treatments-xml/data/03/8E/02/038E021DFFA6FFBE4FA9FC6BFB2DFF3D.xml
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<document id="0D37D189611CD3EF31AEAAFEC6C7DBA4" ID-CLB-Dataset="29419" ID-DOI="10.11646/zootaxa.4482.3.1" ID-GBIF-Dataset="5916d93e-11d4-49c5-8085-8bcd37d3739b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440674" ID-ZooBank="10258391-162F-4C7D-AA5E-1A03A4F3FE19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538374587817" checkinUser="plazi" docAuthor="Pal, Saunak, Vijayakumar, S. P., Shanker, Kartik, Jayarajan, Aditi &amp; Deepak, V." docDate="2018" docId="038E021DFFA6FFBE4FA9FC6BFB2DFF3D" docLanguage="en" docName="zootaxa.4482.3.1.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calotes grandisquamis Gunther 1875" docType="treatment" docVersion="7" lastPageNumber="424" masterDocId="FFB77A65FFB0FFA64F3EFF9CFFEFFFE8" masterDocTitle="A systematic revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds two genera and reveals two new species" masterLastPageNumber="450" masterPageNumber="401" pageNumber="423" updateTime="1698559486032" updateUser="ExternalLinkService">
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<mods:title id="D9A01B354FF39B4ABC947C5D054FBBB1">A systematic revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds two genera and reveals two new species</mods:title>
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<mods:namePart id="8751B2FF119AE9A68ABD1C0594C677CF">Pal, Saunak</mods:namePart>
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<mods:namePart id="1355606CAB4ECAE48FF4B851E366F4C2">Vijayakumar, S. P.</mods:namePart>
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<mods:namePart id="C2A6C1F6A0B244B6C5761E7FEA4728DB">Shanker, Kartik</mods:namePart>
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<mods:namePart id="7E4E02DBCD4742A93EBF6E2B4C3E2CAC">Jayarajan, Aditi</mods:namePart>
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<mods:namePart id="1A8DE40038A40ED9794F8E59F3B64CDD">Deepak, V.</mods:namePart>
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<mods:title id="7B07DDCF8FAD0B9E59794B71A58C6CFE">Zootaxa</mods:title>
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<mods:date id="BB828F13A7EBBAC2BCAD7242B0D10E0F">2018</mods:date>
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<mods:number id="E6EDC99887203B28BE90A08590A288B4">4482</mods:number>
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<treatment id="038E021DFFA6FFBE4FA9FC6BFB2DFF3D" ID-DOI="http://doi.org/10.5281/zenodo.5996686" ID-GBIF-Taxon="148421706" ID-Zenodo-Dep="5996686" LSID="urn:lsid:plazi:treatment:038E021DFFA6FFBE4FA9FC6BFB2DFF3D" httpUri="http://treatment.plazi.org/id/038E021DFFA6FFBE4FA9FC6BFB2DFF3D" lastPageId="24" lastPageNumber="424" pageId="22" pageNumber="423">
<subSubSection id="C33DE080FFA6FFB04FA9FC6BFCA2FBBC" pageId="22" pageNumber="423" type="nomenclature">
<paragraph id="8B98B30BFFA6FFB04FA9FC6BFD8BFBFA" blockId="22.[151,612,1015,1042]" box="[151,612,1015,1042]" pageId="22" pageNumber="423">
<heading id="D0D00467FFA6FFB04FA9FC6BFD8BFBFA" bold="true" box="[151,612,1015,1042]" fontSize="11" level="1" pageId="22" pageNumber="423" reason="1">
<taxonomicName id="4C27C888FFA6FFB04FA9FC6BFD8BFBFA" authority="Gunther, 1875" authorityName="Gunther" authorityYear="1875" box="[151,612,1015,1042]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="grandisquamis">
<emphasis id="B9536F19FFA6FFB04FA9FC6BFD8BFBFA" bold="true" box="[151,612,1015,1042]" pageId="22" pageNumber="423">
<emphasis id="B9536F19FFA6FFB04FA9FC6BFE44FBF9" bold="true" box="[151,427,1015,1041]" italics="true" pageId="22" pageNumber="423">Calotes grandisquamis</emphasis>
<bibRefCitation id="EFB6CEFAFFA6FFB04E8CFC64FD8BFBFA" box="[434,612,1016,1042]" pageId="22" pageNumber="423" refId="ref35549" refString="Gunther, A. (1875) Second report on collections of Indian Reptiles obtained by the British Museum. Proceedings of the Zoological Society of London, 1875, 224 - 234." type="journal article">Günther, 1875</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="8B98B30BFFA6FFB04FA9FBA2FCA2FBBC" blockId="22.[151,845,1086,1108]" box="[151,845,1086,1108]" pageId="22" pageNumber="423">
<emphasis id="B9536F19FFA6FFB04FA9FBA2FE77FBBC" box="[151,408,1086,1108]" italics="true" pageId="22" pageNumber="423">
<taxonomicName id="4C27C888FFA6FFB04FA9FBA2FF0BFBBC" box="[151,228,1086,1108]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="genus">Calotes</taxonomicName>
grandisquamis—
</emphasis>
<bibRefCitation id="EFB6CEFAFFA6FFB04EA9FBA2FDC3FBBC" box="[407,556,1086,1108]" pageId="22" pageNumber="423" refId="ref35549" refString="Gunther, A. (1875) Second report on collections of Indian Reptiles obtained by the British Museum. Proceedings of the Zoological Society of London, 1875, 224 - 234." type="journal article">Günther, 1875</bibRefCitation>
. Proc. Zool. Soc. 1875: 226.
</paragraph>
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<paragraph id="8B98B30BFFA6FFB04FA9FB1DFB03FB71" blockId="22.[151,1436,1153,2006]" box="[151,1260,1153,1178]" pageId="22" pageNumber="423">
<emphasis id="B9536F19FFA6FFB04FA9FB1DFE7FFB72" bold="true" box="[151,400,1153,1178]" pageId="22" pageNumber="423">Original description.</emphasis>
<bibRefCitation id="EFB6CEFAFFA6FFB04EA9FB1DFDD5FB72" box="[407,570,1153,1178]" pageId="22" pageNumber="423" refId="ref35549" refString="Gunther, A. (1875) Second report on collections of Indian Reptiles obtained by the British Museum. Proceedings of the Zoological Society of London, 1875, 224 - 234." type="journal article">Günther, 1875</bibRefCitation>
. Proceedings of the Zoological Society, 1875: 226, Plate xxx.
</paragraph>
<paragraph id="8B98B30BFFA6FFB04FF9FB39FCD5FAED" blockId="22.[151,1436,1153,2006]" pageId="22" pageNumber="423">
<emphasis id="B9536F19FFA6FFB04FF9FB39FE39FB56" bold="true" box="[199,470,1189,1214]" pageId="22" pageNumber="423">Taxonomic comments.</emphasis>
<taxonomicName id="4C27C888FFA6FFB04EDEFB39FD07FB55" authorityName="Gunther" authorityYear="1875" box="[480,744,1189,1213]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="grandisquamis">
<emphasis id="B9536F19FFA6FFB04EDEFB39FD07FB55" box="[480,744,1189,1213]" italics="true" pageId="22" pageNumber="423">Calotes grandisquamis</emphasis>
</taxonomicName>
was described based on specimens collected from foot of Canoot Ghat, near Manantoddy, Bramagherry hills. This large bodied, green
<taxonomicName id="4C27C888FFA6FFB04CC3FB55FBBCFB09" box="[1021,1107,1225,1249]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="genus">
<emphasis id="B9536F19FFA6FFB04CC3FB55FBBCFB09" box="[1021,1107,1225,1249]" italics="true" pageId="22" pageNumber="423">Calotes</emphasis>
</taxonomicName>
is a well-defined species and can be distinguished from the closely related
<taxonomicName id="4C27C888FFA6FFB04DADFB72FCDDFAED" authorityName=", Jerdon" authorityYear="1853" box="[659,818,1261,1285]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="nemoricola">
<emphasis id="B9536F19FFA6FFB04DADFB72FCDDFAED" box="[659,818,1261,1285]" italics="true" pageId="22" pageNumber="423">C. nemoricola</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B98B30BFFA6FFB04FF9FA8DFBC5FAA6" blockId="22.[151,1436,1153,2006]" pageId="22" pageNumber="423">
<materialsCitation id="3B4FB956FFA6FFB04FF9FA8DFBC5FAA6" ID-GBIF-Occurrence="1913889692" collectionCode="BMNH" county="Canoot Ghat" location="Bramagherry" municipality="Manantoddy" pageId="22" pageNumber="423" specimenCode="BMNH 1946.8" specimenCount="1" stateProvince="Kerala" typeStatus="syntype">
<emphasis id="B9536F19FFA6FFB04FF9FA8DFED6FAC2" bold="true" box="[199,313,1297,1322]" pageId="22" pageNumber="423">
<typeStatus id="549C0DA9FFA6FFB04FF9FA8DFEDAFAC2" box="[199,309,1297,1322]" pageId="22" pageNumber="423" type="syntype">Syntypes</typeStatus>
.
</emphasis>
<specimenCode id="DB811B70FFA6FFB04E75FA8EFE10FAC2" box="[331,511,1297,1322]" collectionCode="BMNH" pageId="22" pageNumber="423">BMNH 1946.8</specimenCode>
.11.44 to 47, collected from foot of the
<collectingCounty id="62F9CB87FFA6FFB04B2BFA8EFB5EFAC2" box="[1045,1201,1297,1322]" pageId="22" pageNumber="423">Canoot Ghat</collectingCounty>
, near
<collectingMunicipality id="6BFC2971FFA6FFB04A37FA8DFA77FAC2" box="[1289,1432,1297,1322]" pageId="22" pageNumber="423">Manantoddy</collectingMunicipality>
,
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hills (presently near
<location id="8EF8E5D0FFA6FFB04D1EFAA9FD29FAA6" LSID="urn:lsid:plazi:treatment:038E021DFFA6FFBE4FA9FC6BFB2DFF3D:8EF8E5D0FFA6FFB04D1EFAA9FD29FAA6" box="[544,710,1333,1358]" county="Canoot Ghat" municipality="Manantoddy" name="Mananthawadi" pageId="22" pageNumber="423" stateProvince="Kerala">Mananthawadi</location>
in the
<location id="8EF8E5D0FFA6FFB04C2DFAA9FC7CFAA6" LSID="urn:lsid:plazi:treatment:038E021DFFA6FFBE4FA9FC6BFB2DFF3D:8EF8E5D0FFA6FFB04C2DFAA9FC7CFAA6" box="[787,915,1333,1358]" county="Canoot Ghat" municipality="Manantoddy" name="Brahmagiri" pageId="22" pageNumber="423" stateProvince="Kerala">Brahmagiri</location>
hills,
<collectingRegion id="49E37DE9FFA6FFB04CE6FAA9FBC9FAA6" box="[984,1062,1333,1358]" country="India" name="Kerala" pageId="22" pageNumber="423">Kerala</collectingRegion>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFA6FFB04FF9FAC5FDE2F9CE" pageId="22" pageNumber="423" type="materials_examined">
<paragraph id="8B98B30BFFA6FFB04FF9FAC5FDE2F9CE" blockId="22.[151,1436,1153,2006]" pageId="22" pageNumber="423">
<materialsCitation id="3B4FB956FFA6FFB04FF9FAC5FF0AFA7E" ID-GBIF-Occurrence="1913889695" collectionCode="CESL" location="Peppara Wildlife Sanctuary" municipality="Chemunji" pageId="22" pageNumber="423" specimenCount="35" specimenCount-adult="35" stateProvince="Kerala">
<emphasis id="B9536F19FFA6FFB04FF9FAC5FE59FA9A" bold="true" box="[199,438,1369,1394]" pageId="22" pageNumber="423">Material examined.</emphasis>
<collectionCode id="ED362BCEFFA6FFB04EFCFAC6FDE9FA9A" box="[450,518,1370,1394]" pageId="22" pageNumber="423">CESL</collectionCode>
0
<specimenCount id="9D217882FFA6FFB04D1FFAC5FD6EFA9A" box="[545,641,1369,1394]" pageId="22" pageNumber="423" type="adult">35 adult</specimenCount>
female collected from near
<collectingMunicipality id="6BFC2971FFA6FFB04CE8FAC5FBA8FA9A" box="[982,1095,1369,1394]" pageId="22" pageNumber="423">Chemunji</collectingMunicipality>
,
<location id="8EF8E5D0FFA6FFB04B66FAC6FA77FA9A" LSID="urn:lsid:plazi:treatment:038E021DFFA6FFBE4FA9FC6BFB2DFF3D:8EF8E5D0FFA6FFB04B66FAC6FA77FA9A" box="[1112,1432,1369,1394]" municipality="Chemunji" name="Peppara Wildlife Sanctuary" pageId="22" pageNumber="423" stateProvince="Kerala">Peppara Wildlife Sanctuary</location>
,
<collectingRegion id="49E37DE9FFA6FFB04FA9FAE1FF0AFA7E" box="[151,229,1405,1430]" country="India" name="Kerala" pageId="22" pageNumber="423">Kerala</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B4FB956FFA6FFB04FCDFAE2FBA1FA7D" ID-GBIF-Occurrence="1913889705" box="[243,1102,1405,1430]" collectionCode="CESL" location="Periyar" municipality="Upper Manalar" pageId="22" pageNumber="423" specimenCount="120" specimenCount-adult="120" stateProvince="Kerala">
<collectionCode id="ED362BCEFFA6FFB04FCDFAE2FED8FA7E" box="[243,311,1406,1430]" pageId="22" pageNumber="423">CESL</collectionCode>
<specimenCount id="9D217882FFA6FFB04E7FFAE2FE42FA7E" box="[321,429,1405,1430]" pageId="22" pageNumber="423" type="adult">120 adult</specimenCount>
male collected from
<collectingMunicipality id="6BFC2971FFA6FFB04D98FAE2FCBAFA7E" box="[678,853,1405,1430]" pageId="22" pageNumber="423">Upper Manalar</collectingMunicipality>
,
<location id="8EF8E5D0FFA6FFB04C5CFAE1FC5CFA7E" LSID="urn:lsid:plazi:treatment:038E021DFFA6FFBE4FA9FC6BFB2DFF3D:8EF8E5D0FFA6FFB04C5CFAE1FC5CFA7E" box="[866,947,1405,1430]" municipality="Upper Manalar" name="Periyar" pageId="22" pageNumber="423" stateProvince="Kerala">Periyar</location>
tiger reserve
</materialsCitation>
,
<materialsCitation id="3B4FB956FFA6FFB04B62FAE1FC8DFA51" ID-GBIF-Occurrence="1913889698" collectionCode="CESL" location="Kerala" pageId="22" pageNumber="423" specimenCount="191" specimenCount-adult="191" stateProvince="Kerala">
<collectingRegion id="49E37DE9FFA6FFB04B62FAE1FB48FA7E" box="[1116,1191,1405,1430]" country="India" name="Kerala" pageId="22" pageNumber="423">Kerala</collectingRegion>
and
<collectionCode id="ED362BCEFFA6FFB04BDCFAE2FAC9FA7E" box="[1250,1318,1406,1430]" pageId="22" pageNumber="423">CESL</collectionCode>
<specimenCount id="9D217882FFA6FFB04A0EFAE2FA73FA7E" box="[1328,1436,1405,1430]" pageId="22" pageNumber="423" type="adult">191 adult</specimenCount>
male collected from near Pulmeedu, Periyar tiger reserve
</materialsCitation>
,
<materialsCitation id="3B4FB956FFA6FFB04C49FA3DFADDFA52" ID-GBIF-Occurrence="1913889710" box="[887,1330,1441,1466]" collectionCode="BMNH" location="Kerala" pageId="22" pageNumber="423" specimenCode="BMNH 1946.8" specimenCount="1" stateProvince="Kerala">
<collectingRegion id="49E37DE9FFA6FFB04C49FA3DFC2AFA52" box="[887,965,1441,1466]" country="India" name="Kerala" pageId="22" pageNumber="423">Kerala</collectingRegion>
;
<specimenCode id="DB811B70FFA6FFB04CE5FA3EFB61FA52" box="[987,1166,1441,1466]" collectionCode="BMNH" pageId="22" pageNumber="423">BMNH 1946.8</specimenCode>
.11.44 female
</materialsCitation>
, BMNH 1946.8.11.45 juvenile, BMNH 1946.8.11.46 juvenile, BMNH 1946.8.11.47 male all deposited by R.H. Beddome collected from the foothills of “Coonoor Ghat. Details of collection locality, specimen voucher and GenBank accession number in Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFA6FFB14FF9F9ADFC16FE38" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423" type="diagnosis">
<paragraph id="8B98B30BFFA6FFB04FF9F9ADFE3DF882" blockId="22.[151,1436,1153,2006]" pageId="22" pageNumber="423">
<emphasis id="B9536F19FFA6FFB04FF9F9ADFDE5F9A2" bold="true" box="[199,522,1585,1610]" pageId="22" pageNumber="423">Diagnosis and comparison.</emphasis>
A large sized
<taxonomicName id="4C27C888FFA6FFB04D8EF9ADFCEAF9A1" box="[688,773,1585,1609]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="genus">
<emphasis id="B9536F19FFA6FFB04D8EF9ADFCEAF9A1" box="[688,773,1585,1609]" italics="true" pageId="22" pageNumber="423">Calotes</emphasis>
</taxonomicName>
(SVL up to
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) characterized by the posterodorsal orientation of lateral scales; antehumeral fold absent; 2735 midbody scale rows; nuchal and dorsal crest well developed, continuous, nuchal crest composed of long, curved spines, dorsal spines smaller, gradually reducing in length and reaches till above the tail base; row of 34 compressed supratympanic spines; postorbital spine absent; a crescent-shaped patch of granular scales in front of the shoulder present; dorsal and lateral scales large, smooth, subtriangular, ventral scales keeled, mucronate; paired postmentals, first pair separated by a single median scale; 2225 subdigital lamellae under fourth finger, 2730 subdigital lamellae under fourth toe; 10 supralabials and 910 infralabials; green above, often with 23 broad black transverse bars on the lateral side, lateral side of head darker, blackish, ventral pale green.
</paragraph>
<paragraph id="8B98B30BFFA6FFB14FF9F8E9FC16FE38" blockId="22.[151,1436,1153,2006]" lastBlockId="23.[151,1437,151,2013]" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">
<taxonomicName id="4C27C888FFA6FFB04FF9F8E9FE26F865" authorityName="Gunther" authorityYear="1875" box="[199,457,1909,1933]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="grandisquamis">
<emphasis id="B9536F19FFA6FFB04FF9F8E9FE26F865" box="[199,457,1909,1933]" italics="true" pageId="22" pageNumber="423">Calotes grandisquamis</emphasis>
</taxonomicName>
can be distinguished from members of Smiths
<taxonomicName id="4C27C888FFA6FFB04CE3F8EAFB80F865" authorityName="Daudin" authorityYear="1802" box="[989,1135,1909,1933]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="B9536F19FFA6FFB04CE3F8EAFB80F865" box="[989,1135,1909,1933]" italics="true" pageId="22" pageNumber="423">C. versicolor</emphasis>
</taxonomicName>
group by a combination of the following characters: larger body size: adult SVL 110.0
<quantity id="4CDF1EEEFFA6FFB04C06F805FC45F859" box="[824,938,1945,1970]" metricMagnitude="-1" metricUnit="m" metricValue="1.365" pageId="22" pageNumber="423" unit="mm" value="136.5">136.5 mm</quantity>
, n=5 (vs.
<emphasis id="B9536F19FFA6FFB04B28F806FB43F859" box="[1046,1196,1945,1969]" italics="true" pageId="22" pageNumber="423">
<taxonomicName id="4C27C888FFA6FFB04B28F806FB48F859" authorityName="Daudin" authorityYear="1802" box="[1046,1191,1945,1969]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="versicolor">C. versicolor</taxonomicName>
,
</emphasis>
adult SVL
<quantity id="4CDF1EEEFFA6FFB04A12F806FF2AF83D" metricMagnitude="-2" metricUnit="m" metricValue="8.66" metricValueMax="9.94" metricValueMin="7.38" pageId="22" pageNumber="423" unit="mm" value="86.6" valueMax="99.4" valueMin="73.8">73.899.4 mm</quantity>
, n=9 and
<emphasis id="B9536F19FFA6FFB04E06F822FE5DF83D" box="[312,434,1981,2005]" italics="true" pageId="22" pageNumber="423">
<taxonomicName id="4C27C888FFA6FFB04E06F822FE42F83D" authorityName="calotes (Linnaeus" authorityYear="1758" box="[312,429,1981,2005]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="calotes">C. calotes</taxonomicName>
,
</emphasis>
adult SVL
<quantity id="4CDF1EEEFFA6FFB04D07F821FD0FF83D" box="[569,736,1981,2006]" metricMagnitude="-2" metricUnit="m" metricValue="8.52" metricValueMax="9.68" metricValueMin="7.359999999999999" pageId="22" pageNumber="423" unit="mm" value="85.19999999999999" valueMax="96.8" valueMin="73.6">73.696.8 mm</quantity>
, n=3); 2735 midbody scale rows (
<quantity id="4CDF1EEEFFA6FFB04B49F821FB31F83E" box="[1143,1246,1981,2006]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.89" metricValueMin="7.62" pageId="22" pageNumber="423" unit="in" value="32.5" valueMax="35.0" valueMin="30.0">3035 in</quantity>
<taxonomicName id="4C27C888FFA6FFB04BD8F822FAB7F83D" authorityName="calotes (Linnaeus" authorityYear="1758" box="[1254,1368,1981,2005]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="423" phylum="Chordata" rank="species" species="calotes">
<emphasis id="B9536F19FFA6FFB04BD8F822FAB7F83D" box="[1254,1368,1981,2005]" italics="true" pageId="22" pageNumber="423">C. calotes</emphasis>
</taxonomicName>
, n=3;
<quantity id="4CDF1EEEFFA7FFB14FA9FF04FF15FF58" box="[151,250,151,176]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.1176" metricValueMin="0.9652" pageId="23" pageNumber="424" unit="in" value="41.0" valueMax="44.0" valueMin="38.0">3844 in</quantity>
<emphasis id="B9536F19FFA7FFB14E3EFF04FE74FF58" box="[256,411,152,176]" italics="true" pageId="23" pageNumber="424">
<taxonomicName id="4C27C888FFA7FFB14E3EFF04FE7BFF58" authorityName="Daudin" authorityYear="1802" box="[256,404,152,176]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="versicolor">C. versicolor</taxonomicName>
;
</emphasis>
<quantity id="4CDF1EEEFFA7FFB14E9CFF0BFDEBFF58" box="[418,516,151,176]" metricMagnitude="0" metricUnit="m" metricValue="1.5367000000000002" metricValueMax="1.6002000000000003" metricValueMin="1.4731999999999998" pageId="23" pageNumber="424" unit="in" value="60.5" valueMax="63.0" valueMin="58.0">5863 in</quantity>
<taxonomicName id="4C27C888FFA7FFB14D35FF04FD81FF47" authorityName="Gray" authorityYear="1845" box="[523,622,152,175]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="maria">
<emphasis id="B9536F19FFA7FFB14D35FF04FD81FF47" box="[523,622,152,175]" italics="true" pageId="23" pageNumber="424">C. maria</emphasis>
</taxonomicName>
;
<quantity id="4CDF1EEEFFA7FFB14D43FF0BFD30FF58" box="[637,735,151,176]" metricMagnitude="0" metricUnit="m" metricValue="1.2953999999999999" metricValueMax="1.4478" metricValueMin="1.143" pageId="23" pageNumber="424" unit="in" value="51.0" valueMax="57.0" valueMin="45.0">4557 in</quantity>
<taxonomicName id="4C27C888FFA7FFB14DDBFF04FCB9FF58" authorityName="Gunther" authorityYear="1870" box="[741,854,152,176]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="jerdoni">
<emphasis id="B9536F19FFA7FFB14DDBFF04FCB9FF58" box="[741,854,152,176]" italics="true" pageId="23" pageNumber="424">C. jerdoni</emphasis>
</taxonomicName>
;
<quantity id="4CDF1EEEFFA7FFB14C5AFF0BFC28FF58" box="[868,967,151,176]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" metricValueMax="1.6509999999999998" metricValueMin="1.2446" pageId="23" pageNumber="424" unit="in" value="57.0" valueMax="65.0" valueMin="49.0">4965 in</quantity>
<taxonomicName id="4C27C888FFA7FFB14CF0FF04FBDEFF58" authorityName="Gray" authorityYear="1845" box="[974,1073,152,176]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="emma">
<emphasis id="B9536F19FFA7FFB14CF0FF04FBDEFF58" box="[974,1073,152,176]" italics="true" pageId="23" pageNumber="424">C. emma</emphasis>
</taxonomicName>
;
<quantity id="4CDF1EEEFFA7FFB14B01FF0BFB4DFF58" box="[1087,1186,151,176]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" metricValueMax="1.4731999999999998" metricValueMin="1.2192" pageId="23" pageNumber="424" unit="in" value="53.0" valueMax="58.0" valueMin="48.0">4858 in</quantity>
<taxonomicName id="4C27C888FFA7FFB14B97FF04FAD3FF58" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[1193,1340,152,176]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="mystaceus">
<emphasis id="B9536F19FFA7FFB14B97FF04FAD3FF58" box="[1193,1340,152,176]" italics="true" pageId="23" pageNumber="424">C. mystaceus</emphasis>
</taxonomicName>
and
<quantity id="4CDF1EEEFFA7FFB14A4CFF04FF3DFF3D" metricMagnitude="0" metricUnit="m" metricValue="1.3716" metricValueMax="1.524" metricValueMin="1.2192" pageId="23" pageNumber="424" unit="in" value="54.0" valueMax="60.0" valueMin="48.0">48 60 in</quantity>
<taxonomicName id="4C27C888FFA7FFB14FE2FF20FEADFF3C" baseAuthorityName="Hardwicke &amp; Gray" baseAuthorityYear="1827" box="[220,322,188,212]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="minor">
<emphasis id="B9536F19FFA7FFB14FE2FF20FEADFF3C" box="[220,322,188,212]" italics="true" pageId="23" pageNumber="424">C. minor</emphasis>
</taxonomicName>
); nuchal crest well developed with much longer spines than dorsal spines (vs. nuchal and dorsal crest well developed, composed of almost equal spines in
<taxonomicName id="4C27C888FFA7FFB14C7FFF7CFC38FF10" authorityName="Daudin" authorityYear="1802" box="[833,983,224,248]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="B9536F19FFA7FFB14C7FFF7CFC38FF10" box="[833,983,224,248]" italics="true" pageId="23" pageNumber="424">C. versicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C27C888FFA7FFB14B2AFF7CFB44FF10" authorityName="Dumeril &amp; Bibron" authorityYear="1837" box="[1044,1195,224,248]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="mystaceus">
<emphasis id="B9536F19FFA7FFB14B2AFF7CFB44FF10" box="[1044,1195,224,248]" italics="true" pageId="23" pageNumber="424">C. mystaceus</emphasis>
</taxonomicName>
; nuchal spines long, dorsal crest reduced in
<taxonomicName id="4C27C888FFA7FFB14E86FE98FDCCFEF4" authorityName="Gray" authorityYear="1845" box="[440,547,260,284]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="maria">
<emphasis id="B9536F19FFA7FFB14E86FE98FDCCFEF4" box="[440,547,260,284]" italics="true" pageId="23" pageNumber="424">C. maria</emphasis>
</taxonomicName>
); row of 34 compressed supratympanic spines (vs. two well separated supratympanic spines in
<taxonomicName id="4C27C888FFA7FFB14E8FFEB4FDAAFEA8" authorityName="Daudin" authorityYear="1802" box="[433,581,296,320]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="B9536F19FFA7FFB14E8FFEB4FDAAFEA8" box="[433,581,296,320]" italics="true" pageId="23" pageNumber="424">C. versicolor</emphasis>
</taxonomicName>
and 89 compressed spines in
<emphasis id="B9536F19FFA7FFB14C97FEB4FBCDFEA8" box="[937,1058,296,320]" italics="true" pageId="23" pageNumber="424">
<taxonomicName id="4C27C888FFA7FFB14C97FEB4FBF1FEA8" authorityName="calotes (Linnaeus" authorityYear="1758" box="[937,1054,296,320]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="calotes">C. calotes</taxonomicName>
,
</emphasis>
two parallel rows of compressed supratympanic scales in
<taxonomicName id="4C27C888FFA7FFB14E94FED0FDE1FE8C" authorityName="Gray" authorityYear="1845" box="[426,526,332,356]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="maria">
<emphasis id="B9536F19FFA7FFB14E94FED0FDE1FE8C" box="[426,526,332,356]" italics="true" pageId="23" pageNumber="424">C. maria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C27C888FFA7FFB14D7AFED0FD5AFE8C" authorityName="Gunther" authorityYear="1870" box="[580,693,332,356]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="jerdoni">
<emphasis id="B9536F19FFA7FFB14D7AFED0FD5AFE8C" box="[580,693,332,356]" italics="true" pageId="23" pageNumber="424">C. jerdoni</emphasis>
</taxonomicName>
; single well developed postorbital spine in
<taxonomicName id="4C27C888FFA7FFB14BA1FED0FAECFE8C" authorityName="Gray" authorityYear="1845" box="[1183,1283,332,356]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="emma">
<emphasis id="B9536F19FFA7FFB14BA1FED0FAECFE8C" box="[1183,1283,332,356]" italics="true" pageId="23" pageNumber="424">C. emma</emphasis>
</taxonomicName>
); presence of a crescent-shaped patch of granular scales in front of the shoulder (vs. absent in
<taxonomicName id="4C27C888FFA7FFB14B0DFEECFB29FE60" authorityName="Daudin" authorityYear="1802" box="[1075,1222,368,392]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="B9536F19FFA7FFB14B0DFEECFB29FE60" box="[1075,1222,368,392]" italics="true" pageId="23" pageNumber="424">C. versicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="4C27C888FFA7FFB14BE8FEECFAA7FE60" authorityName="calotes (Linnaeus" authorityYear="1758" box="[1238,1352,368,392]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="calotes">
<emphasis id="B9536F19FFA7FFB14BE8FEECFAA7FE60" box="[1238,1352,368,392]" italics="true" pageId="23" pageNumber="424">C. calotes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C27C888FFA7FFB14ABCFEECFF35FE44" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="maria">
<emphasis id="B9536F19FFA7FFB14ABCFEECFF35FE44" italics="true" pageId="23" pageNumber="424">C. maria</emphasis>
</taxonomicName>
) and lateral scales much larger than ventrals (vs. almost equal to the ventrals in
<taxonomicName id="4C27C888FFA7FFB14BE2FE08FABAFE44" authorityName="calotes (Linnaeus" authorityYear="1758" box="[1244,1365,404,428]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="calotes">
<emphasis id="B9536F19FFA7FFB14BE2FE08FABAFE44" box="[1244,1365,404,428]" italics="true" pageId="23" pageNumber="424">C. calotes</emphasis>
</taxonomicName>
). For distinguishing from
<emphasis id="B9536F19FFA7FFB14E44FE24FDCEFE38" box="[378,545,440,464]" italics="true" pageId="23" pageNumber="424">
<taxonomicName id="4C27C888FFA7FFB14E44FE24FDF2FE38" authorityName=", Jerdon" authorityYear="1853" box="[378,541,440,464]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="nemoricola">C. nemoricola</taxonomicName>
,
</emphasis>
see comparison section of
<emphasis id="B9536F19FFA7FFB14C6FFE24FC16FE38" box="[849,1017,440,464]" italics="true" pageId="23" pageNumber="424">
<taxonomicName id="4C27C888FFA7FFB14C6FFE24FC1AFE38" authorityName=", Jerdon" authorityYear="1853" box="[849,1013,440,464]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="nemoricola">C. nemoricola</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFA7FFB14FF9FE40FD16F955" pageId="23" pageNumber="424" type="description">
<paragraph id="8B98B30BFFA7FFB14FF9FE40FA73FC68" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="424">
<emphasis id="B9536F19FFA7FFB14FF9FE40FEB6FE1D" bold="true" box="[199,345,476,501]" pageId="23" pageNumber="424">Description.</emphasis>
Based on specimen CESL 191. A large sized adult male (SVL-
<quantity id="4CDF1EEEFFA7FFB14B03FE40FB59FE1C" box="[1085,1206,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.365" pageId="23" pageNumber="424" unit="mm" value="136.5">136.5 mm</quantity>
), morphometric and meristic data are summarised in Appendix 2 &amp; 3. General habitus compressed. Head moderately large (HL/SVL ratio 0.28), elongate (HW/HL ratio 0.60), maximum height slightly less than maximum width; snout pointed; rostral broader than high; nostrils in single nasal shield which is separated from rostral by two scales; mental shield narrower than rostral; two postmentals, first pair separated from each other by a single scale; genials smooth; gular sac well developed, composed of mostly smooth scales, weakly keeled only at the medial row, slightly smaller than genials; scales on top of snout smooth except median row, which is keeled; scales on top of head heterogenous in shape and size, keeled towards the edges; supraorbital scales weakly keeled; canthus-rostralis and supraciliary edge sharp; 3 supratympanic spines in a row, compressed; orbit diameter 71% of distance between anterior border of orbit and snout tip; tympanum exposed, its greatest diameter 41% of horizontal diameter of orbit; partially keeled, scales between tympanum and orbit smooth, enlarged, slightly smaller than the tympanum; posterior region of jaws distinctly swollen with large scales, equal to or slightly larger than tympanum; supralabials 10/10; infralabials 10/9.
</paragraph>
<paragraph id="8B98B30BFFA7FFB14FF9FC10FAD6FBDD" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="424">Nuchal and dorsal crest well developed, continuous; composed of 28 long, conical spines, the first being the smallest; dorsal crest reaching above the tail base gradually decreasing in length, continues till the end of tail as a serrated ridge; longest nuchal spine 107% of the orbit; 30 longitudinal scale rows around midbody; dorsal and lateral scales sub-triangular, smooth, oriented postero-dorsally; ventrals strongly keeled, mucronate, smaller than dorsal and lateral scales; a crescent-shaped patch of small, granular scales present in front of the shoulder.</paragraph>
<paragraph id="8B98B30BFFA7FFB14FF9FBA3FBD8FAE5" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="424">
Limbs moderate and covered with smooth scales, weakly keeled scales towards the end; scales under thighs weakly keeled; length of hindlimb ca. 70 % SVL; relative length of fingers 4&gt;3&gt;2&gt;5&gt;1, third and fourth finger almost equal; relative lengths of toes 4&gt;3&gt;5&gt;2&gt;1; fourth toe much longer than fifth finger; 22 subdigital lamellae under fourth finger; 29 subdigital lamellae under fourth toe; subdigital lamellae with sharp keels, bicarinate; slender, swollen at the base; dorsal tail scales keeled, median row forming a serrated ridge, ventral tail scales mucronate, strongly keeled; tail length
<quantity id="4CDF1EEEFFA7FFB14D70FB68FD44FAE4" box="[590,683,1268,1293]" metricMagnitude="-1" metricUnit="m" metricValue="2.67" pageId="23" pageNumber="424" unit="mm" value="267.0">267 mm</quantity>
(tail incomplete, broken at the tip).
</paragraph>
<paragraph id="8B98B30BFFA7FFB14FF9FA8BFE95F9C5" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="424">
<emphasis id="B9536F19FFA7FFB14FF9FA8BFEB0FAD8" bold="true" box="[199,351,1303,1328]" pageId="23" pageNumber="424">Colouration.</emphasis>
In life: dorsum and head uniform, green above with slightly yellowish lateral scales; three broad black bands on the lateral side; dorsal and scales around the orbit greenish yellow with thick black edges forming an indistinct black band from behind the nostril till above the tympanum; head laterally yellow, bluish below the cheeks; pupil black surrounded by a brick red iris; a thin black stripe across the horizontally orbit; lip scales yellow; gular pouch yellow with few slightly orange scales; legs uniform lighter shade of green; tympanum pale bluish green; ventral pale green; tail banded with thick dark and light bands, dark bands almost black near the end. Representative image showing live colouration (
<figureCitation id="131CAF8EFFA7FFB14D89FA73FCE6F9E0" box="[695,777,1519,1544]" captionStart="FIGURE 5" captionStartId="14.[151,250,1665,1687]" captionTargetBox="[344,1243,373,1644]" captionTargetId="figure@14.[344,1243,373,1644]" captionTargetPageId="14" captionText="FIGURE 5. Lateral photograph showing live coloration of A. adult male C. grandisquamis and B. adult male C. nemoricola." httpUri="https://zenodo.org/record/1440684/files/figure.png" pageId="23" pageNumber="424">Fig. 5a</figureCitation>
). In Preservative: colouration mostly similar to that in life, except overall paler.
</paragraph>
<paragraph id="8B98B30BFFA7FFB14FF9F9ABFD16F955" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="424">
<emphasis id="B9536F19FFA7FFB14FF9F9ABFD16F9B8" bold="true" box="[199,761,1591,1616]" pageId="23" pageNumber="424">Variation and secondary sexual characteristics.</emphasis>
Meristic and morphometric data of the examined specimens are given in Appendix 2 &amp; 3. The examined specimens agree with each other in general morphology and scalation. The female specimen (CESL 035) has a reduced nuchal crest composed of shorter spines, dorsal crest almost indistinct and ending at mid body; gular pouch absent.
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFA7FFB14FF9F95BFD03F898" pageId="23" pageNumber="424" type="distribution">
<paragraph id="8B98B30BFFA7FFB14FF9F95BFD03F898" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="424">
<emphasis id="B9536F19FFA7FFB14FF9F95BFE8FF908" bold="true" box="[199,352,1735,1760]" pageId="23" pageNumber="424">Distribution.</emphasis>
<taxonomicName id="4C27C888FFA7FFB14E52F954FD9BF908" authorityName="Gunther" authorityYear="1875" box="[364,628,1736,1760]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="grandisquamis">
<emphasis id="B9536F19FFA7FFB14E52F954FD9BF908" box="[364,628,1736,1760]" italics="true" pageId="23" pageNumber="424">Calotes grandisquamis</emphasis>
</taxonomicName>
is endemic to the
<collectingRegion id="49E37DE9FFA7FFB14C62F954FC58F937" box="[860,951,1736,1759]" country="Sri Lanka" name="Western" pageId="23" pageNumber="424">Western</collectingRegion>
Ghats and is distributed across the high elevation evergreen (above
<quantity id="4CDF1EEEFFA7FFB14EF1F970FDCAF8EC" box="[463,549,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="424" unit="m" value="1000.0">1000 m</quantity>
msl) forests of the southern part of central
<collectingRegion id="49E37DE9FFA7FFB14B2EF971FB84F8EC" box="[1040,1131,1773,1796]" country="Sri Lanka" name="Western" pageId="23" pageNumber="424">Western</collectingRegion>
Ghats (south of Brahmgiri hills) and the southern
<collectingRegion id="49E37DE9FFA7FFB14EA6F88CFE1CF8CF" box="[408,499,1808,1831]" country="Sri Lanka" name="Western" pageId="23" pageNumber="424">Western</collectingRegion>
Ghats. During the present study, this species was recorded from various sites in the hills of the southern
<collectingRegion id="49E37DE9FFA7FFB14EBFF8A9FE33F8A4" box="[385,476,1845,1868]" country="Sri Lanka" name="Western" pageId="23" pageNumber="424">Western</collectingRegion>
Ghats (See Appendix 1 for details). The only record of
<taxonomicName id="4C27C888FFA7FFB14B65F8A8FACCF8A4" authorityName="Gunther" authorityYear="1875" box="[1115,1315,1844,1868]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="grandisquamis">
<emphasis id="B9536F19FFA7FFB14B65F8A8FACCF8A4" box="[1115,1315,1844,1868]" italics="true" pageId="23" pageNumber="424">C. grandisquamis</emphasis>
</taxonomicName>
in
<collectingRegion id="49E37DE9FFA7FFB14A74F8A8FA73F8A5" box="[1354,1436,1844,1869]" country="Sri Lanka" name="Central" pageId="23" pageNumber="424">Central</collectingRegion>
<collectingRegion id="49E37DE9FFA7FFB14FA9F8C4FF1DF887" box="[151,242,1880,1903]" country="Sri Lanka" name="Western" pageId="23" pageNumber="424">Western</collectingRegion>
Ghats is from Agumbe (Ganesh
<emphasis id="B9536F19FFA7FFB14D5AF8C5FD7AF898" box="[612,661,1880,1904]" italics="true" pageId="23" pageNumber="424">et al</emphasis>
. 2013).
</paragraph>
</subSubSection>
<subSubSection id="C33DE080FFA7FFBE4FF9F8E0FB2DFF3D" lastPageId="24" lastPageNumber="425" pageId="23" pageNumber="424" type="biology_ecology">
<paragraph id="8B98B30BFFA7FFBE4FF9F8E0FB2DFF3D" blockId="23.[151,1437,151,2013]" lastBlockId="24.[151,1436,151,213]" lastPageId="24" lastPageNumber="425" pageId="23" pageNumber="424">
<emphasis id="B9536F19FFA7FFB14FF9F8E0FDF1F87D" bold="true" box="[199,542,1916,1941]" pageId="23" pageNumber="424">Ecology and natural history.</emphasis>
<taxonomicName id="4C27C888FFA7FFB14D1BF8E0FCC5F87C" authorityName="Gunther" authorityYear="1875" box="[549,810,1916,1940]" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="424" phylum="Chordata" rank="species" species="grandisquamis">
<emphasis id="B9536F19FFA7FFB14D1BF8E0FCC5F87C" box="[549,810,1916,1940]" italics="true" pageId="23" pageNumber="424">Calotes grandisquamis</emphasis>
</taxonomicName>
is a diurnal lizard, mostly arboreal in habit, and so far, has been recorded from primary evergreen forests and montane shola forests. Individuals were mostly seen perched on branches, high in the canopy and actively moving on tree trunks inside forests. In a few instances, it was found sleeping on lower branches at night. This species has mostly been recorded from higher elevations (above
<quantity id="4CDF1EEEFFA8FFBE4A79FF04FA73FF47" box="[1351,1436,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="425" unit="m" value="1000.0">1000 m</quantity>
msl) in the southern
<collectingRegion id="49E37DE9FFA8FFBE4E43FF21FE37FF3C" box="[381,472,189,212]" country="Sri Lanka" name="Western" pageId="24" pageNumber="425">Western</collectingRegion>
Ghats although there are some reports of it from lower elevations.
</paragraph>
</subSubSection>
</treatment>
</document>