442 lines
72 KiB
XML
442 lines
72 KiB
XML
<document id="CB75C5B3B052C91211CAAEE8E0059AC8" ID-DOI="10.5281/zenodo.179290" ID-GBIF-Dataset="3218835d-a188-406e-96ec-346bd5911888" ID-ISSN="1175-5326" ID-Zenodo-Dep="179290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459961374711" checkinUser="plazi" docAuthor="Anker, Arthur, Hurt, Carla & Knowlton, Nancy" docDate="2007" docId="038E6046FFF0FF9F37E3FA2DFC6EAFE6" docLanguage="en" docName="zt01626p023.pdf" docOrigin="Zootaxa 1626" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Alpheus naos Anker, Hurt & Knowlton, 2007, n. sp." docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="FFB7183EFFFBFF8F3774FF98FFC6AB24" masterDocTitle="Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae) in Panama" masterLastPageNumber="23" masterPageNumber="1" pageNumber="12" updateTime="1698230876662" updateUser="plazi">
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<mods:titleInfo id="0CD1857192AD8FB7C8470015D87CB545">
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<mods:title id="115E4C99B0350BA18F07803A6B34A2E4">Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae) in Panama</mods:title>
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<mods:name id="3641EFD49AF277D378122ACC8B059B1E" type="personal">
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<mods:namePart id="B64314D68F2E956A0BD5ED15EC83F943">Anker, Arthur</mods:namePart>
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<mods:namePart id="A910C21313190D50F670B66455E9590E">Hurt, Carla</mods:namePart>
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<mods:name id="BE33C58B2F780EF1C44C43565FFA8FDA" type="personal">
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<mods:namePart id="7316AEAC68A30EDF68488B228CA40106">Knowlton, Nancy</mods:namePart>
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<mods:typeOfResource id="401F663C9E9796FF2408577AEFD2E0D2">text</mods:typeOfResource>
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<mods:title id="7A072583704583F2C330C08EAC6762EF">Zootaxa</mods:title>
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<mods:part id="8FF73B01742CB6B44C19D242BF01C6C1">
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<mods:date id="E89E6E1583BDFEBCF8BD8463FC847A4D">2007</mods:date>
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<mods:identifier id="6ADDCECAB4487488A8EB6E343C2D54E4" type="DOI">10.5281/zenodo.179290</mods:identifier>
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<treatment id="038E6046FFF0FF9F37E3FA2DFC6EAFE6" ID-DOI="http://doi.org/10.5281/zenodo.6247387" ID-GBIF-Taxon="119376642" ID-Zenodo-Dep="6247387" LSID="urn:lsid:plazi:treatment:038E6046FFF0FF9F37E3FA2DFC6EAFE6" httpUri="http://treatment.plazi.org/id/038E6046FFF0FF9F37E3FA2DFC6EAFE6" lastPageId="16" lastPageNumber="17" pageId="11" pageNumber="12">
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<subSubSection id="C33D82DBFFF0FF8437E3FA2DFE9FAED6" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8B98D150FFF0FF8437E3FA2DFE42AEEB" blockId="11.[151,388,1461,1522]" box="[151,388,1461,1487]" pageId="11" pageNumber="12">
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<heading id="D0D0663CFFF0FF8437E3FA2DFE42AEEB" bold="true" box="[151,388,1461,1487]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
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<emphasis id="B9530D42FFF0FF8437E3FA2DFE42AEEB" bold="true" box="[151,388,1461,1487]" pageId="11" pageNumber="12">
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<taxonomicName id="4C27AAD3FFF0FF8437E3FA2DFEFCAEEB" ID-CoL="85LLV" box="[151,314,1461,1487]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
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<emphasis id="B9530D42FFF0FF8437E3FA2DFEFCAEEB" bold="true" box="[151,314,1461,1487]" italics="true" pageId="11" pageNumber="12">Alpheus naos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A260B039FFF0FF843635FA2DFE42AEEB" box="[321,388,1461,1487]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B98D150FFF0FF8437E3FA40FE9FAED6" blockId="11.[151,388,1461,1522]" box="[151,345,1496,1522]" pageId="11" pageNumber="12">
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<figureCitation id="131CCDD5FFF0FF8437E3FA40FEC1AED6" box="[151,263,1496,1522]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="12.[151,255,1881,1905]" captionStartId-1="13.[151,255,1933,1957]" captionStartId-2="14.[151,255,1350,1374]" captionTargetBox-0="[176,1390,196,1842]" captionTargetBox-1="[163,1421,201,1909]" captionTargetBox-2="[160,1404,210,1300]" captionTargetId-0="figure@12.[151,1436,164,1857]" captionTargetId-1="figure@13.[151,1436,194,1910]" captionTargetId-2="figure@14.[151,1436,194,1326]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." captionText-1="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." captionText-2="FIGURE 9. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm." httpUri-0="https://zenodo.org/record/179297/files/figure.png" httpUri-1="https://zenodo.org/record/179298/files/figure.png" httpUri-2="https://zenodo.org/record/179299/files/figure.png" pageId="11" pageNumber="12">Figs. 7–9</figureCitation>
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,
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<figureCitation id="131CCDD5FFF0FF843666FA40FEE8AED6" box="[274,302,1496,1522]" captionStart="FIGURE 11" captionStartId="18.[151,255,1495,1519]" captionTargetBox="[151,1434,434,1471]" captionTargetId="figure@18.[151,1436,434,1471]" captionTargetPageId="18" captionText="FIGURE 11. Color patterns in life. Alpheus christofferseni n. sp., male from Bocas del Toro, Panama (USNM 1100667) (a – c): a, b, dorsal view (note partly faded chromatopores in a); c, same, lateral view. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668) (d, e): d, dorsal view; e, lateral view. Alpheus aequus Kim & Abele, 1988, female from Coibita, Coiba Marine National Park, Panama (MNHN-Na 16389) (f): dorsal view." httpUri="https://zenodo.org/record/179301/files/figure.png" pageId="11" pageNumber="12">11</figureCitation>
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d, e
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D82DBFFF0FF8437E3F9BAFF3DAD90" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8B98D150FFF0FF8437E3F9BAFF3DAD90" blockId="11.[151,1437,1570,1996]" pageId="11" pageNumber="12">
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<emphasis id="B9530D42FFF0FF8437E3F9BAFE43AD18" bold="true" box="[151,389,1570,1596]" pageId="11" pageNumber="12">Material examined</emphasis>
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.
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<specimenCount id="9D211AD9FFF0FF8436E2F9BAFE21AD18" box="[406,487,1570,1596]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
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(CL 6.8, TL 20.1),
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<collectionCode id="ED364995FFF0FF8435ACF9BAFCEBAD18" LSID="urn:lsid:biocol.org:col:34871" box="[728,813,1570,1596]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12">USNM</collectionCode>
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1100668,
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<collectingCountry id="F33091C0FFF0FF8434D9F9BAFBCBAD18" box="[941,1037,1570,1596]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
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, Pacific coast, Amador causeway near
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<collectingCountry id="F33091C0FFF0FF8437A6F9D2FEE9AD40" box="[210,303,1610,1636]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
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City, Punta Culebra, rocky shore, under large mud-covered rocks on muddy sand, in burrow of
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<taxonomicName id="4C27AAD3FFF0FF8437E3F9EBFEE2ADA8" box="[151,292,1651,1676]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="11" pageNumber="12" phylum="Echiura" rank="genus">
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<emphasis id="B9530D42FFF0FF8437E3F9EBFEE2ADA8" box="[151,292,1651,1676]" italics="true" pageId="11" pageNumber="12">Listriolobus</emphasis>
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</taxonomicName>
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sp. (host collected and preserved), extreme low tide,
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<date id="FF99F790FFF0FF8434E6F9EAFBDBADA8" box="[914,1053,1650,1676]" pageId="11" pageNumber="12" value="2006-03-02">2 Mar 2006</date>
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, coll., A. Anker and C. Hurt [fcn 06-270].
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D82DBFFF0FF8037B2F95AFC30ACB6" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12" type="description">
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<paragraph id="8B98D150FFF0FF8437B2F95AFB2CACE8" blockId="11.[151,1437,1570,1996]" pageId="11" pageNumber="12">
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<emphasis id="B9530D42FFF0FF8437B2F95AFE91ADF8" bold="true" box="[198,343,1730,1756]" pageId="11" pageNumber="12">Description</emphasis>
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. Carapace smooth, not setose, laterally not compressed. Rostrum short, subtriangular, distally subacute (
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<figureCitation id="131CCDD5FFF0FF843666F972FE9AAC20" box="[274,348,1770,1796]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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a), slightly descendant and situated below dorsal-most portion of orbital hoods (
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<figureCitation id="131CCDD5FFF0FF843246F972FABBAC20" box="[1330,1405,1770,1796]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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b); mediorostral carina feebly marked, blunt, reaching beyond eye level, posteriorly flattening (
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<figureCitation id="131CCDD5FFF0FF8433A1F88AFADBAC08" box="[1237,1309,1810,1836]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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a); orbitorostral grooves very shallow; orbitorostral process not conspicuous. Orbital hoods moderately inflated, rounded, unarmed (
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<figureCitation id="131CCDD5FFF0FF84360BF8FAFE01AC58" box="[383,455,1890,1916]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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a). Pterygostomial angle rounded, not protruding (
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<figureCitation id="131CCDD5FFF0FF84336EF8FAFBA4AC58" box="[1050,1122,1890,1916]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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b). Branchiostegial margin fringed with setae. Eyes completely concealed in dorsal, lateral and partly frontal view. Ocellar beak welldeveloped, protruding vertically towards rostrum. Epistomial sclerite without projecting tooth.
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</paragraph>
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<caption id="DF5881D8FFF7FF8337E3F8C1FCD2ACFD" httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="12" pageNumber="13" targetBox="[176,1390,196,1842]" targetPageId="12">
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<paragraph id="8B98D150FFF7FF8337E3F8C1FCD2ACFD" blockId="12.[151,1436,1881,2009]" pageId="12" pageNumber="13">
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<emphasis id="B9530D42FFF7FF8337E3F8C1FEDDAC55" bold="true" box="[151,283,1881,1905]" pageId="12" pageNumber="13">FIGURE 7.</emphasis>
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<taxonomicName id="4C27AAD3FFF7FF833650F8C2FE73AC55" box="[292,437,1882,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
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<emphasis id="B9530D42FFF7FF833650F8C2FE73AC55" box="[292,437,1882,1905]" italics="true" pageId="12" pageNumber="13">Alpheus naos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A260B039FFF7FF8336C9F8C1FE3DAC55" box="[445,507,1881,1905]" pageId="12" pageNumber="13" rank="species">
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<emphasis id="B9530D42FFF7FF8336C9F8C1FE3DAC55" bold="true" box="[445,507,1881,1905]" pageId="12" pageNumber="13">n. sp.</emphasis>
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</taxonomicNameLabel>
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, male holotype from Punta Culebra, Isla
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<taxonomicName id="4C27AAD3FFF7FF8334B6F8C2FC3DAC55" box="[962,1019,1882,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
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, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption id="DF5881D8FFF6FF8237E3F815FCC0ACCE" httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="13" pageNumber="14" targetBox="[163,1421,201,1909]" targetPageId="13">
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<paragraph id="8B98D150FFF6FF8237E3F815FCC0ACCE" blockId="13.[151,1436,1933,2026]" pageId="13" pageNumber="14">
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<emphasis id="B9530D42FFF6FF8237E3F815FEDFAC81" bold="true" box="[151,281,1933,1957]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
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<taxonomicName id="4C27AAD3FFF6FF82366BF816FE6BAC81" box="[287,429,1934,1957]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
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<emphasis id="B9530D42FFF6FF82366BF816FE6BAC81" box="[287,429,1934,1957]" italics="true" pageId="13" pageNumber="14">Alpheus naos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A260B039FFF6FF8236C7F815FE28AC81" box="[435,494,1933,1957]" pageId="13" pageNumber="14" rank="species">
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<emphasis id="B9530D42FFF6FF8236C7F815FE28AC81" bold="true" box="[435,494,1933,1957]" pageId="13" pageNumber="14">n. sp.</emphasis>
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</taxonomicNameLabel>
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, male holotype from Punta Culebra, Isla
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<taxonomicName id="4C27AAD3FFF6FF8234D7F816FC1AAC81" box="[931,988,1934,1957]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
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, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm.
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||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5881D8FFF5FF8137E3FADEFB2CAEA4" httpUri="https://zenodo.org/record/179299/files/figure.png" pageId="14" pageNumber="15" targetBox="[160,1404,210,1300]" targetPageId="14">
|
||
<paragraph id="8B98D150FFF5FF8137E3FADEFB2CAEA4" blockId="14.[151,1436,1350,1408]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9530D42FFF5FF8137E3FADEFEDEAE7A" bold="true" box="[151,280,1350,1374]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C27AAD3FFF5FF81366BFADFFE6BAE7A" box="[287,429,1351,1374]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
|
||
<emphasis id="B9530D42FFF5FF81366BFADFFE6BAE7A" box="[287,429,1351,1374]" italics="true" pageId="14" pageNumber="15">Alpheus naos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFF5FF8136C0FADEFE29AE7A" box="[436,495,1350,1374]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B9530D42FFF5FF8136C0FADEFE29AE7A" bold="true" box="[436,495,1350,1374]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype from Punta Culebra, Isla
|
||
<taxonomicName id="4C27AAD3FFF5FF8134D2FADFFC19AE7A" box="[934,991,1351,1374]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
|
||
, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B98D150FFF5FF8137B1FA2CFB0FADC2" blockId="14.[151,1437,1460,2006]" pageId="14" pageNumber="15">
|
||
Antennular peduncles moderately stout, second segment slightly longer than dorsally visible portion of first segment, about 1.7–1.9 times as long as wide (
|
||
<figureCitation id="131CCDD5FFF5FF81358FFA44FC85AED2" box="[763,835,1500,1526]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
a); stylocerite short, distally blunt, almost reaching distal margin of first segment (
|
||
<figureCitation id="131CCDD5FFF5FF81368AF99CFD82AD3A" box="[510,580,1540,1566]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
a); mesioventral carina of first segment with tooth as illustrated (
|
||
<figureCitation id="131CCDD5FFF5FF81324CF99CFAB9AD3A" box="[1336,1407,1540,1566]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
c); lateral flagellum without independent secondary ramus, groups of aesthetasc tufts present starting from about ninth segment (
|
||
<figureCitation id="131CCDD5FFF5FF813624F9CCFE5EAD4A" box="[336,408,1620,1646]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
a, b). Antenna with basicerite lacking ventrolateral tooth (
|
||
<figureCitation id="131CCDD5FFF5FF81333AF9CCFB51AD4A" box="[1102,1175,1620,1646]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
b); carpocerite robust, reaching slightly beyond distal margin of antennular peduncle and scaphocerite; scaphocerite exceeding distal margin of antennular peduncle, with deep cleft between distolateral tooth and blade, lateral margin almost straight (
|
||
<figureCitation id="131CCDD5FFF5FF81378BF954FE83ADC2" box="[255,325,1740,1766]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
a), distolateral tooth reaching well beyond anterior margin of blade (
|
||
<figureCitation id="131CCDD5FFF5FF813311F954FB6AADC2" box="[1125,1196,1740,1766]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF5FF8137B1F96CFC38ACA2" blockId="14.[151,1437,1460,2006]" pageId="14" pageNumber="15">
|
||
Mouthparts (mandible, maxillule, maxilla, first and second maxillipeds) typical for
|
||
<taxonomicName id="4C27AAD3FFF5FF8133EEF96CFB3EAC29" box="[1178,1272,1780,1805]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9530D42FFF5FF8133EEF96CFB3EAC29" box="[1178,1272,1780,1805]" italics="true" pageId="14" pageNumber="15">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
. Third maxilliped moderately stout (
|
||
<figureCitation id="131CCDD5FFF5FF8136DAF884FE30AC12" box="[430,502,1820,1846]" captionStart="FIGURE 9" captionStartId="14.[151,255,1350,1374]" captionTargetBox="[160,1404,210,1300]" captionTargetId="figure@14.[151,1436,194,1326]" captionTargetPageId="14" captionText="FIGURE 9. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179299/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
|
||
a); coxa with lateral plate ear-shaped, distally acute, furnished with some setae; antepenultimate segment flattened, with ventral margin somewhat rugose; ultimate segment very setose; exopod reaching penultimate segment; arthrobranch well-developed (
|
||
<figureCitation id="131CCDD5FFF5FF8134EFF8F4FC24ACA2" box="[923,994,1900,1926]" captionStart="FIGURE 9" captionStartId="14.[151,255,1350,1374]" captionTargetBox="[160,1404,210,1300]" captionTargetId="figure@14.[151,1436,194,1326]" captionTargetPageId="14" captionText="FIGURE 9. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179299/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF5FF8137B1F80CFA5AACF2" blockId="14.[151,1437,1460,2006]" pageId="14" pageNumber="15">
|
||
Major cheliped (
|
||
<figureCitation id="131CCDD5FFF5FF8136FFF80CFE15AC8A" box="[395,467,1940,1966]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
a, b) with short, robust ischium; merus stout, about twice as long as wide, dorsal margin distally blunt, not projecting, ventromesial margin finely toothed, distally with small blunt tooth (
|
||
<figureCitation id="131CCDD5FFF5FF813219F824FA5AACF2" box="[1389,1436,1980,2006]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="14" pageNumber="15">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037E3FF00FF05AA86" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
|
||
8a); carpus cup-shaped, without distinct distal lobes; chela ovate, laterally compressed, subrectangular in cross-section; dorsal margin of palm with deep transverse notch subdistally (
|
||
<figureCitation id="131CCDD5FFF4FF80336DFF58FB99ABFE" box="[1049,1119,192,218]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
a, b), forming a broadly Ushaped groove in dorsal view; mesial face of palm smooth except for small depression near dorsal notch (
|
||
<figureCitation id="131CCDD5FFF4FF803218FF70FF60AA0E" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
b); lateral face with small depression near dorsal notch and broad, very shallow depression extending from base of pollex to about mid-length of palm (
|
||
<figureCitation id="131CCDD5FFF4FF8035C7FEA0FD3DAA76" box="[691,763,312,338]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
a); slightly more than 1/2 length of palm; dactylus with short, distally truncate plunger, latter with stamen-shaped sensillae (
|
||
<figureCitation id="131CCDD5FFF4FF8034BCFEF8FBDBAA5E" box="[968,1053,352,378]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Figs. 8</figureCitation>
|
||
a. b); adhesive discs small (
|
||
<figureCitation id="131CCDD5FFF4FF803218FEF8FF60AA86" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037B1FE28FDE7A9B6" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
|
||
Male minor cheliped (
|
||
<figureCitation id="131CCDD5FFF4FF8036B9FE28FDD4AAEE" box="[461,530,432,458]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
c–e) with robust merus, more than twice as long as wide; dorsal margin distally unarmed; ventromesial margin straight, distally blunt (
|
||
<figureCitation id="131CCDD5FFF4FF803426FE40FC5EAAD6" box="[850,920,472,498]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
c); carpus short, cup-shaped, with blunt distomesial lobe; chela with palm about 3/5 as long as fingers, without sculpture, ventral margin almost straight (
|
||
<figureCitation id="131CCDD5FFF4FF8037EBFDB0FF20A966" box="[159,230,552,578]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
d, e), distomesial margin with promenent blunt tooth (
|
||
<figureCitation id="131CCDD5FFF4FF803404FDB0FC71A966" box="[880,951,552,578]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
d); fingers long, slender, slightly gaping, laterally and mesially with rows of stiff setae along cutting edges, tips crossing when chela closed (
|
||
<figureCitation id="131CCDD5FFF4FF803268FDC8FAA6A94E" box="[1308,1376,592,618]" captionStart="FIGURE 8" captionStartId="13.[151,255,1933,1957]" captionTargetBox="[163,1421,201,1909]" captionTargetId="figure@13.[151,1436,194,1910]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, major cheliped, lateral view; b, same, distal merus, carpus and chela, mesial view; c, minor cheliped, dorsolateral view; d, same, chela, mesial view; e, same, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179298/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
d, e). Female minor cheliped unknown.
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037B1FD38FE5BA8F6" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
|
||
Second pereiopod relatively slender; ischium slightly longer than merus; carpus with five articles, with ratios (from proximal to distal) of approximately 5: 3: 1: 1: 2 (
|
||
<figureCitation id="131CCDD5FFF4FF8034E4FD50FC11A9C6" box="[912,983,712,738]" captionStart="FIGURE 9" captionStartId="14.[151,255,1350,1374]" captionTargetBox="[160,1404,210,1300]" captionTargetId="figure@14.[151,1436,194,1326]" captionTargetPageId="14" captionText="FIGURE 9. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179299/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||
b); chela simple, with fingers subequal to palm; palm as long as distal carpal article. Third pereiopod (
|
||
<figureCitation id="131CCDD5FFF4FF803401FD68FC7DA82E" box="[885,955,752,778]" captionStart="FIGURE 9" captionStartId="14.[151,255,1350,1374]" captionTargetBox="[160,1404,210,1300]" captionTargetId="figure@14.[151,1436,194,1326]" captionTargetPageId="14" captionText="FIGURE 9. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179299/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||
c) with ischium bearing one ventrolateral spine; merus unarmed, at least five times as long as wide; carpus unarmed; propodus with four slender spines along ventral margin and a pair of distal spines (
|
||
<figureCitation id="131CCDD5FFF4FF803597FCD8FCEBA87E" box="[739,813,832,858]" captionStart="FIGURE 9" captionStartId="14.[151,255,1350,1374]" captionTargetBox="[160,1404,210,1300]" captionTargetId="figure@14.[151,1436,194,1326]" captionTargetPageId="14" captionText="FIGURE 9. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, third maxilliped, lateral view; b, second pereiopod, lateral view; c, third pereiopod, lateral view. Scale bar: 1 mm." httpUri="https://zenodo.org/record/179299/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||
c); dactylus simple, conical, not spatulate, gradually curved towards subacute tip, about 1/2 propodus length. Fourth pereiopod similar to third. Fifth pereiopod with different article proportions; ischium with ventrolateral spine; propodus with ventral spines and welldeveloped setal brush.
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037B1FC78FA5BAEAE" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
|
||
Abdominal somites with broadly rounded posteroventral margins; fifth somite with rounded-angular posteroventral margin; sixth segment without articulated flap, posterior margin straight, dorsolateral projections rounded; preanal plate rounded. Male second pleopod (
|
||
<figureCitation id="131CCDD5FFF4FF803452FBA8FCABAF6E" box="[806,877,1072,1098]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
|
||
d) with appendix masculina longer than appendix interna, possibly partly fused to endopod, with slender spine-like setae distally and along endopod-facing margin (
|
||
<figureCitation id="131CCDD5FFF4FF80378DFB18FEF9AFBE" box="[249,319,1152,1178]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
|
||
d, e). Uropod (
|
||
<figureCitation id="131CCDD5FFF4FF80369DFB18FDEBAFBE" box="[489,557,1152,1178]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
|
||
f) with protopod bearing distally two strong acute teeth; exopod with diaeresis bearing one subtriangular lateral lobe proximal to distolateral spine and two well-marked rounded lobes at mid-length and mesially; lateral spine relatively strong, elongate; distal margins of exopod furnished with spinules (
|
||
<figureCitation id="131CCDD5FFF4FF80367EFB60FE95AE36" box="[266,339,1272,1298]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
|
||
f). Telson relatively narrow, tapering towards posterior margin (
|
||
<figureCitation id="131CCDD5FFF4FF803323FB60FB67AE36" box="[1111,1185,1272,1298]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">
|
||
Fig.
|
||
<quantity id="4CDF7CB5FFF4FF8033E5FB60FB67AE36" box="[1169,1185,1272,1298]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="15" pageNumber="16" unit="g" value="7.0">7</quantity>
|
||
</figureCitation>
|
||
g), with two pairs of short dorsal spines inserted at some distance from lateral margins, first pair at about mid-length, second pair at 3/4 of telson length (
|
||
<figureCitation id="131CCDD5FFF4FF8036FCFAD0FE08AE46" box="[392,462,1352,1378]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">
|
||
Fig.
|
||
<quantity id="4CDF7CB5FFF4FF8036CAFAD0FE08AE46" box="[446,462,1352,1378]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="15" pageNumber="16" unit="g" value="7.0">7</quantity>
|
||
</figureCitation>
|
||
g); posterior margin broadly rounded, with one pair of small posterolateral spines at each angle, mesial spines being at least twice as long as lateral spines (
|
||
<figureCitation id="131CCDD5FFF4FF8034B8FAE8FBD4AEAE" box="[972,1042,1392,1418]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
|
||
h); anal tubercles well-developed.
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037B1FA00FC87AE96" blockId="15.[151,1437,152,2018]" box="[197,833,1432,1458]" pageId="15" pageNumber="16">Gill/exopod formula as indicated for previous species.</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037B2FA27FD74AC4E" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9530D42FFF4FF8037B2FA27FECAAEFD" bold="true" box="[198,268,1471,1497]" pageId="15" pageNumber="16">Color</emphasis>
|
||
. Background mostly white semitransparent, with numerous red chromatophores forming several bands, including a broad lateral band on abdominal pleurae, interconnecting with two broad transverse bands on second and fifth somites, and narrow, more irregular bands on first and fourth somites, sixth somite with narrow transverse band along posterior margin; telson and uropods red except for most proximal portions; dorsal portion of carapace with broad band of red chromatophores, descending to flanks; central dorsal portions of abdominal somites, especially third, fourth and sixth, as well as most of posterior and anterior carapace colorless; rostral region, anterolateral margins of carapace and antennular peduncles semitransparent speckled (more or less densely) with red chromatophores; antennal peduncle and walking legs mostly semitransparent white; cheliped merus and carpus hyaline white with red chromatophores; chelae pinkish red due to numerous red chromatophores, distally (proximal to fingers) with pale orange tinge; fingers of major and minor chelae mostly ivory white (
|
||
<figureCitation id="131CCDD5FFF4FF803550F8C8FDB1AC4E" box="[548,631,1872,1898]" captionStart="FIGURE 11" captionStartId="18.[151,255,1495,1519]" captionTargetBox="[151,1434,434,1471]" captionTargetId="figure@18.[151,1436,434,1471]" captionTargetPageId="18" captionText="FIGURE 11. Color patterns in life. Alpheus christofferseni n. sp., male from Bocas del Toro, Panama (USNM 1100667) (a – c): a, b, dorsal view (note partly faded chromatopores in a); c, same, lateral view. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668) (d, e): d, dorsal view; e, lateral view. Alpheus aequus Kim & Abele, 1988, female from Coibita, Coiba Marine National Park, Panama (MNHN-Na 16389) (f): dorsal view." httpUri="https://zenodo.org/record/179301/files/figure.png" pageId="15" pageNumber="16">Fig. 11</figureCitation>
|
||
d, e).
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFF4FF8037B2F8EFFC30ACB6" blockId="15.[151,1437,152,2018]" box="[198,1014,1911,1938]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9530D42FFF4FF8037B2F8EFFF3FACB5" bold="true" box="[198,249,1911,1937]" pageId="15" pageNumber="16">Size</emphasis>
|
||
. The CL of the
|
||
<typeStatus id="549C6FF2FFF4FF8036C0F8E0FDDCACB6" box="[436,538,1912,1938]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
male specimen is
|
||
<quantity id="4CDF7CB5FFF4FF803586F8E0FC89ACB6" box="[754,847,1912,1938]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="15" pageNumber="16" unit="mm" value="6.8">6.8 mm</quantity>
|
||
, TL
|
||
<quantity id="4CDF7CB5FFF4FF8034F2F8E0FC34ACB6" box="[902,1010,1912,1938]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="15" pageNumber="16" unit="mm" value="20.1">20.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D82DBFFF4FF8037B2F807FD19ACC6" pageId="15" pageNumber="16" type="etymology">
|
||
<paragraph id="8B98D150FFF4FF8037B2F807FD19ACC6" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9530D42FFF4FF8037B2F807FE8DAC9D" bold="true" box="[198,331,1951,1977]" pageId="15" pageNumber="16">Etymology</emphasis>
|
||
. The new species is named after
|
||
<collectingCountry id="F33091C0FFF4FF803594F838FCCCAC9E" box="[736,778,1952,1978]" name="Iceland" pageId="15" pageNumber="16">Isla</collectingCountry>
|
||
<taxonomicName id="4C27AAD3FFF4FF803460F838FC95AC9E" box="[788,851,1952,1978]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
|
||
, where the
|
||
<typeStatus id="549C6FF2FFF4FF803494F838FBD4AC9E" box="[992,1042,1952,1978]" pageId="15" pageNumber="16">type</typeStatus>
|
||
locality (Punta Culebra) and the
|
||
<taxonomicName id="4C27AAD3FFF4FF8037E3F850FF15ACC6" box="[151,211,1992,2018]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
|
||
Marine Laboratory of the STRI are situated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D82DBFFEBFF9F37B2FF0FFA4FAB96" box="[198,1417,151,178]" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="8B98D150FFEBFF9F37B2FF0FFA4FAB96" blockId="16.[151,1437,151,1218]" box="[198,1417,151,178]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9530D42FFEBFF9F37B2FF0FFEA2AB95" bold="true" box="[198,356,151,177]" pageId="16" pageNumber="17">
|
||
<typeStatus id="549C6FF2FFEBFF9F37B2FF0FFEC2AB95" box="[198,260,151,177]" pageId="16" pageNumber="17">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. Punta Culebra,
|
||
<collectingCountry id="F33091C0FFEBFF9F355CFF00FD94AB96" box="[552,594,152,178]" name="Iceland" pageId="16" pageNumber="17">Isla</collectingCountry>
|
||
<taxonomicName id="4C27AAD3FFEBFF9F352EFF00FD5FAB96" box="[602,665,152,178]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
|
||
, Amador Causeway near
|
||
<collectingCountry id="F33091C0FFEBFF9F34B1FF00FBE4AB96" box="[965,1058,152,178]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
|
||
City, Pacific coast of
|
||
<collectingCountry id="F33091C0FFEBFF9F3250FF00FA43AB96" box="[1316,1413,152,178]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D82DBFFEBFF9F37B2FF27FE61AA86" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="8B98D150FFEBFF9F37B2FF27FF16AA26" blockId="16.[151,1437,151,1218]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9530D42FFEBFF9F37B2FF27FE99ABFD" bold="true" box="[198,351,191,217]" pageId="16" pageNumber="17">Distribution</emphasis>
|
||
. Eastern Pacific: presently known only from the
|
||
<typeStatus id="549C6FF2FFEBFF9F34C5FF58FC25ABFE" box="[945,995,192,218]" pageId="16" pageNumber="17">type</typeStatus>
|
||
locality on the Pacific coast of
|
||
<collectingCountry id="F33091C0FFEBFF9F3212FF58FF0AAA26" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFEBFF9F37B2FE97FE61AA86" blockId="16.[151,1437,151,1218]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9530D42FFEBFF9F37B2FE97FEEEAA0D" bold="true" box="[198,296,271,297]" pageId="16" pageNumber="17">Ecology</emphasis>
|
||
. The
|
||
<typeStatus id="549C6FF2FFEBFF9F3607FE88FE1CAA0E" box="[371,474,272,298]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
was collected by hand from a burrow of a large thalassematid echiuran (body length
|
||
<quantity id="4CDF7CB5FFEBFF9F3793FEA0FE8FAA76" box="[231,329,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="mm" value="150.0">150 mm</quantity>
|
||
, body diameter
|
||
<quantity id="4CDF7CB5FFEBFF9F3570FEA0FD42AA76" box="[516,644,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="16" pageNumber="17" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
|
||
, proboscis length
|
||
<quantity id="4CDF7CB5FFEBFF9F3423FEA0FC6BAA76" box="[855,941,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="17" unit="mm" value="30.0">30 mm</quantity>
|
||
),
|
||
<taxonomicName id="4C27AAD3FFEBFF9F34B5FEA0FB89AA75" box="[961,1103,312,337]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="genus">
|
||
<emphasis id="B9530D42FFEBFF9F34B5FEA0FB89AA75" box="[961,1103,312,337]" italics="true" pageId="16" pageNumber="17">Listriolobus</emphasis>
|
||
</taxonomicName>
|
||
sp. (see below). The burrow was dug horizontally into muddy sand under large mud-covered rocks, in the rocky intertidal, near the extreme low tide mark.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D82DBFFEBFF9F37B2FE37FAACA82D" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8B98D150FFEBFF9F37B2FE37FAACA82D" blockId="16.[151,1437,151,1218]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9530D42FFEBFF9F37B2FE37FDE3AAED" bold="true" box="[198,549,431,457]" pageId="16" pageNumber="17">Taxonomic remarks on host</emphasis>
|
||
. The original identification of the host as “
|
||
<taxonomicName id="4C27AAD3FFEBFF9F3352FE28FB72AAED" box="[1062,1204,432,457]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="genus">
|
||
<emphasis id="B9530D42FFEBFF9F3352FE28FB72AAED" box="[1062,1204,432,457]" italics="true" pageId="16" pageNumber="17">Listriolobus</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
<taxonomicName id="4C27AAD3FFEBFF9F3392FE28FEBFAAD6" authority="Chen & Yeh, 1958" authorityName="Chen & Yeh" authorityYear="1958" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
aff.
|
||
<emphasis id="B9530D42FFEBFF9F3261FE28FA5DAAED" box="[1301,1435,432,457]" italics="true" pageId="16" pageNumber="17">brevirostris</emphasis>
|
||
<bibRefCitation id="EFB6ACA1FFEBFF9F37E3FE40FEBFAAD6" author="Chen" box="[151,377,472,498]" pageId="16" pageNumber="22" refString="Chen, Y. & Yeh, C. C. (1958) Notes on some Gephyrea of China with descriptions of four new species. Acta Zoologica Sinica, 10, 265 - 278." type="journal article" year="1958">Chen & Yeh, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
” was tentative (G.-V. Murina, pers. comm.), partly because of the imperfect condition of the specimen from Punta Culebra (the animal died a few hours prior to preservation).
|
||
<taxonomicName id="4C27AAD3FFEBFF9F33F5FD98FA5DA93D" box="[1153,1435,512,537]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="species" species="brevirostris">
|
||
<emphasis id="B9530D42FFEBFF9F33F5FD98FA5DA93D" box="[1153,1435,512,537]" italics="true" pageId="16" pageNumber="17">Listriolobus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
was originally described from
|
||
<collectingCountry id="F33091C0FFEBFF9F3573FDB0FD8BA966" box="[519,589,552,578]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB6ACA1FFEBFF9F352BFDB0FC80A966" author="Chen" box="[607,838,552,578]" pageId="16" pageNumber="22" refString="Chen, Y. & Yeh, C. C. (1958) Notes on some Gephyrea of China with descriptions of four new species. Acta Zoologica Sinica, 10, 265 - 278." type="journal article" year="1958">Chen & Yeh, 1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB6ACA1FFEBFF9F3420FDB0FB5FA966" author="Stephen" box="[852,1177,552,578]" pageId="16" pageNumber="23" refString="Stephen, A. C. & Edmonds, S. J. (1972) The Phyla Sipuncula and Echiura. Trustees of the British Museum of Natural History, London, 588 pp." type="book" year="1972">Stephen & Edmonds, 1972</bibRefCitation>
|
||
) and is not known to occur in the eastern Pacific. On the other hand, specimens of its congener,
|
||
<taxonomicName id="4C27AAD3FFEBFF9F3375FDC8FF12A9B6" authority="Fisher, 1926" authorityName="Fisher" authorityYear="1926" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="species" species="bahamensis">
|
||
<emphasis id="B9530D42FFEBFF9F3375FDC8FB49A94D" box="[1025,1167,592,617]" italics="true" pageId="16" pageNumber="17">Listriolobus</emphasis>
|
||
cf.
|
||
<emphasis id="B9530D42FFEBFF9F33CFFDC8FA80A94D" box="[1211,1350,592,617]" italics="true" pageId="16" pageNumber="17">bahamensis</emphasis>
|
||
Fisher, 1926
|
||
</taxonomicName>
|
||
, were recently collected at Punta Culebra and other localities along the Pacific coast of
|
||
<collectingCountry id="F33091C0FFEBFF9F33A3FDE0FAF2A9B6" box="[1239,1332,632,658]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
|
||
(Murina & Anker, in prep.). They are morphologically (at least externally) and ecologically very similar to the individual found with
|
||
<taxonomicName id="4C27AAD3FFEBFF9F3633FD50FE58A9C5" box="[327,414,712,737]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F3633FD50FE58A9C5" box="[327,414,712,737]" italics="true" pageId="16" pageNumber="17">A. naos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F36D0FD5FFE20A9C5" box="[420,486,711,737]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F36D0FD5FFE20A9C5" bold="true" box="[420,486,711,737]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
As no other species of
|
||
<emphasis id="B9530D42FFEBFF9F3580FD50FCB5A9C5" box="[756,883,712,737]" italics="true" pageId="16" pageNumber="17">Listrilobus</emphasis>
|
||
are known from the shallow waters of the tropical eastern Pacific (
|
||
<bibRefCitation id="EFB6ACA1FFEBFF9F36F7FD68FD03A82E" author="Stephen" box="[387,709,752,778]" pageId="16" pageNumber="23" refString="Stephen, A. C. & Edmonds, S. J. (1972) The Phyla Sipuncula and Echiura. Trustees of the British Museum of Natural History, London, 588 pp." type="book" year="1972">Stephen & Edmonds, 1972</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C27AAD3FFEBFF9F35ADFD68FC6EA82D" box="[729,936,752,778]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="species" species="bahamensis">
|
||
<emphasis id="B9530D42FFEBFF9F35ADFD68FD2FA82D" box="[729,745,752,777]" italics="true" pageId="16" pageNumber="17">L</emphasis>
|
||
. cf.
|
||
<emphasis id="B9530D42FFEBFF9F3469FD68FC6EA82D" box="[797,936,752,777]" italics="true" pageId="16" pageNumber="17">bahamensis</emphasis>
|
||
</taxonomicName>
|
||
may well be the host of
|
||
<taxonomicName id="4C27AAD3FFEBFF9F33BCFD68FAE6A82D" box="[1224,1312,752,777]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F33BCFD68FAE6A82D" box="[1224,1312,752,777]" italics="true" pageId="16" pageNumber="17">A. naos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F325CFD77FAACA82D" box="[1320,1386,751,777]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F325CFD77FAACA82D" bold="true" box="[1320,1386,751,777]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D82DBFFEBFF9F37B2FC8FFC6EAFE6" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8B98D150FFEBFF9F37B2FC8FFCF2AFBE" blockId="16.[151,1437,151,1218]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9530D42FFEBFF9F37B2FC8FFEFEA815" bold="true" box="[198,312,791,817]" pageId="16" pageNumber="17">Remarks</emphasis>
|
||
. Within the
|
||
<taxonomicName id="4C27AAD3FFEBFF9F36BCFC80FD96A815" box="[456,592,792,817]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B9530D42FFEBFF9F36BCFC80FD96A815" box="[456,592,792,817]" italics="true" pageId="16" pageNumber="17">A. barbatus</emphasis>
|
||
</taxonomicName>
|
||
complex,
|
||
<taxonomicName id="4C27AAD3FFEBFF9F35BDFC80FCE6A815" box="[713,800,792,817]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F35BDFC80FCE6A815" box="[713,800,792,817]" italics="true" pageId="16" pageNumber="17">A. naos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F3453FC8FFCAFA815" box="[807,873,791,817]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F3453FC8FFCAFA815" bold="true" box="[807,873,791,817]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is morphologically closest to the eastern Pacific
|
||
<taxonomicName id="4C27AAD3FFEBFF9F37E3FCD8FEC8A87D" box="[151,270,832,857]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aequus">
|
||
<emphasis id="B9530D42FFEBFF9F37E3FCD8FEC8A87D" box="[151,270,832,857]" italics="true" pageId="16" pageNumber="17">A. aequus</emphasis>
|
||
</taxonomicName>
|
||
and the western Atlantic
|
||
<taxonomicName id="4C27AAD3FFEBFF9F353AFCD8FCDDA87D" box="[590,795,832,857]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="christofferseni" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F353AFCD8FCDDA87D" box="[590,795,832,857]" italics="true" pageId="16" pageNumber="17">A. christofferseni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F3453FCA7FCABA87D" box="[807,877,831,857]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F3453FCA7FCABA87D" bold="true" box="[807,877,831,857]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(see above). It differs from
|
||
<taxonomicName id="4C27AAD3FFEBFF9F33B9FCD8FA82A87D" box="[1229,1348,832,857]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aequus">
|
||
<emphasis id="B9530D42FFEBFF9F33B9FCD8FA82A87D" box="[1229,1348,832,857]" italics="true" pageId="16" pageNumber="17">A. aequus</emphasis>
|
||
</taxonomicName>
|
||
by the slightly narrower telson; the slightly longer second segment of the antennular peduncle; the broader blade of the scaphocerite; the lack of a slight sinus on the ventral margin of the palm of the minor chela; the less gaping fingers of the minor chela; a different palm/fingers ratio of the major chela; and the distinctly narrower merus of the third pereiopod. For differences between
|
||
<taxonomicName id="4C27AAD3FFEBFF9F35BFFC78FC53A8DD" box="[715,917,992,1017]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="christofferseni" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F35BFFC78FC53A8DD" box="[715,917,992,1017]" italics="true" pageId="16" pageNumber="17">A. christofferseni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F34E9FC47FC26A8DD" box="[925,992,991,1017]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F34E9FC47FC26A8DD" bold="true" box="[925,992,991,1017]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C27AAD3FFEBFF9F336EFC78FBB5A8DD" box="[1050,1139,992,1017]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F336EFC78FBB5A8DD" box="[1050,1139,992,1017]" italics="true" pageId="16" pageNumber="17">A. naos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F330FFC47FB78A8DD" box="[1147,1214,991,1017]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F330FFC47FB78A8DD" bold="true" box="[1147,1214,991,1017]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
see remarks under the former species. Another possible difference between these two species may lie in the appendix masculina: its proximal half is apparently partly fused to the endopod in
|
||
<taxonomicName id="4C27AAD3FFEBFF9F34F9FBA8FC2EAF6D" box="[909,1000,1072,1097]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F34F9FBA8FC2EAF6D" box="[909,1000,1072,1097]" italics="true" pageId="16" pageNumber="17">A. naos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F3480FBB7FBFCAF6D" box="[1012,1082,1071,1097]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F3480FBB7FBFCAF6D" bold="true" box="[1012,1082,1071,1097]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="131CCDD5FFEBFF9F3339FBA8FB51AF6E" box="[1101,1175,1072,1098]" captionStart="FIGURE 7" captionStartId="12.[151,255,1881,1905]" captionTargetBox="[176,1390,196,1842]" captionTargetId="figure@12.[151,1436,164,1857]" captionTargetPageId="12" captionText="FIGURE 7. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, second pleopod, lateral view; e, same, appendix masculina and appendix interna; f, left uropod, dorsal view; g, telson, dorsal view; h, same, detail of posterior margin. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179297/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
|
||
e), whereas the entire appendix masculina is free-standing in
|
||
<taxonomicName id="4C27AAD3FFEBFF9F3515FBC0FCEDAF55" box="[609,811,1112,1137]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="christofferseni" status="sp. nov.">
|
||
<emphasis id="B9530D42FFEBFF9F3515FBC0FCEDAF55" box="[609,811,1112,1137]" italics="true" pageId="16" pageNumber="17">A. christofferseni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A260B039FFEBFF9F3446FBCFFCB2AF55" box="[818,884,1111,1137]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B9530D42FFEBFF9F3446FBCFFCB2AF55" bold="true" box="[818,884,1111,1137]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="131CCDD5FFEBFF9F34F0FBC0FC15AF56" box="[900,979,1112,1138]" captionStart="FIGURE 1" captionStartId="4.[151,255,1718,1742]" captionTargetBox="[207,1397,218,1671]" captionTargetId="figure@4.[175,1422,201,1683]" captionTargetPageId="4" captionText="FIGURE 1. Alpheus christofferseni n. sp., female paratypes from Atol das Rocas, Brazil (MNRJ 17926) (a – h) and male holotype (MNRJ 17905) (i): a, frontal region, dorsal view; b, same, lateral view; c, tooth on mesioventral carina of first antennular segment, lateral view; d, scaphocerite of antenna, dorsal view; e, third maxilliped, lateral view; f, telson and left uropod, dorsal view; g, distal margin of exopod of right uropod; h, orbital hoods and rostrum of another specimen, dorsal view; i, appendix masculina and appendix interna of male second pleopod. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179291/files/figure.png" pageId="16" pageNumber="17">
|
||
Figs.
|
||
<date id="FF99F790FFEBFF9F34B2FBC0FC15AF56" box="[966,979,1112,1138]" pageId="16" pageNumber="17">1</date>
|
||
</figureCitation>
|
||
i, 6e) All these features, however, need to be confirmed by observations in additional specimens.
|
||
</paragraph>
|
||
<paragraph id="8B98D150FFEBFF9F37B2FB3FFC6EAFE6" blockId="16.[151,1437,151,1218]" box="[198,936,1191,1218]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9530D42FFEBFF9F37B2FB3FFE60AFE5" bold="true" box="[198,422,1191,1217]" pageId="16" pageNumber="17">GenBank number</emphasis>
|
||
.
|
||
<accessionNumber id="94744CB3FFEBFF9F36C8FB30FDF1AFE6" box="[444,567,1192,1218]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF092275" pageId="16" pageNumber="17">
|
||
EF
|
||
<date id="FF99F790FFEBFF9F36ABFB30FDF1AFE6" box="[479,567,1192,1218]" pageId="16" pageNumber="17" value="75-09-22">092275</date>
|
||
</accessionNumber>
|
||
(fcn 06-270, USNM 1100668).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |