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<document id="340F12344A0667EEC0A28C9F04F07E02" ID-CLB-Dataset="24006" ID-DOI="10.5281/zenodo.3670493" ID-GBIF-Dataset="77690e3f-3f08-490a-b0e8-0bbcac1fa05c" ID-ISSN="1942-1354" ID-Zenodo-Dep="3670493" ID-ZooBank="4774A311-748B-4D03-932F-2FA60A2DEDD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581944803342" checkinUser="felipe" docAuthor="Nikodým, Milan &amp; Sabatinelli, Guido" docDate="2019" docId="A860EC0C25324902FF32F7DC4DC7FB59" docLanguage="en" docName="InsectaMundi.0693.1-5.pdf.imf" docOrigin="Insecta Mundi 693" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Amphicoma gandhara Nikodym and Sabatinelli 2019, n. sp." docType="treatment" docVersion="9" lastPageNumber="3" masterDocId="5459947425314906FFF2F7054F2FFFD1" masterDocTitle="Amphicoma gandhara, a new species of Glaphyridae (Coleoptera: Scarabaeoidea) from Swat District in northern Pakistan" masterLastPageNumber="5" masterPageNumber="1" pageNumber="2" updateTime="1698783790611" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="F9E119C56FC587DE559CAA74DACFC2DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="05DCD013D2F791464382D8C729F1D45B">
<mods:title id="A352A1B333CA41F457A9E19D72E129A3">Amphicoma gandhara, a new species of Glaphyridae (Coleoptera: Scarabaeoidea) from Swat District in northern Pakistan</mods:title>
</mods:titleInfo>
<mods:name id="925044D0B39B18849183380BEF76B632" type="personal">
<mods:role id="7687935CE1A583A2DCE9A61C81A1C770">
<mods:roleTerm id="C8E8FA2B1CD34CAE3E964AECFF58EF97">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1391F48A2FC7B0F20F9E9D73D264FBE5">Nikodým, Milan</mods:namePart>
</mods:name>
<mods:name id="86C027D62841646D66296EC195E1BD1A" type="personal">
<mods:role id="6E96D2EEA658733C96E5CA900A75878D">
<mods:roleTerm id="72C12D954324B912EFE7C1621186F649">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D27A040639C970596AFE9B9A5F07AB0C">Sabatinelli, Guido</mods:namePart>
</mods:name>
<mods:typeOfResource id="0054536431072CE01AA6B14B388E7B12">text</mods:typeOfResource>
<mods:relatedItem id="1AD9EBC2DECB7039822226A7E7D9B6F9" type="host">
<mods:titleInfo id="BFE724E24250E82BDE6A193F7DF3024B">
<mods:title id="A25482F95CCB51D7A2F15E8D23917D53">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="1A45C106A3D2EE6C1278D0737A72DA60">
<mods:date id="A2381699E3D1709CE96D48A982C255A1">2019</mods:date>
<mods:detail id="D3C3B70FEC4F4C278484D83F0ED36D23" type="pubDate">
<mods:number id="006F789E2AE05B1ECB947CB20EB3A4B6">2019-03-29</mods:number>
</mods:detail>
<mods:detail id="F44149BF4CEEDEC76DE99047E46497BB" type="issue">
<mods:number id="C8D09BCA1BC7FC42AA83778980CE1914">693</mods:number>
</mods:detail>
<mods:extent id="F0FF31D5099EF74CF4029A3A40A327C4" unit="page">
<mods:start id="CEBBD328515E97C93D348A115FCC0CF9">1</mods:start>
<mods:end id="354738D41E7D58EFA088560461260E85">5</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="340D2E4CED433B3D61CF6D0DF792BF0A">journal article</mods:classification>
<mods:identifier id="66E744FF3721549AFF141F93D4AE1B53" type="CLB-Dataset">24006</mods:identifier>
<mods:identifier id="D265AB598A05843E2289C2FE40BB52A8" type="DOI">10.5281/zenodo.3670493</mods:identifier>
<mods:identifier id="0383AFA85CE1024ECCA6B71186724E13" type="GBIF-Dataset">77690e3f-3f08-490a-b0e8-0bbcac1fa05c</mods:identifier>
<mods:identifier id="C3C337CE4EBDDD3DFB2111080CE4C8E4" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="33BBFEB9010ED96B66497CAD709C12E5" type="Zenodo-Dep">3670493</mods:identifier>
<mods:identifier id="8772B910692C918EABEAF7D50C9F6319" type="ZooBank">4774A311-748B-4D03-932F-2FA60A2DEDD7</mods:identifier>
</mods:mods>
<treatment id="A860EC0C25324902FF32F7DC4DC7FB59" ID-DOI="http://doi.org/10.5281/zenodo.3671049" ID-GBIF-Taxon="162146783" ID-Zenodo-Dep="3671049" LSID="urn:lsid:plazi:treatment:A860EC0C25324902FF32F7DC4DC7FB59" httpUri="http://treatment.plazi.org/id/A860EC0C25324902FF32F7DC4DC7FB59" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<subSubSection id="68D30E9125324905FF32F7DC4C8EFF20" box="[192,929,217,241]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="20765D1A25324905FF32F7DC4C8EFF20" blockId="3.[192,929,217,274]" box="[192,929,217,241]" pageId="3" pageNumber="2">
<heading id="7B3EEA7625324905FF32F7DC4C8EFF20" bold="true" box="[192,929,217,241]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<emphasis id="12BD810825324905FF32F7DC4C8EFF20" bold="true" box="[192,929,217,241]" pageId="3" pageNumber="2">
<taxonomicName id="E7C9269925324905FF32F7DC4C7DFF20" ID-CoL="CZX8" authority="Nikodym and Sabatinelli" authorityName="Nikodym and Sabatinelli" authorityYear="2019" box="[192,850,217,241]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gandhara" status="sp. nov.">
<emphasis id="12BD810825324905FF32F7DC4EDCFF20" bold="true" box="[192,499,217,241]" italics="true" pageId="3" pageNumber="2">Amphicoma gandhara</emphasis>
Nikodým and Sabatinelli
</taxonomicName>
<taxonomicNameLabel id="098E3C7325324905FCABF7DC4C8EFF20" box="[857,929,217,241]" pageId="3" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125324905FF32F7FF4E0CFEC3" box="[192,291,250,274]" pageId="3" pageNumber="2" type="description">
<paragraph id="20765D1A25324905FF32F7FF4E0CFEC3" blockId="3.[192,929,217,274]" box="[192,291,250,274]" pageId="3" pageNumber="2">
<figureCitation id="B8F2419F25324905FF32F7FF4E0CFEC3" box="[192,291,250,274]" captionStart="Figures 1–9" captionStartId="6.[192,286,1848,1870]" captionTargetBox="[280,1333,319,1767]" captionTargetId="figure@6.[268,1364,316,1778]" captionTargetPageId="6" captionText="Figures 1–9. Amphicoma spp. 1-3) Amphicoma gandhara, new species, dorsal habitus. 1) Holotypus Ƌ. 2) Paratypus Ƌ#1. 3) Paratypus Ƌ#2. 4-6). Amphicoma gandhara, new species, parameres. 4) Right view. 5) Dorsal. 6) Left. 7-9) Amphicoma schneideri Nikodým, parameres. 7) Right view. 8) Dorsal. 9) Left." figureDoi="http://doi.org/10.5281/zenodo.3670495" httpUri="https://zenodo.org/record/3670495/files/figure.png" pageId="3" pageNumber="2">Fig. 1–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125324905FF32F62F4E1CFDD2" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="20765D1A25324905FF32F62F4E1CFDD2" blockId="3.[192,1441,297,515]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F62F4E4BFE90" bold="true" box="[192,356,297,322]" pageId="3" pageNumber="2">Type series.</emphasis>
<materialsCitation id="90A1574725324905FE9EF62F4C00FEB3" ID-GBIF-Occurrence="2565890966" collectingDate="2008-07" collectorName="Arihta &amp; M. Nikodym &amp; G. Sabatinelli" country="Pakistan" location="Tauzin" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus id="FF72E3B825324905FE9EF62F4EC6FE93" box="[364,489,298,322]" pageId="3" pageNumber="2" type="holotype">Holotypus</typeStatus>
Ƌ, Paratypus Ƌ#1 and Paratypus Ƌ#2, labeled as follow: white label, printed, “ex H.-P.
<location id="25160BC125324905FEC7F64F4EA4FEB3" LSID="urn:lsid:plazi:treatment:A860EC0C25324902FF32F7DC4DC7FB59:25160BC125324905FEC7F64F4EA4FEB3" box="[309,395,330,354]" country="Pakistan" name="Tauzin" pageId="3" pageNumber="2">Tauzin</location>
/ collection” // white label, printed
</materialsCitation>
<materialsCitation id="90A1574725324905FCC5F64F4C6CFE53" ID-GBIF-Occurrence="2565890964" collectingDate="2008-07" collectorName="Arihta &amp; M. Nikodym &amp; G. Sabatinelli" country="Pakistan" location="Tauzin" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
“
<collectingCountry id="58DE1D8A25324905FCB3F64F4CFEFEB3" box="[833,977,330,354]" name="Pakistan" pageId="3" pageNumber="2">PAKISTAN</collectingCountry>
- Khyber / Pakhtunkhwa Prov., Swat / District,
<date id="54777BDA25324905FECEF66F4E98FE53" box="[316,439,362,386]" pageId="3" pageNumber="2" value="2008-07">
<collectingDate id="4433823225324905FECEF66F4E98FE53" box="[316,439,362,386]" pageId="3" pageNumber="2" value="2008-07">July 2008</collectingDate>
</date>
,
<collectorName id="8D3C38CC25324905FE31F66F4D3BFE53" box="[451,532,362,386]" pageId="3" pageNumber="2">Arihta</collectorName>
leg.” // red label printed:
</materialsCitation>
<materialsCitation id="90A1574725324905FCB9F66F4E08FE13" ID-GBIF-Occurrence="2565890967" collectingDate="2008-07" collectorName="Arihta &amp; M. Nikodym &amp; G. Sabatinelli" country="Pakistan" location="Tauzin" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
“
<typeStatus id="FF72E3B825324905FCA7F66F4B2CFE53" box="[853,1027,362,386]" pageId="3" pageNumber="2" type="holotype">HOLOTYPUS</typeStatus>
Ƌ /
<taxonomicName id="E7C9269925324905FBC7F66F4A68FE53" authority="Nikodým &amp; Sabatinelli, 2019" authorityName="Nikodým &amp; Sabatinelli" authorityYear="2019" box="[1077,1351,362,386]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gandhara" status="sp. nov.">
<emphasis id="12BD810825324905FBC7F66F4A68FE53" box="[1077,1351,362,386]" italics="true" pageId="3" pageNumber="2">Amphicoma gandhara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="098E3C7325324905FABDF66E4ABEFE53" box="[1359,1425,363,386]" pageId="3" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
/
<collectorName id="8D3C38CC25324905FF32F68E4E78FE73" box="[192,343,394,418]" pageId="3" pageNumber="2">M. Nikodým</collectorName>
and
<collectorName id="8D3C38CC25324905FE61F68F4D11FE73" box="[403,574,394,418]" pageId="3" pageNumber="2">G. Sabatinelli</collectorName>
, 2019” // (deposited in Muséum d’Histoire Naturelle, Geneva, Switzerland). //
</materialsCitation>
<materialsCitation id="90A1574725324905FEDDF6AF4E1DFE32" ID-GBIF-Occurrence="2565890968" collectingDate="2008-07" collectorName="Arihta &amp; M. Nikodym &amp; G. Sabatinelli" country="Pakistan" location="Tauzin" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
“
<typeStatus id="FF72E3B825324905FECBF6AF4ECEFE13" box="[313,481,426,450]" pageId="3" pageNumber="2" type="paratype">PARATYPUS</typeStatus>
Ƌ#1” // with other data as the
<typeStatus id="FF72E3B825324905FC90F6AF4CE5FE13" box="[866,970,426,450]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
in
<collectorName id="8D3C38CC25324905FC06F6AF4BB9FE13" box="[1012,1174,426,450]" pageId="3" pageNumber="2">G.Sabatinelli</collectorName>
collection, Prévessin, France. //
</materialsCitation>
<materialsCitation id="90A1574725324905FEC5F6CE4E1CFDD2" ID-GBIF-Occurrence="2565890965" collectingDate="2008-07" collectorName="Arihta &amp; M. Nikodym &amp; G. Sabatinelli" country="Pakistan" location="Tauzin" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
“
<typeStatus id="FF72E3B825324905FEB3F6CE4EC6FE32" box="[321,489,459,483]" pageId="3" pageNumber="2" type="paratype">PARATYPUS</typeStatus>
Ƌ#2” with other data as the
<typeStatus id="FF72E3B825324905FCCDF6CF4C88FE33" box="[831,935,458,482]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
in M. Nikodým collection, Roztoky, Czech Republic.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125324905FF32F51F4A1DFD02" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="20765D1A25324905FF32F51F4A1DFD02" blockId="3.[192,1440,538,723]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F51F4E60FDE3" bold="true" box="[192,335,538,562]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="E7C9269925324905FEA6F51F4D4BFDE3" authorityName="Nikodým &amp; Sabatinelli" authorityYear="2019" box="[340,612,538,562]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gandhara">
<emphasis id="12BD810825324905FEA6F51F4D4BFDE3" box="[340,612,538,562]" italics="true" pageId="3" pageNumber="2">Amphicoma gandhara</emphasis>
</taxonomicName>
belongs to the “
<taxonomicName id="E7C9269925324905FCD2F51E4CA8FDE3" authorityName="Solsky" authorityYear="1876" box="[800,903,538,562]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="12BD810825324905FCD2F51E4CA8FDE3" box="[800,903,538,562]" italics="true" pageId="3" pageNumber="2">A. dubia</emphasis>
</taxonomicName>
group-species” (
<bibRefCitation id="445820EB25324905FBB8F51E4BD3FDE2" author="Nikodym, M." box="[1098,1276,539,563]" pageId="3" pageNumber="2" pagination="1 - 35" refId="ref2870" refString="Nikodym, M. 2005. New species of the genus Amphicoma from China, Vietnam, Laos and Thailand (Coleoptera: Scarabaeoidea: Glaphyridae). Animma. X (Plzen) 10: 1 - 35." type="journal article" year="2005">Nikodým 2005</bibRefCitation>
) based on the following characters: mesotibia with apical spur, antennal club more than 1.5 times longer than stalk and markedly out curved. The most similar species is
<taxonomicName id="E7C9269925324905FC92F55E4A10FDA2" authority="Nikodym, 2005" authorityName="Nikodym" authorityYear="2005" box="[864,1343,603,627]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="12BD810825324905FC92F55E4B57FDA2" box="[864,1144,603,627]" italics="true" pageId="3" pageNumber="2">Amphicoma schneideri</emphasis>
<bibRefCitation id="445820EB25324905FB73F55E4A10FDA2" author="Nikodym, M." box="[1153,1343,603,627]" pageId="3" pageNumber="2" pagination="1 - 35" refId="ref2870" refString="Nikodym, M. 2005. New species of the genus Amphicoma from China, Vietnam, Laos and Thailand (Coleoptera: Scarabaeoidea: Glaphyridae). Animma. X (Plzen) 10: 1 - 35." type="journal article" year="2005">Nikodým, 2005</bibRefCitation>
</taxonomicName>
, having similar body length
<quantity id="E731F0FF25324905FE5CF57E4D2CFD42" box="[430,515,635,659]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="2" unit="mm" value="14.0">14 mm</quantity>
, antennomeres 5–7 disc-shaped, elytra uniformly coppery green and terminal maxillary palpomere quite long. In
<emphasis id="12BD810825324905FD9BF59E4C26FD62" box="[617,777,667,691]" italics="true" pageId="3" pageNumber="2">
<taxonomicName id="E7C9269925324905FD9BF59E4C2BFD62" authorityName="Nikodým &amp; Sabatinelli" authorityYear="2019" box="[617,772,667,691]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gandhara">A. gandhara</taxonomicName>
,
</emphasis>
the elytra are rather dull with sutural angles right but with blunt angle while in
<taxonomicName id="E7C9269925324905FE0EF5BE4DB5FD02" box="[508,666,699,723]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="12BD810825324905FE0EF5BE4DB5FD02" box="[508,666,699,723]" italics="true" pageId="3" pageNumber="2">A. schneideri</emphasis>
</taxonomicName>
elytra are shiny with sutural angles broadly rounded.
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125324905FF32F5EE4C63F966" pageId="3" pageNumber="2" type="description">
<paragraph id="20765D1A25324905FF32F5EE4CF0FCF5" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F5EE4DDAFCD2" bold="true" box="[192,757,747,772]" pageId="3" pageNumber="2">
Description of the
<typeStatus id="FF72E3B825324905FE36F5E94D13FCD5" box="[452,572,748,772]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
(adult male).
</emphasis>
Body elongate, length (from clypeal anterior margin to apex of elytra)
<quantity id="E731F0FF25324905FE85F4094EE1FCF2" box="[375,462,780,804]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="2" unit="mm" value="13.0">13 mm</quantity>
, width across elytral humeri
<quantity id="E731F0FF25324905FCC6F4094C56FCF2" box="[820,889,780,804]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="2" unit="mm" value="5.0">5 mm</quantity>
(
<figureCitation id="B8F2419F25324905FC78F4094CE1FCF5" box="[906,974,780,804]" captionStart="Figures 1–9" captionStartId="6.[192,286,1848,1870]" captionTargetBox="[280,1333,319,1767]" captionTargetId="figure@6.[268,1364,316,1778]" captionTargetPageId="6" captionText="Figures 1–9. Amphicoma spp. 1-3) Amphicoma gandhara, new species, dorsal habitus. 1) Holotypus Ƌ. 2) Paratypus Ƌ#1. 3) Paratypus Ƌ#2. 4-6). Amphicoma gandhara, new species, parameres. 4) Right view. 5) Dorsal. 6) Left. 7-9) Amphicoma schneideri Nikodým, parameres. 7) Right view. 8) Dorsal. 9) Left." figureDoi="http://doi.org/10.5281/zenodo.3670495" httpUri="https://zenodo.org/record/3670495/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="20765D1A25324905FF1AF4294AB6FC55" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF4294E05FC95" box="[232,298,812,836]" italics="true" pageId="3" pageNumber="2">Color</emphasis>
. Head from above dark green with irregular golden luster, pronotum black green with weak golden luster. Scutellum and elytra even darker, luster is irregular from golden green to dark purple. Pygidium yellowish brown without reflection. Antennae brown, legs black-brown, tarsi lighter brown.
</paragraph>
<paragraph id="20765D1A25324905FF1AF4884EA0FBD4" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF4884E7AFC75" box="[232,341,909,932]" italics="true" pageId="3" pageNumber="2">Vestiture</emphasis>
. Dorsal setation yellow-brown, erect on clypeus and scutellum. Head setation longer, light brown and oblique toward apex. Elytra setation as long as on pronotum, irregularly inclined or erect and with solitary long brown setae in posterior quarter. Ventral setation yellow, legs setation light yellowish brown.
</paragraph>
<paragraph id="20765D1A25324905FF1AF3084B65FB14" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF3084E06FBF4" box="[232,297,1037,1061]" italics="true" pageId="3" pageNumber="2">Head</emphasis>
. Anterior margin of labrum slightly emarginate medially. Clypeus 1.1 × wider than long, lateral margins slightly rounded, anterior angles rounded, clypeofrontal suture fine but clearly distinct. Head strongly and densely punctuate. Clypeal punctation clearly weaker and not closely spaced. Antenna consisting of 10 antennomeres; 5, 6 and 7 disc-shaped; antennal club markedly out-curved, more than 2 × longer than wide. Terminal maxillary palpomere long, slightly widened in the middle, truncate at the end, without impression and with several punctures in apical quarter.
</paragraph>
<paragraph id="20765D1A25324905FF1AF3CB4CDCFAF7" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF3CB4E4EFB34" box="[232,353,1230,1253]" italics="true" pageId="3" pageNumber="2">Pronotum</emphasis>
. Width 1.1 × length, widest medially, narrowing anteriorly. Anterior angles produced and 90°, posterior angles obtuse. Basal margin markedly emarginate medially. Punctation dense and coarse, distance between punctures. With median longitudinal depression.
</paragraph>
<paragraph id="20765D1A25324905FF1AF22B4A79FA97" blockId="3.[192,1440,747,1543]" box="[232,1366,1326,1350]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF22B4E4BFA97" box="[232,356,1326,1350]" italics="true" pageId="3" pageNumber="2">Scutellum</emphasis>
. Triangular, elongate with slightly rounded apex, punctation similar to pronotum.
</paragraph>
<paragraph id="20765D1A25324905FF1AF24B4D22FA76" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF24B4E19FAB7" box="[232,310,1358,1382]" italics="true" pageId="3" pageNumber="2">Elytra</emphasis>
. Punctation finer than that of pronotum, slightly irregularly wrinkled. Posterolateral angles broadly rounded, sutural angles 90° but with blunt angles. Sutural margin distinctly raised apically from second third of elytra.
</paragraph>
<paragraph id="20765D1A25324905FF1AF2AA4CEDFA37" blockId="3.[192,1440,747,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF2AA4E30FA17" box="[232,287,1455,1478]" italics="true" pageId="3" pageNumber="2">Legs</emphasis>
. Protibia tridentate, mesotibia with apical spur shorter, less than half of mesotarsomere length. Basal metatarsomere 1.6 × longer than second metatarsomere.
</paragraph>
<paragraph id="20765D1A25324905FF1AF2EA4A5AF9D6" blockId="3.[192,1440,747,1543]" box="[232,1397,1519,1543]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF1AF2EA4E73F9D6" box="[232,348,1519,1543]" italics="true" pageId="3" pageNumber="2">Aedeagus</emphasis>
. Parameres asymmetrical with the right paramere larger than the left one (
<figureCitation id="B8F2419F25324905FAF3F2EA4A47F9D6" box="[1281,1384,1519,1543]" captionStart="Figures 1–9" captionStartId="6.[192,286,1848,1870]" captionTargetBox="[280,1333,319,1767]" captionTargetId="figure@6.[268,1364,316,1778]" captionTargetPageId="6" captionText="Figures 1–9. Amphicoma spp. 1-3) Amphicoma gandhara, new species, dorsal habitus. 1) Holotypus Ƌ. 2) Paratypus Ƌ#1. 3) Paratypus Ƌ#2. 4-6). Amphicoma gandhara, new species, parameres. 4) Right view. 5) Dorsal. 6) Left. 7-9) Amphicoma schneideri Nikodým, parameres. 7) Right view. 8) Dorsal. 9) Left." figureDoi="http://doi.org/10.5281/zenodo.3670495" httpUri="https://zenodo.org/record/3670495/files/figure.png" pageId="3" pageNumber="2">Fig. 4–6</figureCitation>
).
</paragraph>
<paragraph id="20765D1A25324905FF32F11A4C63F966" blockId="3.[192,1440,1567,1720]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F11A4EF3F9E6" bold="true" box="[192,476,1567,1591]" pageId="3" pageNumber="2">Relevant variability.</emphasis>
The three specimens known so far show a remarkable difference in the integument color.
<typeStatus id="FF72E3B825324905FEA5F1454EF9F989" box="[343,470,1600,1624]" pageId="3" pageNumber="2" type="paratype">Paratypus</typeStatus>
Ƌ#1 (
<figureCitation id="B8F2419F25324905FDD5F13A4D5FF986" box="[551,624,1599,1623]" captionStart="Figures 1–9" captionStartId="6.[192,286,1848,1870]" captionTargetBox="[280,1333,319,1767]" captionTargetId="figure@6.[268,1364,316,1778]" captionTargetPageId="6" captionText="Figures 1–9. Amphicoma spp. 1-3) Amphicoma gandhara, new species, dorsal habitus. 1) Holotypus Ƌ. 2) Paratypus Ƌ#1. 3) Paratypus Ƌ#2. 4-6). Amphicoma gandhara, new species, parameres. 4) Right view. 5) Dorsal. 6) Left. 7-9) Amphicoma schneideri Nikodým, parameres. 7) Right view. 8) Dorsal. 9) Left." figureDoi="http://doi.org/10.5281/zenodo.3670495" httpUri="https://zenodo.org/record/3670495/files/figure.png" pageId="3" pageNumber="2">Fig. 2</figureCitation>
) has the dorsal surface black violet and irregular golden luster.
<typeStatus id="FF72E3B825324905FF32F1654E10F9A9" box="[192,319,1632,1656]" pageId="3" pageNumber="2" type="paratype">Paratypus</typeStatus>
Ƌ#2 (
<figureCitation id="B8F2419F25324905FE77F15A4EE7F9A9" box="[389,456,1631,1656]" captionStart="Figures 1–9" captionStartId="6.[192,286,1848,1870]" captionTargetBox="[280,1333,319,1767]" captionTargetId="figure@6.[268,1364,316,1778]" captionTargetPageId="6" captionText="Figures 1–9. Amphicoma spp. 1-3) Amphicoma gandhara, new species, dorsal habitus. 1) Holotypus Ƌ. 2) Paratypus Ƌ#1. 3) Paratypus Ƌ#2. 4-6). Amphicoma gandhara, new species, parameres. 4) Right view. 5) Dorsal. 6) Left. 7-9) Amphicoma schneideri Nikodým, parameres. 7) Right view. 8) Dorsal. 9) Left." figureDoi="http://doi.org/10.5281/zenodo.3670495" httpUri="https://zenodo.org/record/3670495/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
) has the dorsal surface black green and irregular golden luster. In
<typeStatus id="FF72E3B825324905FB18F1654A46F9A9" box="[1258,1385,1632,1656]" pageId="3" pageNumber="2" type="paratype">Paratypus</typeStatus>
Ƌ#1 the antennal club is markedly deformed, segments are narrowed and distorted apically. This feature is also known in other species of the genus
<taxonomicName id="E7C9269925324905FD41F19A4C6AF966" authorityName="Latreille" authorityYear="1807" box="[691,837,1695,1719]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="12BD810825324905FD41F19A4C6AF966" box="[691,837,1695,1719]" italics="true" pageId="3" pageNumber="2">Amphicoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125324905FF32F1D54E9CF8C9" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="20765D1A25324905FF32F1D54B14F939" blockId="3.[192,1083,1743,1768]" box="[192,1083,1743,1768]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F1D54E54F936" bold="true" box="[192,379,1743,1768]" pageId="3" pageNumber="2">
<typeStatus id="FF72E3B825324905FF32F1D54E2BF939" box="[192,260,1744,1768]" pageId="3" pageNumber="2">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="58DE1D8A25324905FE70F1D54EDCF939" box="[386,499,1744,1768]" name="Pakistan" pageId="3" pageNumber="2">Pakistan</collectingCountry>
,
<collectingRegion id="E20D93F825324905FE0CF1D54CACF939" box="[510,899,1744,1768]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="3" pageNumber="2">Khyber Pakhtunkhwa Province</collectingRegion>
,
<location id="25160BC125324905FC62F1D54B19F939" LSID="urn:lsid:plazi:treatment:A860EC0C25324902FF32F7DC4DC7FB59:25160BC125324905FC62F1D54B19F939" box="[912,1078,1744,1768]" country="Pakistan" name="Swat District" pageId="3" pageNumber="2" stateProvince="Khyber Pakhtunkhwa">Swat District</location>
.
</paragraph>
<paragraph id="20765D1A25324905FF32F0054E9CF8C9" blockId="3.[192,435,1792,1816]" box="[192,435,1792,1816]" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F0054E03F8C9" bold="true" box="[192,300,1792,1816]" pageId="3" pageNumber="2">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125324902FF32F0354B3FFF20" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="etymology">
<paragraph id="20765D1A25324902FF32F0354B3FFF20" blockId="3.[192,1440,1840,1928]" lastBlockId="4.[192,1040,217,241]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<emphasis id="12BD810825324905FF32F0354E76F899" bold="true" box="[192,345,1840,1864]" pageId="3" pageNumber="2">Etymology.</emphasis>
The specimens were collected in Swat, a valley and an administrative district in the
<collectingRegion id="E20D93F825324905FAB4F0354EF5F8B9" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="3" pageNumber="2">Khyber Pakhtunkhwa province</collectingRegion>
of
<collectingCountry id="58DE1D8A25324905FE09F0554D43F8B9" box="[507,620,1872,1896]" name="Pakistan" pageId="3" pageNumber="2">Pakistan</collectingCountry>
. Swat is renowned for its outstanding natural beauty and this region was also a major center of early Buddhist thought as part of the Gandhara kingdom. The new species is named after this ancient Indo-Aryan kingdom. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="68D30E9125354902FF32F60F4DC7FB59" pageId="4" pageNumber="3" type="discussion">
<paragraph id="20765D1A25354902FF32F60F4B7FFE74" blockId="4.[192,1440,266,488]" pageId="4" pageNumber="3">
<emphasis id="12BD810825354902FF32F60F4E6EFEF3" bold="true" box="[192,321,266,290]" pageId="4" pageNumber="3">Remarks.</emphasis>
The genus
<taxonomicName id="E7C9269925354902FE3AF60F4D76FEF3" authorityName="Latreille" authorityYear="1807" box="[456,601,266,290]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="12BD810825354902FE3AF60F4D76FEF3" box="[456,601,266,290]" italics="true" pageId="4" pageNumber="3">Amphicoma</emphasis>
</taxonomicName>
is known from Europe (
<collectingCountry id="58DE1D8A25354902FC81F60F4C93FEF3" box="[883,956,266,290]" name="Spain" pageId="4" pageNumber="3">Spain</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FC34F60F4B2DFEF3" box="[966,1026,266,290]" name="Italy" pageId="4" pageNumber="3">Italy</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FBFFF60F4B8DFEF3" box="[1037,1186,266,290]" name="Switzerland" pageId="4" pageNumber="3">Switzerland</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FB5EF60F4BD0FEF3" box="[1196,1279,266,290]" name="Greece" pageId="4" pageNumber="3">Greece</collectingCountry>
and
<collectingCountry id="58DE1D8A25354902FACAF60F4AB7FEF3" box="[1336,1432,266,290]" name="Albania" pageId="4" pageNumber="3">Albania</collectingCountry>
) and Asia (
<collectingCountry id="58DE1D8A25354902FEC4F62E4EADFE92" box="[310,386,299,323]" name="China" pageId="4" pageNumber="3">China</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FE79F6294ECDFE92" box="[395,482,300,323]" name="Myanmar" pageId="4" pageNumber="3">Burma</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FE1EF62E4D73FE92" box="[492,604,299,323]" name="Thailand" pageId="4" pageNumber="3">Thailand</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FD97F6294D8FFE92" box="[613,672,300,323]" name="Laos" pageId="4" pageNumber="3">Laos</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FD59F62E4C3AFE92" box="[683,789,299,323]" name="Vietnam" pageId="4" pageNumber="3">Vietnam</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FCEDF62E4C53FE92" box="[799,892,299,323]" name="Taiwan" pageId="4" pageNumber="3">Taiwan</collectingCountry>
,
<collectingCountry id="58DE1D8A25354902FC74F6294CFFFE92" box="[902,976,300,323]" name="Japan" pageId="4" pageNumber="3">Japan</collectingCountry>
and Ryukyu Islands). The new location from northern
<collectingCountry id="58DE1D8A25354902FE86F6494ECDFEB5" box="[372,482,332,356]" name="Pakistan" pageId="4" pageNumber="3">Pakistan</collectingCountry>
is disjoint from all known localities for the genus
<emphasis id="12BD810825354902FBCCF6494BF6FEB5" box="[1086,1241,332,356]" italics="true" pageId="4" pageNumber="3">
<taxonomicName id="E7C9269925354902FBCCF6494BFBFEB5" authorityName="Latreille" authorityYear="1807" box="[1086,1236,332,356]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">Amphicoma</taxonomicName>
.
</emphasis>
This indicates a potential for more species to be found in the Himalayan region:
<taxonomicName id="E7C9269925354902FC3DF6684B47FE54" authorityName="MacLeay" authorityYear="1819" box="[975,1128,365,389]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
can be expected from the eastern part of
<collectingCountry id="58DE1D8A25354902FE8EF6884E92FE74" box="[380,445,397,421]" name="India" pageId="4" pageNumber="3">India</collectingCountry>
(
<collectingRegion id="E20D93F825354902FE3FF6884D67FE74" box="[461,584,397,421]" country="India" name="Nagaland" pageId="4" pageNumber="3">Nagaland</collectingRegion>
,
<collectingRegion id="E20D93F825354902FDA6F6884DEDFE74" box="[596,706,397,421]" country="India" name="Manipur" pageId="4" pageNumber="3">Manipur</collectingRegion>
,
<collectingRegion id="E20D93F825354902FD3CF6884C13FE74" box="[718,828,397,421]" country="India" name="Mizoram" pageId="4" pageNumber="3">Mizoram</collectingRegion>
regions) or
<collectingCountry id="58DE1D8A25354902FC3FF6884B62FE74" box="[973,1101,397,421]" name="Cambodia" pageId="4" pageNumber="3">Cambodia</collectingCountry>
.
</paragraph>
<paragraph id="20765D1A25354902FF1AF6AA4DC7FB59" blockId="4.[192,1440,266,488]" lastBlockId="4.[192,744,1136,1160]" pageId="4" pageNumber="3">
For comparisons with the most similar species, we studied
<specimenCount id="36CF969325354902FC58F6AB4B64FE17" box="[938,1099,430,454]" pageId="4" pageNumber="3" type="generic">47 specimens</specimenCount>
of
<taxonomicName id="E7C9269925354902FB9FF6AB4A26FE17" box="[1133,1289,430,454]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="12BD810825354902FB9FF6AB4A26FE17" box="[1133,1289,430,454]" italics="true" pageId="4" pageNumber="3">A. schneideri</emphasis>
</taxonomicName>
from
<collectingCountry id="58DE1D8A25354902FABCF6AB4AB3FE17" box="[1358,1436,430,454]" name="China" pageId="4" pageNumber="3">China</collectingCountry>
:
<collectingRegion id="E20D93F825354902FF32F6CA4E08FE36" box="[192,295,463,487]" country="China" name="Sichuan" pageId="4" pageNumber="3">Sichuan</collectingRegion>
, Guan Xian and Songpan and
<collectingRegion id="E20D93F825354902FD6CF6CA4C29FE36" box="[670,774,463,487]" country="China" name="Shaanxi" pageId="4" pageNumber="3">Shaanxi</collectingRegion>
, Luoyang. The two species can be easily distinguished by several characters as indicated in
<tableCitation id="6D4B68A125354902FD74F3754DCDFB59" box="[646,738,1136,1160]" captionStart="Table 1" captionStartId="4.[192,261,504,525]" captionTargetBox="[203,1425,580,1070]" captionTargetId="graphics@4.[192,1440,566,1074]" captionText="Table 1. Comparative analysis of main characters in Amphicoma gandhara n. sp. and Amphicoma schneideri Nikodým." httpUri="http://table.plazi.org/id/74B60D9225354902FF32F6FD4E03FDFA" pageId="4" pageNumber="3" tableUuid="74B60D9225354902FF32F6FD4E03FDFA">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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