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<document id="82C861E70102B07714A0185C029749E6" ID-DOI="10.11646/zootaxa.4743.2.12" ID-GBIF-Dataset="b2d2e507-f695-4ea7-b2cb-69b28d885d8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3687732" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582709235737" checkinUser="plazi" docAuthor="Hu, Fang-Shuo &amp; Bogri, Amalia" docDate="2020" docId="038E4022B109FFEDFF6DFEA95A42E8D2" docLanguage="en" docName="zootaxa.4743.2.12.pdf" docOrigin="Zootaxa 4743 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quwatanabius flavicornis" docType="treatment" docVersion="6" lastPageNumber="288" masterDocId="FFB7385AB10BFFEEFFFAFFE55B06E807" masterDocTitle="Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae)" masterLastPageNumber="288" masterPageNumber="285" pageNumber="287" updateTime="1698721560719" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DAABC320C7DB9EFB28273FD50767CD3E">Taxonomic notes on the genus Quwatanabius in Taiwan (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="9348FC83A4AABD208E0B86FB46395C9F">Hu, Fang-Shuo</mods:namePart>
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<mods:date id="FE73E3C97B1A50F4220095D1BC2F3B4C">2020</mods:date>
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<treatment id="038E4022B109FFEDFF6DFEA95A42E8D2" ID-DOI="http://doi.org/10.5281/zenodo.3688024" ID-GBIF-Taxon="162267519" ID-Zenodo-Dep="3688024" LSID="urn:lsid:plazi:treatment:038E4022B109FFEDFF6DFEA95A42E8D2" httpUri="http://treatment.plazi.org/id/038E4022B109FFEDFF6DFEA95A42E8D2" lastPageId="3" lastPageNumber="288" pageId="2" pageNumber="287">
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<paragraph id="8B98F134B109FFECFF6DFEA9580FE961" blockId="2.[151,1437,332,754]" box="[151,777,332,359]" pageId="2" pageNumber="287">
<heading id="D0D04658B109FFECFF6DFEA9580FE961" bold="true" box="[151,777,332,359]" fontSize="11" level="1" pageId="2" pageNumber="287" reason="1">
<emphasis id="B9532D26B109FFECFF6DFEA9580FE961" bold="true" box="[151,777,332,359]" pageId="2" pageNumber="287">
<taxonomicName id="4C278AB7B109FFECFF6DFEA95986E961" ID-CoL="4R7PZ" authority="(Sharp, 1889)" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[151,640,332,359]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B9532D26B109FFECFF6DFEA95AD6E960" bold="true" box="[151,464,332,359]" italics="true" pageId="2" pageNumber="287">Quwatanabius flavicornis</emphasis>
(Sharp, 1889)
</taxonomicName>
(
<figureCitation id="131CEDB1B109FFECFD6AFEA95807E961" box="[656,769,332,358]" captionStart="FIGURES 78" captionStartId="2.[151,258,1809,1831]" captionTargetBox="[163,1435,777,1782]" captionTargetId="figure@2.[151,1435,777,1782]" captionTargetPageId="2" captionText="FIGURES 78. Quwatanabius flavicornis 7. habitus, scale bar represents 1mm; 8. habitat at the location where the additional specimens of Q. flavicornis were collected." figureDoi="http://doi.org/10.5281/zenodo.3687736" httpUri="https://zenodo.org/record/3687736/files/figure.png" pageId="2" pageNumber="287">Figs. 78</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33DA2BFB109FFECFF6DFE975890E9D5" pageId="2" pageNumber="287" type="materials_examined">
<paragraph id="8B98F134B109FFECFF6DFE975890E9D5" blockId="2.[151,1437,332,754]" pageId="2" pageNumber="287">
<emphasis id="B9532D26B109FFECFF6DFE975A73E98D" bold="true" box="[151,373,370,394]" pageId="2" pageNumber="287">Material examined.</emphasis>
<materialsCitation id="3B4FFB69B109FFECFE81FE975FEEE9AA" ID-GBIF-Occurrence="2571176112" collectingDate="2019-03-23" collectionCode="BBM" collectorName="Renai To. &amp; F. S. Hu &amp; Det. By F. S. Hu" country="Taiwan" determinerName="Det. By F. S. Hu" elevation="1470" latitude="24.0191" location="Guandao Mt." longLatPrecision="7" longitude="121.0108" pageId="2" pageNumber="287" specimenCount="1" specimenCount-male="1" stateProvince="Nantou">
<collectingCountry id="F330B1A4B109FFECFE81FE975AE0E98D" box="[379,486,370,394]" name="Taiwan" pageId="2" pageNumber="287">TAIWAN</collectingCountry>
:
<specimenCount id="9D213ABDB109FFECFE15FE97593FE98D" box="[495,569,370,394]" pageId="2" pageNumber="287" type="male">1 male</specimenCount>
,
<collectingRegion id="49E33FD6B109FFECFDB9FE975994E98E" box="[579,658,370,393]" country="Taiwan" name="Nantou" pageId="2" pageNumber="287">Nantou</collectingRegion>
Co.,
<location id="8EF8A7EFB109FFECFD31FE97585EE98E" LSID="urn:lsid:plazi:treatment:038E4022B109FFEDFF6DFEA95A42E8D2:8EF8A7EFB109FFECFD31FE97585EE98E" box="[715,856,370,394]" country="Taiwan" latitude="24.0191" longLatPrecision="7" longitude="121.0108" name="Guandao Mt." pageId="2" pageNumber="287" stateProvince="Nantou">Guandao Mt.</location>
[
<emphasis id="B9532D26B109FFECFC9CFE8A58B0E982" box="[870,950,367,389]" italics="true" pageId="2" pageNumber="287">
<collectionCode id="ED3669F1B109FFECFC9CFE8A58B0E982" box="[870,950,367,389]" pageId="2" pageNumber="287">ḆBƜ</collectionCode>
</emphasis>
],
<collectorName id="26D294E2B109FFECFC36FE975F33E98E" box="[972,1077,370,394]" pageId="2" pageNumber="287">Renai To.</collectorName>
,
<geoCoordinate id="EE1397F3B109FFECFBC5FE975F9EE98D" box="[1087,1176,370,394]" degrees="24.0191" direction="north" orientation="latitude" pageId="2" pageNumber="287" precision="5" value="24.0191">24.0191</geoCoordinate>
,
<geoCoordinate id="EE1397F3B109FFECFB58FE975E00E98D" box="[1186,1286,370,394]" degrees="121.0108" direction="east" orientation="longitude" pageId="2" pageNumber="287" precision="5" value="121.0108">121.0108</geoCoordinate>
(alt.
<quantity id="4CDF5CD1B109FFECFAC6FE975E96E98D" box="[1340,1424,370,394]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="287" unit="m" value="1470.0">
<elevation id="000A1607B109FFECFAC6FE975E96E98D" box="[1340,1424,370,394]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="287" unit="m" value="1470.0">1470 m</elevation>
</quantity>
),
<date id="FF99D7F4B109FFECFF6DFE735A19E9A9" box="[151,287,406,430]" pageId="2" pageNumber="287" value="2019-03-23">
<collectingDate id="EFDD2E1CB109FFECFF6DFE735A19E9A9" box="[151,287,406,430]" pageId="2" pageNumber="287" value="2019-03-23">23III2019</collectingDate>
</date>
, leg.
<collectorName id="26D294E2B109FFECFEA0FE735AB4E9AA" box="[346,434,406,430]" pageId="2" pageNumber="287">F. S. Hu</collectorName>
(in dead wood) /
<taxonomicName id="4C278AB7B109FFECFD89FE735F20E9A9" authority="(Sharp, 1889)" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[627,1062,406,430]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B9532D26B109FFECFD89FE73588EE9A9" box="[627,904,406,430]" italics="true" pageId="2" pageNumber="287">Quwatanabius flavicornis</emphasis>
(Sharp, 1889)
</taxonomicName>
,
<determinerName id="7E441DF0B109FFECFBC8FE735FEEE9AA" box="[1074,1256,406,430]" pageId="2" pageNumber="287">
<collectorName id="26D294E2B109FFECFBC8FE735FEEE9AA" box="[1074,1256,406,430]" pageId="2" pageNumber="287">Det. By F. S. Hu</collectorName>
</determinerName>
</materialsCitation>
, 2019;
<materialsCitation id="3B4FFB69B109FFECFAC2FE735894E9D5" ID-GBIF-Occurrence="2571176114" collectingDate="2019-11-01" collectionCode="BBM" collectorName="F. S. Hu" country="Taiwan" determinerName="Det. By F. S. Hu" elevation="1470" latitude="24.0191" location="Guandao Mt." longLatPrecision="7" longitude="121.0108" pageId="2" pageNumber="287" specimenCount="1" specimenCount-female="1" stateProvince="Nantou">
<specimenCount id="9D213ABDB109FFECFAC2FE735E9EE9A9" box="[1336,1432,406,430]" pageId="2" pageNumber="287" type="female">1 female</specimenCount>
, same location,
<date id="FF99D7F4B109FFECFEC2FE5F5AC7E9D5" box="[312,449,442,466]" pageId="2" pageNumber="287" value="2019-11-01">
<collectingDate id="EFDD2E1CB109FFECFEC2FE5F5AC7E9D5" box="[312,449,442,466]" pageId="2" pageNumber="287" value="2019-11-01">01XI2019</collectingDate>
</date>
, leg.
<collectorName id="26D294E2B109FFECFE00FE5F5957E9D6" box="[506,593,442,466]" pageId="2" pageNumber="287">F. S. Hu</collectorName>
(sifting pieces of dead wood)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33DA2BFB109FFECFF3DFE3B5FDCE9F1" box="[199,1242,478,502]" pageId="2" pageNumber="287" type="description">
<paragraph id="8B98F134B109FFECFF3DFE3B5FDCE9F1" blockId="2.[151,1437,332,754]" box="[199,1242,478,502]" pageId="2" pageNumber="287">
<emphasis id="B9532D26B109FFECFF3DFE3B5A54E9F1" bold="true" box="[199,338,478,502]" pageId="2" pageNumber="287">Description.</emphasis>
See
<bibRefCitation id="EFB68CC5B109FFECFE7CFE3B5934E9F1" author="Smetana, A." box="[390,562,478,502]" pageId="2" pageNumber="287" pagination="1 - 145" refId="ref1870" refString="Smetana, A. (1995) Revision of the tribes Quediini and Tanygnathinini. Part III. Taiwan. (Coleoptera: Staphylinidae). National Museum of Natural Science, Taichung city, Special Publication, 6, 1 - 145." type="journal article" year="1995">Smetana (1995)</bibRefCitation>
; for illustration of internal sac of aedeagus see
<bibRefCitation id="EFB68CC5B109FFECFBD3FE3B5FD0E9F1" author="Smetana, A." box="[1065,1238,478,502]" pageId="2" pageNumber="287" pagination="271 - 274" refId="ref1962" refString="Smetana, A. (2002) Quwatanabius, a new genus of East Palaearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Special Bulletin of the Japanese Society of Coleopterology, 5, 271 - 274." type="journal article" year="2002">Smetana (2002)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33DA2BFB109FFECFF3DFDE75E3BEAF5" pageId="2" pageNumber="287" type="diagnosis">
<paragraph id="8B98F134B109FFECFF3DFDE758B2EA66" blockId="2.[151,1437,332,754]" pageId="2" pageNumber="287">
<emphasis id="B9532D26B109FFECFF3DFDE75A3AEA1D" bold="true" box="[199,316,514,538]" pageId="2" pageNumber="287">Diagnosis.</emphasis>
<taxonomicName id="4C278AB7B109FFECFEB9FDE7595EEA1D" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[323,600,514,538]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B9532D26B109FFECFEB9FDE7595EEA1D" box="[323,600,514,538]" italics="true" pageId="2" pageNumber="287">Quwatanabius flavicornis</emphasis>
</taxonomicName>
(
<figureCitation id="131CEDB1B109FFECFD9DFDE759AEEA1D" box="[615,680,514,538]" captionStart="FIGURES 78" captionStartId="2.[151,258,1809,1831]" captionTargetBox="[163,1435,777,1782]" captionTargetId="figure@2.[151,1435,777,1782]" captionTargetPageId="2" captionText="FIGURES 78. Quwatanabius flavicornis 7. habitus, scale bar represents 1mm; 8. habitat at the location where the additional specimens of Q. flavicornis were collected." figureDoi="http://doi.org/10.5281/zenodo.3687736" httpUri="https://zenodo.org/record/3687736/files/figure.png" pageId="2" pageNumber="287">Fig. 7</figureCitation>
) is similar to
<taxonomicName id="4C278AB7B109FFECFCC4FDE75899EA1D" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[830,927,514,538]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="chiaw">
<emphasis id="B9532D26B109FFECFCC4FDE75857EA1E" box="[830,849,514,537]" italics="true" pageId="2" pageNumber="287">Q</emphasis>
.
<emphasis id="B9532D26B109FFECFCA5FDE75899EA1D" box="[863,927,514,538]" italics="true" pageId="2" pageNumber="287">chiaw</emphasis>
</taxonomicName>
but can be distinguished from it by the following characters: pronotum much darker; elytra relatively longer; pale apical seam of palisade fringe on tergite VII present; median lobe rather narrow at apex, internal sac of more complex structure.
</paragraph>
<paragraph id="8B98F134B109FFECFF3DFD8B5E3BEAF5" blockId="2.[151,1437,332,754]" pageId="2" pageNumber="287">
<emphasis id="B9532D26B109FFECFF3DFD8B5A43EA81" bold="true" box="[199,325,622,646]" pageId="2" pageNumber="287">Bionomics.</emphasis>
The specimens in this study were collected from a large, fallen, moist, strongly decomposed piece of dead wood in primary, evergreen forest (
<figureCitation id="131CEDB1B109FFECFDBFFD775982EAAD" box="[581,644,658,682]" captionStart="FIGURES 78" captionStartId="2.[151,258,1809,1831]" captionTargetBox="[163,1435,777,1782]" captionTargetId="figure@2.[151,1435,777,1782]" captionTargetPageId="2" captionText="FIGURES 78. Quwatanabius flavicornis 7. habitus, scale bar represents 1mm; 8. habitat at the location where the additional specimens of Q. flavicornis were collected." figureDoi="http://doi.org/10.5281/zenodo.3687736" httpUri="https://zenodo.org/record/3687736/files/figure.png" pageId="2" pageNumber="287">Fig. 8</figureCitation>
), similar to the habitat where
<taxonomicName id="4C278AB7B109FFECFC3BFD775FA5EAAD" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[961,1187,658,682]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="chiaw">
<emphasis id="B9532D26B109FFECFC3BFD775FA5EAAD" box="[961,1187,658,682]" italics="true" pageId="2" pageNumber="287">Quwatanabius chiaw</emphasis>
</taxonomicName>
was collected. The two species, however, have been collected from different mountains, and most specimens of
<taxonomicName id="4C278AB7B109FFECFB9BFD535FF0EAC9" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[1121,1270,694,718]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B9532D26B109FFECFB9BFD535FF0EAC9" box="[1121,1270,694,718]" italics="true" pageId="2" pageNumber="287">Q. flavicornis</emphasis>
</taxonomicName>
were collected from lower altitudes (approximately
<quantity id="4CDF5CD1B109FFECFDDAFD3F5972EAF5" box="[544,628,730,754]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="287" unit="m" value="1500.0">1500 m</quantity>
), compared to the higher altitudes where
<taxonomicName id="4C278AB7B109FFECFBD7FD3F5F8BEAF5" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[1069,1165,730,754]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="chiaw">
<emphasis id="B9532D26B109FFECFBD7FD3F5F8BEAF5" box="[1069,1165,730,754]" italics="true" pageId="2" pageNumber="287">Q. chiaw</emphasis>
</taxonomicName>
has been found.
</paragraph>
</subSubSection>
<caption id="DF58A1BCB109FFECFF6DF8F45900EF4C" ID-DOI="http://doi.org/10.5281/zenodo.3687736" ID-Zenodo-Dep="3687736" httpUri="https://zenodo.org/record/3687736/files/figure.png" pageId="2" pageNumber="287" startId="2.[151,258,1809,1831]" targetBox="[163,1435,777,1782]" targetPageId="2">
<paragraph id="8B98F134B109FFECFF6DF8F45900EF4C" blockId="2.[151,1436,1809,1867]" pageId="2" pageNumber="287">
<emphasis id="B9532D26B109FFECFF6DF8F45A34EF21" bold="true" box="[151,306,1809,1831]" pageId="2" pageNumber="287">FIGURES 78.</emphasis>
<taxonomicName id="4C278AB7B109FFECFEC3F8F45935EF21" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[313,563,1809,1830]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B9532D26B109FFECFEC3F8F45935EF21" box="[313,563,1809,1830]" italics="true" pageId="2" pageNumber="287">Quwatanabius flavicornis</emphasis>
</taxonomicName>
7. habitus, scale bar represents 1mm; 8. habitat at the location where the additional specimens of
<taxonomicName id="4C278AB7B109FFECFF10F8D05A68EF4D" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[234,366,1845,1866]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="B9532D26B109FFECFF10F8D05A68EF4D" box="[234,366,1845,1866]" italics="true" pageId="2" pageNumber="287">Q. flavicornis</emphasis>
</taxonomicName>
were collected.
</paragraph>
</caption>
<subSubSection id="C33DA2BFB109FFECFF3DF89E5989EFDC" pageId="2" pageNumber="287" type="discussion">
<paragraph id="8B98F134B109FFECFF3DF89E5989EFDC" blockId="2.[151,1436,1915,2012]" pageId="2" pageNumber="287">
<emphasis id="B9532D26B109FFECFF3DF89E5A4FEF94" bold="true" box="[199,329,1915,1939]" pageId="2" pageNumber="287">Comments.</emphasis>
The bionomics of the Chinese species remains unknown, except
<taxonomicName id="4C278AB7B109FFECFBE0F8995FB2EF94" authorityName="Hu, Li &amp; Zhao" authorityYear="2013" box="[1050,1204,1915,1939]" class="Insecta" family="Staphylinidae" genus="Quwatanabius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="287" phylum="Arthropoda" rank="species" species="dayaoensis">
<emphasis id="B9532D26B109FFECFBE0F8995F2BEF94" box="[1050,1069,1916,1939]" italics="true" pageId="2" pageNumber="287">Q</emphasis>
.
<emphasis id="B9532D26B109FFECFBC7F89E5FB2EF94" box="[1085,1204,1915,1939]" italics="true" pageId="2" pageNumber="287">dayaoensis</emphasis>
</taxonomicName>
which we know was collected from dead wood (
<bibRefCitation id="EFB68CC5B109FFECFE47F87A595BEFB0" author="Hu, J. - Y. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[445,605,1951,1975]" pageId="2" pageNumber="287" pagination="297 - 299" refId="ref1802" refString="Hu, J. - Y., Li, L. - Z. &amp; Zhao, M. - J. (2013) A new species of Quwatanabius Smetana (Coleoptera, Staphylinidae, Staphylininae) from Guangxi, South China. Zootaxa, 3646 (3), 297 - 299. https: // doi. org / 10.11646 / zootaxa. 3646.3.8" type="journal article" year="2013">
Hu
<emphasis id="B9532D26B109FFECFE1CF845591AEFB0" box="[486,540,1951,1975]" italics="true" pageId="2" pageNumber="287">et al.</emphasis>
2013
</bibRefCitation>
). Since both species from
<collectingCountry id="F330B1A4B109FFECFC8CF87A58C2EFB0" box="[886,964,1951,1975]" name="Taiwan" pageId="2" pageNumber="287">Taiwan</collectingCountry>
were collected from dead wood as well, we assume that this genus may be saproxylophilic.
</paragraph>
</subSubSection>
<subSubSection id="C33DA2BFB108FFEDFF3DFF7C5A42E8D2" pageId="3" pageNumber="288" type="distribution">
<paragraph id="8B98F134B108FFEDFF3DFF7C5A42E8D2" blockId="3.[151,1437,153,213]" pageId="3" pageNumber="288">
<emphasis id="B9532D26B108FFEDFF3DFF7C5A5CE8B6" bold="true" box="[199,346,153,177]" pageId="3" pageNumber="288">Distribution.</emphasis>
The species is widespread in
<collectingCountry id="F330B1A4B108FFEDFD60FF7F59D1E8B6" box="[666,727,154,177]" name="Japan" pageId="3" pageNumber="288">Japan</collectingCountry>
(Honshu, Kyushu, and Shikoku) and
<collectingCountry id="F330B1A4B108FFEDFB96FF7C5FBCE8B6" box="[1132,1210,153,177]" name="Taiwan" pageId="3" pageNumber="288">Taiwan</collectingCountry>
(
<collectingRegion id="49E33FD6B108FFEDFB33FF7C5E69E8B6" box="[1225,1391,153,177]" country="Taiwan" name="Nantou" pageId="3" pageNumber="288">Nantou County</collectingRegion>
and
<collectingRegion id="49E33FD6B108FFEDFF6DFF585A3EE8D2" box="[151,312,189,213]" country="Taiwan" name="Chiayi" pageId="3" pageNumber="288">Chiayi County</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>