105" httpUri="https://zenodo.org/record/5170105/files/figure.png" pageId="95" pageNumber="94">Fig. 614</figureCitation>
); head with apical ½ yellow (
<figureCitation id="131C2683D84BFFDFBCA4E6462C4CFDF1" box="[1010,1114,547,571]" captionStart="Figures 608–625" captionStartId="152.[192,287,1617,1639]" captionTargetBox="[207,1422,202,1588]" captionTargetId="figure-20@152.[192,1440,187,1616]" captionTargetPageId="152" captionText="Figures 608–625. Prodilis spp. 608–613) Prodilis lynda 608) Habitus. 609) Basal abdominal ventrites. 610) Frons. 611) Phallobase ventral. 612) Phallobase lateral. 613) Sipho. 614–619) Prodilis madeline 614) Habitus. 615) Basal abdominal ventrites. 616) Frons. 617) Phallobase ventral 618) Phallobase lateral. 619) Sipho. 620–625) Prodilis mamie 620) Habitus. 621) Basal abdominal ventrites. 622) Frons. 623) Phallobase ventral. 624) Phallobase lateral. 625) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170105" httpUri="https://zenodo.org/record/5170105/files/figure.png" pageId="95" pageNumber="94">Fig. 616</figureCitation>
); antenna, tibiae and tarsi yellow; mouthparts yellow except maxillary palpus brownish yellow; epipleuron dark reddish brown; femur yellowish brown; abdomen yellow except basal ventrite medially yellowish brown. Head punctures small, separated by a diameter or less; pronotal punctures as large as on head, separated by a diameter or less; elytral punctures larger than on pronotum, separated by a diameter or less, often nearly contiguous; prosternal punctures large, separated by less than 3 times a diameter; mesosternal and metasternal punctures large, separated by a diameter or less except middle of metasternum with punctures more widely spaced; abdomen with punctures on ventrites 1–3 large, separated by less than twice a diameter, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons widened from vertex to clypeus, 1.4 times width of eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin narrow, equal in width from base to apex. Epipleuron strongly descending externally, wide in basal ½, as wide as pronotal hypomeron, with feeble depressions for reception of femoral apices. Prosternum as wide as long, slightly longer than mesosternum, base broadly, weakly emarginate medially, lateral carina narrow, extended to apex of procoxa. Postcoxal line on ventrite 1 long, rounded, extended beyond midpoint of ventrite (
<figureCitation id="131C2683D84BFFDFBDFDE0662B03FBD1" box="[683,789,1027,1051]" captionStart="Figures 608–625" captionStartId="152.[192,287,1617,1639]" captionTargetBox="[207,1422,202,1588]" captionTargetId="figure-20@152.[192,1440,187,1616]" captionTargetPageId="152" captionText="Figures 608–625. Prodilis spp. 608–613) Prodilis lynda 608) Habitus. 609) Basal abdominal ventrites. 610) Frons. 611) Phallobase ventral. 612) Phallobase lateral. 613) Sipho. 614–619) Prodilis madeline 614) Habitus. 615) Basal abdominal ventrites. 616) Frons. 617) Phallobase ventral 618) Phallobase lateral. 619) Sipho. 620–625) Prodilis mamie 620) Habitus. 621) Basal abdominal ventrites. 622) Frons. 623) Phallobase ventral. 624) Phallobase lateral. 625) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170105" httpUri="https://zenodo.org/record/5170105/files/figure.png" pageId="95" pageNumber="94">Fig. 615</figureCitation>
). Apex of ventrite 5 arcuate. Genitalia with slender phallobase, basal lobe slender, much longer than paramere, sides nearly parallel to deeply emarginate apex; paramere short, slender, slightly curved to rounded apex, without marginal serrations (
<figureCitation id="131C2683D84BFFDFBA7AE0262D84FB91" box="[1324,1426,1091,1115]" captionStart="Figures 608–625" captionStartId="152.[192,287,1617,1639]" captionTargetBox="[207,1422,202,1588]" captionTargetId="figure-20@152.[192,1440,187,1616]" captionTargetPageId="152" captionText="Figures 608–625. Prodilis spp. 608–613) Prodilis lynda 608) Habitus. 609) Basal abdominal ventrites. 610) Frons. 611) Phallobase ventral. 612) Phallobase lateral. 613) Sipho. 614–619) Prodilis madeline 614) Habitus. 615) Basal abdominal ventrites. 616) Frons. 617) Phallobase ventral 618) Phallobase lateral. 619) Sipho. 620–625) Prodilis mamie 620) Habitus. 621) Basal abdominal ventrites. 622) Frons. 623) Phallobase ventral. 624) Phallobase lateral. 625) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170105" httpUri="https://zenodo.org/record/5170105/files/figure.png" pageId="95" pageNumber="94">Fig. 618</figureCitation>
); sipho long, slender (
<figureCitation id="131C2683D84BFFDFBEE3E0062A08FBB1" box="[437,542,1123,1147]" captionStart="Figures 608–625" captionStartId="152.[192,287,1617,1639]" captionTargetBox="[207,1422,202,1588]" captionTargetId="figure-20@152.[192,1440,187,1616]" captionTargetPageId="152" captionText="Figures 608–625. Prodilis spp. 608–613) Prodilis lynda 608) Habitus. 609) Basal abdominal ventrites. 610) Frons. 611) Phallobase ventral. 612) Phallobase lateral. 613) Sipho. 614–619) Prodilis madeline 614) Habitus. 615) Basal abdominal ventrites. 616) Frons. 617) Phallobase ventral 618) Phallobase lateral. 619) Sipho. 620–625) Prodilis mamie 620) Habitus. 621) Basal abdominal ventrites. 622) Frons. 623) Phallobase ventral. 624) Phallobase lateral. 625) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170105" httpUri="https://zenodo.org/record/5170105/files/figure.png" pageId="95" pageNumber="94">Fig. 619</figureCitation>
).
</paragraph>
<paragraph id="8B983A06D84BFFDFBF96E0C629A5FB71" blockId="95.[192,435,1187,1211]" box="[192,435,1187,1211]" pageId="95" pageNumber="94">
<emphasis id="B953E614D84BFFDFBF96E0C6293AFB71" bold="true" box="[192,300,1187,1211]" pageId="95" pageNumber="94">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B983A06D84BFFDFBF96E08629C7FB31" blockId="95.[192,465,1251,1275]" box="[192,465,1251,1275]" pageId="95" pageNumber="94">
<emphasis id="B953E614D84BFFDFBF96E0862954FB31" bold="true" box="[192,322,1251,1275]" pageId="95" pageNumber="94">Variation</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD84BFFDFBF96E1462A2FFA91" pageId="95" pageNumber="94" type="materials_examined">
<paragraph id="8B983A06D84BFFDFBF96E1462A2FFA91" blockId="95.[192,1440,1315,1371]" pageId="95" pageNumber="94">
<emphasis id="B953E614D84BFFDFBF96E146299CFAF1" bold="true" box="[192,394,1315,1339]" pageId="95" pageNumber="94">Type material.</emphasis>
<materialsCitation id="3B4F305BD84BFFDFBEC7E1462A2FFA91" collectingDate="1965-03-06" collectionCode="USNM" collectorName="J. A. Ramos Collector." country="Colombia" location="Zipaquirra - Pacho" pageId="95" pageNumber="94" specimenCount="1" stateProvince="Cundinamarca" typeStatus="holotype">
<typeStatus id="549C84A4D84BFFDFBEC7E1462A16FAF1" box="[401,512,1315,1339]" pageId="95" pageNumber="94" type="holotype">Holotype</typeStatus>
male;
<collectingCountry id="F3307A96D84BFFDFBD07E1462AFBFAF1" box="[593,749,1315,1339]" name="Colombia" pageId="95" pageNumber="94">COLOMBIA</collectingCountry>
:
<location id="8EF86CDDD84BFFDFBDA9E1462BCAFAF1" LSID="urn:lsid:plazi:treatment:038E8B10D84BFFDFBF96E5C62A3CFA11:8EF86CDDD84BFFDFBDA9E1462BCAFAF1" box="[767,988,1315,1339]" country="Colombia" name="Zipaquirra - Pacho" pageId="95" pageNumber="94" stateProvince="Cundinamarca">Zipaquirra–Pacho</location>
, Cund. (
<collectingRegion id="49E3F4E4D84BFFDFBB16E1462CECFAF1" box="[1088,1274,1315,1339]" country="Colombia" name="Cundinamarca" pageId="95" pageNumber="94">Cundinamarca</collectingRegion>
),
<date id="FF991CC6D84BFFDFBA58E1462D8DFAF1" box="[1294,1435,1315,1339]" pageId="95" pageNumber="94" value="1965-03-06">
<collectingDate id="EFDDE52ED84BFFDFBA58E1462D8DFAF1" box="[1294,1435,1315,1339]" pageId="95" pageNumber="94" value="1965-03-06">6 Mar 1965</collectingDate>
</date>
,
<collectorName id="26D25FD0D84BFFDFBF96E12629D5FA91" box="[192,451,1347,1371]" pageId="95" pageNumber="94">J.A. Ramos Collector.</collectorName>
(
<collectionCode id="ED36A2C3D84BFFDFBE84E1262A3BFA91" box="[466,557,1347,1371]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="95" pageNumber="94" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD84BFFDFBF96E1E62A3CFA11" pageId="95" pageNumber="94" type="discussion">
<paragraph id="8B983A06D84BFFDFBF96E1E62A3CFA11" blockId="95.[192,1441,1411,1499]" pageId="95" pageNumber="94">
<emphasis id="B953E614D84BFFDFBF96E1E62952FA51" bold="true" box="[192,324,1411,1435]" pageId="95" pageNumber="94">Remarks.</emphasis>
<taxonomicName id="4C274185D84BFFDFBE1DE1E62A32FA51" authorityName="Gordon and Hanley" authorityYear="2017" box="[331,548,1411,1435]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="94" phylum="Arthropoda" rank="species" species="madeline">
<emphasis id="B953E614D84BFFDFBE1DE1E62A32FA51" box="[331,548,1411,1435]" italics="true" pageId="95" pageNumber="94">Prodilis madeline</emphasis>
</taxonomicName>
is not particularly distinctive, but the mostly black body, a densely punctured elytron, and epipleuron strongly descending externally will usually serve to distinguish it from other known
<taxonomicName id="4C274185D84BFFDFBE35E1A62A3CFA11" box="[355,554,1475,1499]" class="Insecta" family="Coccinellidae" genus="Prodilis" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="94" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B953E614D84BFFDFBE35E1A629D3FA11" box="[355,453,1475,1499]" italics="true" pageId="95" pageNumber="94">Prodilis</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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