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<document id="C403CC23AAB943ECA99E99E1BE81283E" ID-CLB-Dataset="20929" ID-DOI="10.21248/contrib.entomol.69.2.239-289" ID-GBIF-Dataset="37a37fe7-21ad-45a4-b0d9-880a2b03aa6f" ID-ISSN="0005-805X" ID-Zenodo-Dep="4755240" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620848784293" checkinUser="carolina" docAuthor="Assing, Volker, Brachat, Volker &amp; Meybohm, Heinrich" docDate="2019" docId="C4053B5AFFE1C928FCF58B6980C773AD" docLanguage="en" docName="ContribEntomol.69.2.239-289.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (2)" docSource="http://dx.doi.org/10.21248/contrib.entomol.69.2.239-289" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Stenichnus (Stenichnus) orientalis MEYBOHM 2019, spec. nov." docType="treatment" docUuid="06558FCA-3A62-4019-BFB7-EF06448C1956" docUuidSource="ZooBank" docVersion="14" lastPageNumber="276" masterDocId="383C4322FFC5C90DFF898E638163771A" masterDocTitle="Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera)" masterLastPageNumber="289" masterPageNumber="239" pageNumber="275" updateTime="1698939163438" updateUser="ExternalLinkService">
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<mods:title id="79E7308CE4A3F6601F1F8E1DA7869138">Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera)</mods:title>
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<paragraph id="4C138A4CFFE1C929FCF58B6984CF723F" blockId="36.[892,1452,1290,1382]" box="[892,1452,1290,1317]" pageId="36" pageNumber="275">
<heading id="175B3D20FFE1C929FCF58B6984CF723F" box="[892,1452,1290,1317]" centered="true" fontSize="12" level="2" pageId="36" pageNumber="275" reason="2">
<taxonomicName id="8BACF1CFFFE1C929FCF58B69845C723F" ID-CoL="8Y9TG" authority="MEYBOHM" authorityName="MEYBOHM" authorityYear="2019" box="[892,1343,1290,1317]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="275" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov." subGenus="Stenichnus">
<emphasis id="7ED8565EFFE1C929FCF58B69828F723F" box="[892,1004,1290,1317]" italics="true" pageId="36" pageNumber="275">Stenichnus</emphasis>
(
<emphasis id="7ED8565EFFE1C929FC708B69850A723F" box="[1017,1129,1290,1317]" italics="true" pageId="36" pageNumber="275">Stenichnus</emphasis>
)
<emphasis id="7ED8565EFFE1C929FBFC8B6985B2723F" box="[1141,1233,1290,1317]" italics="true" pageId="36" pageNumber="275">orientalis</emphasis>
MEYBOHM
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<subSubSection id="04B6D9C7FFE1C929FCF58B3184527122" pageId="36" pageNumber="275" type="description">
<paragraph id="4C138A4CFFE1C929FCF58B318288727C" blockId="36.[892,1452,1290,1382]" box="[892,1003,1362,1382]" pageId="36" pageNumber="275">
(
<figureCitation id="D49796C9FFE1C929FC098B31828B727C" box="[896,1000,1362,1382]" captionStart="Figs 127142" captionStartId="14.[212,252,2003,2025]" captionTargetBox="[212,1459,263,1983]" captionTargetId="figure-41@14.[212,1459,255,1994]" captionTargetPageId="14" captionText="Figs 127142: Cephennium chanianum (127129), Euconnus zakrius (130), Stenichnus brachati (131133), S. orientalis (134136), S. aegialioides (137138), S. aegialius (139140), and S. godarti (141142): habitus (127, 134); aedeagus in lateral, ventral, and dorsal view (128130, 132, 136137, 139, 141); male abdomen in ventral view (131, 135); spermatheca (133, 138, 140, 142). Scale bars: 127, 134: 0.5 mm; 131, 135: 0.2 mm; 128130, 132133, 136142: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755268" httpUri="https://zenodo.org/record/4755268/files/figure.png" pageId="36" pageNumber="275">Figs 134136</figureCitation>
)
</paragraph>
<paragraph id="4C138A4CFFE1C929FCDF8BE384527122" blockId="36.[854,1460,1408,1592]" pageId="36" pageNumber="275">
<materialsCitation id="FCC48011FFE1C929FCDF8BE384CC72C2" ID-GBIF-Occurrence="3458714561" collectingDate="2019-03-14" collectionCode="cMey" collectorName="Brachat &amp; Meybohm" determinerName="Meybohm" latitude="35.115833" location="GR Ostkreta" longLatPrecision="19" longitude="35.115833" pageId="36" pageNumber="275" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="931734EEFFE1C929FCDF8BE382DA7282" box="[854,953,1408,1432]" pageId="36" pageNumber="275" type="holotype">Holotype</typeStatus>
<specimenCount id="5AAA41C5FFE1C929FC4D8BE282BA7282" box="[964,985,1409,1432]" pageId="36" pageNumber="275" type="male"></specimenCount>
: “
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<geoCoordinate id="2998EC8BFFE1C929FBE78BE385BA7282" box="[1134,1241,1408,1432]" degrees="26" direction="east" minutes="13" orientation="longitude" pageId="36" pageNumber="275" precision="15" seconds="10" value="26.219444">E26°13'10</geoCoordinate>
,
<location id="4973DC97FFE1C929FB608BE384167282" LSID="urn:lsid:plazi:treatment:C4053B5AFFE1C928FCF58B6980C773AD:4973DC97FFE1C929FB608BE384167282" box="[1257,1397,1408,1432]" latitude="35.115833" longLatPrecision="19" longitude="35.115833" name="GR Ostkreta" pageId="36" pageNumber="275">GR Ostkreta</location>
,
<location id="4973DC97FFE1C929FA0D8BE382C272A2" LSID="urn:lsid:plazi:treatment:C4053B5AFFE1C928FCF58B6980C773AD:4973DC97FFE1C929FA0D8BE382C272A2" latitude="35.115833" longLatPrecision="19" longitude="35.115833" name="Ano Zakros" pageId="36" pageNumber="275">Ano Zakros</location>
<date id="3812AC8CFFE1C929FC228BC3857172A2" box="[939,1042,1440,1464]" pageId="36" pageNumber="275" value="2019-03-14">
<collectingDate id="28565564FFE1C929FC228BC3857172A2" box="[939,1042,1440,1464]" pageId="36" pageNumber="275" value="2019-03-14">14.3.2019</collectingDate>
</date>
,
<collectorName id="E159EF9AFFE1C929FBA98BC3851172A2" box="[1056,1138,1440,1464]" pageId="36" pageNumber="275">Brachat</collectorName>
&amp;
<collectorName id="E159EF9AFFE1C929FB108BC3846472A2" box="[1177,1287,1440,1464]" pageId="36" pageNumber="275">Meybohm</collectorName>
(1) /
<taxonomicName id="8BACF1CFFFE1C929FACA8BC382D472C2" authorityName="MEYBOHM" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="275" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="7ED8565EFFE1C929FACA8BC382D472C2" italics="true" pageId="36" pageNumber="275">Stenichnus orientalis</emphasis>
</taxonomicName>
m.
<determinerName id="B9CF6688FFE1C929FC6A8BA3853272C2" box="[995,1105,1472,1496]" pageId="36" pageNumber="275">Meybohm</determinerName>
2019 det. / Holotypus” (
<collectionCode id="2ABD1289FFE1C929FAE18BA384CB72C2" box="[1384,1448,1472,1496]" pageId="36" pageNumber="275">cMey</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="FCC48011FFE1C929FCDF8B83841872E2" ID-GBIF-Occurrence="3458714397" box="[854,1403,1504,1528]" collectingDate="2019-03-16" elevation="390" location="Lassithi" longitude="35.14917" pageId="36" pageNumber="275" specimenCount="1" stateProvince="Kriti" typeStatus="paratype">
<typeStatus id="931734EEFFE1C929FCDF8B8382A372E2" box="[854,960,1504,1528]" pageId="36" pageNumber="275" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="5AAA41C5FFE1C929FC598B83856A72E2" box="[976,1033,1504,1528]" pageId="36" pageNumber="275" type="female">2 ♀♀</specimenCount>
: same data as
<typeStatus id="931734EEFFE1C929FB348B83847872E2" box="[1213,1307,1504,1528]" pageId="36" pageNumber="275" type="holotype">holotype</typeStatus>
(cMey);
</materialsCitation>
<materialsCitation id="FCC48011FFE1C929FA018B83844F7122" ID-GBIF-Occurrence="3458714588" collectingDate="2019-03-16" elevation="390" latitude="-16.3" location="Lassithi" longLatPrecision="5555" longitude="35.14917" pageId="36" pageNumber="275" specimenCount="1" specimenCount-male="1" stateProvince="Kreta" typeStatus="paratype">
<specimenCount id="5AAA41C5FFE1C929FA018B8384CE72E2" box="[1416,1453,1504,1528]" pageId="36" pageNumber="275" type="male">1 ♂</specimenCount>
: “
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E25°38'30, GR
<collectingRegion id="8E6844AEFFE1C929FBFE886385D1710D" box="[1143,1202,1536,1559]" country="Greece" name="Kriti" pageId="36" pageNumber="275">Kreta</collectingRegion>
<location id="4973DC97FFE1C929FB30886384687102" LSID="urn:lsid:plazi:treatment:C4053B5AFFE1C928FCF58B6980C773AD:4973DC97FFE1C929FB30886384687102" box="[1209,1291,1536,1560]" latitude="-16.3" longLatPrecision="5555" longitude="35.14917" name="Lassithi" pageId="36" pageNumber="275" stateProvince="Kreta">Lassithi</location>
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<elevation id="C7816D7FFFE1C929FA9B8863843B710D" box="[1298,1368,1536,1560]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="36" pageNumber="275" unit="m" value="390.0">390 m</elevation>
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, Kritsa
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s
<date id="3812AC8CFFE1C929FCDF884382E07122" box="[854,899,1568,1592]" pageId="36" pageNumber="275" value="2019-03-16">16.3</date>
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.2019, Brachat &amp; Meybohm (4)” (cMey)
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.
</paragraph>
</subSubSection>
<subSubSection id="04B6D9C7FFE1C929FCDF8803828771A2" pageId="36" pageNumber="275" type="etymology">
<paragraph id="4C138A4CFFE1C929FCDF8803828771A2" blockId="36.[854,1459,1632,1720]" pageId="36" pageNumber="275">
<emphasis id="7ED8565EFFE1C929FCDF880382AE7162" bold="true" box="[854,973,1632,1656]" pageId="36" pageNumber="275">Etymology</emphasis>
: The specific epithet (Latin, adjective: eastern) indicates that the species is endemic to the eastern part of
<collectingRegion id="8E6844AEFFE1C929FC2A88C3828371A2" box="[931,992,1696,1720]" country="Greece" name="Kriti" pageId="36" pageNumber="275">Crete</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="04B6D9C7FFE1C928FCDF88BC835B7322" lastPageId="37" lastPageNumber="276" pageId="36" pageNumber="275" type="description">
<paragraph id="4C138A4CFFE1C929FCDF88BC85A27002" blockId="36.[854,1459,1759,2104]" pageId="36" pageNumber="275">
<emphasis id="7ED8565EFFE1C929FCDF88BC82BB71ED" bold="true" box="[854,984,1759,1783]" pageId="36" pageNumber="275">Description</emphasis>
: Habitus as in
<figureCitation id="D49796C9FFE1C929FB09888385B471E2" box="[1152,1239,1760,1784]" captionStart="Figs 127142" captionStartId="14.[212,252,2003,2025]" captionTargetBox="[212,1459,263,1983]" captionTargetId="figure-41@14.[212,1459,255,1994]" captionTargetPageId="14" captionText="Figs 127142: Cephennium chanianum (127129), Euconnus zakrius (130), Stenichnus brachati (131133), S. orientalis (134136), S. aegialioides (137138), S. aegialius (139140), and S. godarti (141142): habitus (127, 134); aedeagus in lateral, ventral, and dorsal view (128130, 132, 136137, 139, 141); male abdomen in ventral view (131, 135); spermatheca (133, 138, 140, 142). Scale bars: 127, 134: 0.5 mm; 131, 135: 0.2 mm; 128130, 132133, 136142: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755268" httpUri="https://zenodo.org/record/4755268/files/figure.png" pageId="36" pageNumber="275">Fig. 134</figureCitation>
. External characters as in
<taxonomicName id="8BACF1CFFFE1C929FC048963829F7002" authorityName="MEYBOHM" authorityYear="2019" box="[909,1020,1792,1816]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="275" phylum="Arthropoda" rank="species" species="brachati">
<emphasis id="7ED8565EFFE1C929FC048963829F7002" box="[909,1020,1792,1816]" italics="true" pageId="36" pageNumber="275">S. brachati</emphasis>
</taxonomicName>
, except as follows:
</paragraph>
<paragraph id="4C138A4CFFE1C929FCDF894385F07F22" blockId="36.[854,1459,1759,2104]" pageId="36" pageNumber="275">
Body length
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(♂) and
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(♀). Width of head including eyes
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. Pubescence on vertex and frons not sexually dimorphic, in both sexes about
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long, distinctly curved, and directed mediad. Antenna slender, sexually dimorphic:
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(♂) and
<quantity id="8B5427A9FFE1C929FB6489C3842870AD" box="[1261,1355,1952,1976]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="275" unit="mm" value="0.89">0.89 mm</quantity>
(♀) long, antennomeres IX and X in female distinctly broader than in male. Pronotum
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broad and
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(♂) and
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(♀) long, at base laterally with two foveae of similar size as the two median pits.
</paragraph>
<paragraph id="4C138A4CFFE0C928FF278F6383C5768D" blockId="37.[174,779,256,664]" pageId="37" pageNumber="276">
Elytra oblong, 1.00
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long and
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broad, on average 1.57 times as long as broad, strongly convex in cross-section and dorsally not flattened; pubescence very long and erect, setae as long as the combined length of antennomeres III and IV, or nearly so.
</paragraph>
<paragraph id="4C138A4CFFE0C928FF278FC283A0758D" blockId="37.[174,779,256,664]" pageId="37" pageNumber="276">
♂: profemur distally broadly dilated, externally broadly rounded, not angular; median process of sternite III more slender and laterally more distinctly concave than in
<taxonomicName id="8BACF1CFFFE0C928FF278C6380787502" authorityName="MEYBOHM" authorityYear="2019" box="[174,283,512,536]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="276" phylum="Arthropoda" rank="species" species="brachati">
<emphasis id="7ED8565EFFE0C928FF278C6380787502" box="[174,283,512,536]" italics="true" pageId="37" pageNumber="276">S. brachati</emphasis>
</taxonomicName>
, apically with fovea, sternite III without shallow impression, shallow impressions on sternites IV and V more distinctly transverse than in
<taxonomicName id="8BACF1CFFFE0C928FDA48C2383F87542" authorityName="MEYBOHM" authorityYear="2019" box="[557,667,576,600]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="276" phylum="Arthropoda" rank="species" species="brachati">
<emphasis id="7ED8565EFFE0C928FDA48C2383F87542" box="[557,667,576,600]" italics="true" pageId="37" pageNumber="276">S. brachati</emphasis>
</taxonomicName>
, with few minute setae anteriorly (
<figureCitation id="D49796C9FFE0C928FE258C0383677562" box="[428,516,608,632]" captionStart="Figs 127142" captionStartId="14.[212,252,2003,2025]" captionTargetBox="[212,1459,263,1983]" captionTargetId="figure-41@14.[212,1459,255,1994]" captionTargetPageId="14" captionText="Figs 127142: Cephennium chanianum (127129), Euconnus zakrius (130), Stenichnus brachati (131133), S. orientalis (134136), S. aegialioides (137138), S. aegialius (139140), and S. godarti (141142): habitus (127, 134); aedeagus in lateral, ventral, and dorsal view (128130, 132, 136137, 139, 141); male abdomen in ventral view (131, 135); spermatheca (133, 138, 140, 142). Scale bars: 127, 134: 0.5 mm; 131, 135: 0.2 mm; 128130, 132133, 136142: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755268" httpUri="https://zenodo.org/record/4755268/files/figure.png" pageId="37" pageNumber="276">Fig. 135</figureCitation>
); aedeagus as in
<figureCitation id="D49796C9FFE0C928FD388C0382657562" box="[689,774,608,632]" captionStart="Figs 127142" captionStartId="14.[212,252,2003,2025]" captionTargetBox="[212,1459,263,1983]" captionTargetId="figure-41@14.[212,1459,255,1994]" captionTargetPageId="14" captionText="Figs 127142: Cephennium chanianum (127129), Euconnus zakrius (130), Stenichnus brachati (131133), S. orientalis (134136), S. aegialioides (137138), S. aegialius (139140), and S. godarti (141142): habitus (127, 134); aedeagus in lateral, ventral, and dorsal view (128130, 132, 136137, 139, 141); male abdomen in ventral view (131, 135); spermatheca (133, 138, 140, 142). Scale bars: 127, 134: 0.5 mm; 131, 135: 0.2 mm; 128130, 132133, 136142: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755268" httpUri="https://zenodo.org/record/4755268/files/figure.png" pageId="37" pageNumber="276">Fig. 136</figureCitation>
, dorsal sclerotized portion rather short and narrow.
</paragraph>
<paragraph id="4C138A4CFFE0C928FF278CDC80FD7482" blockId="37.[174,779,703,920]" pageId="37" pageNumber="276">
<emphasis id="7ED8565EFFE0C928FF278CDC80C675CD" bold="true" box="[174,421,703,727]" pageId="37" pageNumber="276">Intraspecific variation</emphasis>
: The male
<typeStatus id="931734EEFFE0C928FDAC8CA383E175C2" box="[549,642,704,728]" pageId="37" pageNumber="276" type="paratype">paratype</typeStatus>
from Kritsa differs from the
<typeStatus id="931734EEFFE0C928FEDE8C8380D675E2" box="[343,437,736,760]" pageId="37" pageNumber="276" type="holotype">holotype</typeStatus>
by distally more dilated profemora, less erect, shorter, but more strongly curved setae on the elytra, and a more distinct fovea on the median process of the male sternite III. However, since the aedeagi are identical, these differences are interpreted as intraspecific variation.
</paragraph>
<paragraph id="4C138A4CFFE0C928FF278DDC835B7322" blockId="37.[174,779,959,1080]" pageId="37" pageNumber="276">
<emphasis id="7ED8565EFFE0C928FF278DDC80E674C2" bold="true" box="[174,389,959,984]" pageId="37" pageNumber="276">Comparative notes</emphasis>
: The similar modifications of the male abdominal sternites IIIV suggest that the species is closely related to
<taxonomicName id="8BACF1CFFFE0C928FEEF8A6383517302" authorityName="MEYBOHM" authorityYear="2019" box="[358,562,1024,1048]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="276" phylum="Arthropoda" rank="species" species="brachati">
<emphasis id="7ED8565EFFE0C928FEEF8A6383517302" box="[358,562,1024,1048]" italics="true" pageId="37" pageNumber="276">Stenichnus brachati</emphasis>
</taxonomicName>
,
<taxonomicName id="8BACF1CFFFE0C928FDB78A6381807322" authority="MEYBOHM, 2016" authorityName="MEYBOHM" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="276" phylum="Arthropoda" rank="species" species="chius">
<emphasis id="7ED8565EFFE0C928FDB78A6383EF7302" box="[574,652,1024,1048]" italics="true" pageId="37" pageNumber="276">S. chius</emphasis>
MEYBOHM, 2016
</taxonomicName>
, and
<taxonomicName id="8BACF1CFFFE0C928FE928A4383577322" authority="MEYBOHM, 2016" authorityName="MEYBOHM" authorityYear="2016" box="[283,564,1056,1080]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="276" phylum="Arthropoda" rank="species" species="lesbius">
<emphasis id="7ED8565EFFE0C928FE928A43801A7322" box="[283,377,1056,1080]" italics="true" pageId="37" pageNumber="276">S. lesbius</emphasis>
MEYBOHM, 2016
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="04B6D9C7FFE0C928FF278A3C80C773AD" pageId="37" pageNumber="276" type="distribution">
<paragraph id="4C138A4CFFE0C928FF278A3C80C773AD" blockId="37.[174,779,1119,1208]" pageId="37" pageNumber="276">
<emphasis id="7ED8565EFFE0C928FF278A3C837F736D" bold="true" box="[174,540,1119,1144]" pageId="37" pageNumber="276">Distribution and natural history</emphasis>
: This species is most likely endemic to East
<collectingRegion id="8E6844AEFFE0C928FE2E8AE380807382" box="[423,483,1152,1176]" country="Greece" name="Kriti" pageId="37" pageNumber="276">Crete</collectingRegion>
, where it was found at low elevations up to
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<quantity id="8B5427A9FFE0C928FED28AC380C373AD" box="[347,416,1184,1208]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="37" pageNumber="276" unit="m" value="390.0">390 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>