treatments-xml/data/91/75/4C/91754C375F36FFCBFF17DBDFFCA5FF69.xml

<document id="CE8C8DBBFE300F83ADC57898641CDCFA" ID-CLB-Dataset="32029" ID-DOI="10.11646/zootaxa.4323.2.5" ID-GBIF-Dataset="a73b94ca-14ed-48f6-833d-ef44674c3fbe" ID-ISSN="1175-5326" ID-Zenodo-Dep="898940" ID-ZooBank="E50Ceeac-1219-417D-8Bb8-Fdde0Dcb6Bde" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505970918151" checkinUser="plazi" docAuthor="Devi, Suvarna S." docDate="2017" docId="91754C375F36FFCBFF17DBDFFCA5FF69" docLanguage="en" docName="zootaxa.4323.2.5.pdf" docOrigin="Zootaxa 4323 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laleonectes kuriya Devi 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="227" masterDocId="6D4C344F5F37FFC2FF80DA71FFF3FF9E" masterDocTitle="A new species of the swimming crab genus, Laleonectes Manning &amp; Chace, 1990 (Crustacea: Brachyura: Portunidae), from the western Indian Ocean" masterLastPageNumber="228" masterPageNumber="219" pageNumber="220" updateTime="1698468837196" updateUser="ExternalLinkService">
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<mods:title id="96E09FCE812D4C663F198D51AA1B9693">A new species of the swimming crab genus, Laleonectes Manning &amp; Chace, 1990 (Crustacea: Brachyura: Portunidae), from the western Indian Ocean</mods:title>
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<paragraph id="1963FD215F36FFC3FF17DBDFFE13FE56" blockId="1.[151,510,430,490]" box="[151,480,430,456]" pageId="1" pageNumber="220">
<heading id="422B4A4D5F36FFC3FF17DBDFFE13FE56" bold="true" box="[151,480,430,456]" fontSize="11" level="1" pageId="1" pageNumber="220" reason="1">
<emphasis id="2BA821335F36FFC3FF17DBDFFE13FE56" bold="true" box="[151,480,430,456]" pageId="1" pageNumber="220">
<taxonomicName id="DEDC86A25F36FFC3FF17DBDFFE88FE56" ID-CoL="84FXK" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[151,379,430,456]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F36FFC3FF17DBDFFE88FE56" bold="true" box="[151,379,430,456]" italics="true" pageId="1" pageNumber="220">Laleonectes kuriya</emphasis>
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<taxonomicNameLabel id="309B9C485F36FFC3FE03DBDFFE13FE56" box="[387,480,430,456]" pageId="1" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="1963FD215F36FFC3FF17DBA0FE0DFE74" blockId="1.[151,510,430,490]" box="[151,510,465,490]" pageId="1" pageNumber="220">
(
<figureCitation id="81E7E1A45F36FFC3FF1EDBA0FED0FE77" box="[158,291,465,489]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="1" pageNumber="220">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FEAFDBA0FECCFE77" box="[303,319,465,489]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="1" pageNumber="220">2</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FECBDBA0FEA8FE77" box="[331,347,465,489]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="1" pageNumber="220">3</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FEE7DBA0FE5FFE77" box="[359,428,465,489]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="1" pageNumber="220">4E–H</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FE36DBA0FE35FE74" box="[438,454,465,490]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="1" pageNumber="220">5</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FE53DBA0FE05FE74" box="[467,502,465,490]" captionStart="FIGURE 6" captionStartId="7.[151,250,1969,1991]" captionTargetBox="[211,1387,193,1949]" captionTargetId="figure@7.[211,1387,193,1949]" captionTargetPageId="7" captionText="FIGURE 6. Fresh coloration. A, Laleonectes kuriya sp. nov., male (ZRC 2017.0335), Tamil Nadu; B, L. nipponensis (Sakai, 1938), male (RUMF-ZC- 2462), Ryukyu Is.; C, L. nipponensis (Sakai, 1938), male (ZRC 2017.0009), Bohol." httpUri="https://zenodo.org/record/898952/files/figure.png" pageId="1" pageNumber="220">6A</figureCitation>
)
</paragraph>
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<paragraph id="1963FD215F36FFC3FF17D86BFBEFFDD0" blockId="1.[151,1386,537,590]" pageId="1" pageNumber="220">
<treatmentCitationGroup id="39CCDA0F5F36FFC3FF17D86BFBEFFDD0" pageId="1" pageNumber="220">
<taxonomicName id="DEDC86A25F36FFC3FF17D86BFD14FDB1" authority=", Crosnier &amp; Thomassin, 1975: 1106" authorityName=", Crosnier &amp; Thomassin, 1975:" authorityPageNumber="1106" authorityYear="1106" box="[151,743,537,559]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F36FFC3FF17D86BFE80FDB1" box="[151,371,538,559]" italics="true" pageId="1" pageNumber="220">Portunus nipponensis</emphasis>
, 
<treatmentCitation id="987DDB305F36FFC3FEFFD868FD14FDB1" author="Crosnier" box="[383,743,537,559]" page="1106" pageId="1" pageNumber="220" year="1975">
<bibRefCitation id="7D4D80D05F36FFC3FEFFD868FD5AFDB1" author="Crosnier" box="[383,681,537,559]" pageId="1" pageNumber="220" refString="Crosnier, A. &amp; Thomassin, B. (1975) Sur des crabes de la famille des Portunidae (Crustacea Decapoda) nouveaux pour Madagascar ou rares. Bulletin du Museum National d'Histoire Naturelle, 3 e Serie, 241 (Zoologie 165), 1097 - 1118." type="journal article" year="1975">Crosnier &amp; Thomassin, 1975</bibRefCitation>
: 1106
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</taxonomicName>
, fig. 5c, d [
<collectingCountry id="61CBBDB15F36FFC3FCD7D868FC43FDB1" box="[855,944,537,559]" name="Réunion" pageId="1" pageNumber="220">Reunion</collectingCountry>
I.]. Not 
<taxonomicName id="DEDC86A25F36FFC3FB86D86BFA96FDB1" ID-CoL="4747H" authority="Sakai, 1938" authorityName="Sakai" authorityYear="1938" box="[1030,1381,537,559]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F36FFC3FB86D86BFB16FDB1" box="[1030,1253,538,559]" italics="true" pageId="1" pageNumber="220">Neptunus nipponensis</emphasis>
<bibRefCitation id="7D4D80D05F36FFC3FB6BD868FA96FDB1" author="Sakai" box="[1259,1381,537,559]" pageId="1" pageNumber="220" refString="Sakai, T. (1938) On three systematically interesting crabs from Japan, one of Which is neW to science. Annotationes Zoologicae Japonenses, 17 (3), 301 - 309." type="journal article" year="1938">Sakai, 1938</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="DEDC86A25F36FFC3FF17D849FDBAFDD0" ID-CoL="3RYC2" authority=", Poupin, 2008: 76" authorityName="Poupin" authorityPageNumber="76" authorityYear="2008" box="[151,585,568,590]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F36FFC3FF17D849FE7CFDD0" box="[151,399,568,590]" italics="true" pageId="1" pageNumber="220">Laleonectes nipponensis</emphasis>
, 
<treatmentCitation id="987DDB305F36FFC3FE1BD849FDBAFDD0" author="Poupin" box="[411,585,568,590]" page="76" pageId="1" pageNumber="220" year="2008">
<bibRefCitation id="7D4D80D05F36FFC3FE1BD849FDD5FDD0" author="Poupin" box="[411,550,568,590]" pageId="1" pageNumber="220" refString="Poupin, J. (2008) Crustaces de l'ile de la Reunion (Decapodes &amp; Stomatopodes). Rapport scientifique preliminaire de l'Institut de Recherche de l'Ecole Navale, 85 pp." type="book" year="2008">Poupin, 2008</bibRefCitation>
: 76
</treatmentCitation>
</taxonomicName>
[list]. Not 
<taxonomicName id="DEDC86A25F36FFC3FD38D848FBEAFDD0" ID-CoL="4747H" authority="Sakai, 1938" authorityName="Sakai" authorityYear="1938" box="[696,1049,568,590]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F36FFC3FD38D848FC64FDD0" box="[696,919,569,590]" italics="true" pageId="1" pageNumber="220">Neptunus nipponensis</emphasis>
<bibRefCitation id="7D4D80D05F36FFC3FC1DD849FBEAFDD0" author="Sakai" box="[925,1049,568,590]" pageId="1" pageNumber="220" refString="Sakai, T. (1938) On three systematically interesting crabs from Japan, one of Which is neW to science. Annotationes Zoologicae Japonenses, 17 (3), 301 - 309." type="journal article" year="1938">Sakai, 1938</bibRefCitation>
</taxonomicName>
.
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</paragraph>
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<paragraph id="1963FD215F36FFC3FF17D80BFC01FD29" blockId="1.[151,1438,634,2016]" pageId="1" pageNumber="220">
<materialsCitation id="A9B4F77C5F36FFC3FF17D80BFC1DFD29" ID-GBIF-Occurrence="1653530387" collectingDate="1972-12" collectionCode="MALE, NORTH" collectorName="Of LA" country="Réunion" location="ReUNION" pageId="1" pageNumber="220" specimenCode="MNHN-B6019" specimenCount="2" typeStatus="holotype">
<emphasis id="2BA821335F36FFC3FF17D80BFEBBFD0D" bold="true" box="[151,328,634,659]" pageId="1" pageNumber="220">
<typeStatus id="C66743835F36FFC3FF17D80BFF21FD0D" box="[151,210,634,659]" pageId="1" pageNumber="220">Type</typeStatus>
material.
</emphasis>
<typeStatus id="C66743835F36FFC3FED3D80AFE33FD0D" box="[339,448,635,659]" pageId="1" pageNumber="220" type="holotype">HOLOTYPE</typeStatus>
, 
<collectionCode id="7FCD65E45F36FFC3FE4DD80AFDFBFD0D" box="[461,520,635,659]" pageId="1" pageNumber="220">MALE</collectionCode>
, 72.6 [61.0] × 38.4 MM (MNHN-IU-2014-18254 [= 
<specimenCode id="497A555A5F36FFC3FBE5D80BFAEAFD0D" box="[1125,1305,634,659]" collectionCode="MNHN-B" pageId="1" pageNumber="220">MNHN-B6019</specimenCode>
]), 
<collectingCountry id="61CBBDB15F36FFC3FAB6D80AFA6AFD0D" box="[1334,1433,635,659]" name="Réunion" pageId="1" pageNumber="220">RéUNION</collectingCountry>
, 
<collectionCode id="7FCD65E45F36FFC3FF17D8EEFF21FD29" box="[151,210,671,695]" pageId="1" pageNumber="220">NORTH</collectionCode>
<collectorName id="B42998F75F36FFC3FF59D8EFFEE7FD29" box="[217,276,670,695]" pageId="1" pageNumber="220">Of LA</collectorName>
POSSESSION, cRAb POT (‘cASIER’), 250 M, cOLL. P. GUéZé, 
<collectingDate id="7D2622095F36FFC3FCF7D8EEFC1DFD29" box="[887,1006,671,695]" pageId="1" pageNumber="220" value="1972-12">
<date id="6D62DBE15F36FFC3FCF7D8EEFC1DFD29" box="[887,1006,671,695]" pageId="1" pageNumber="220" value="1972-12">DEc. 1972</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="1963FD215F36FFC3FF47D8B3FD07FCBD" blockId="1.[151,1438,634,2016]" pageId="1" pageNumber="220">
<materialsCitation id="A9B4F77C5F36FFC3FF47D8B3FD1CFCBD" ID-GBIF-Occurrence="1653530390" collectingDate="2016-01-06" collectionCode="MALE, WGRC, SOUTHWEST, TRAWL, DEPTH, KUMAR, DEVI" collectorName="STEPHENSON" country="India" latitude="7.75" location="Other" longLatPrecision="1303" longitude="76.96667" municipality="Off" pageId="1" pageNumber="220" specimenCode="ZRC 2017.0335" specimenCount="1" stateProvince="TAMIL NADU">
<emphasis id="2BA821335F36FFC3FF47D8B3FE05FD45" bold="true" box="[199,502,706,731]" pageId="1" pageNumber="220">
<location id="1C03ABFA5F36FFC3FF47D8B3FEFCFD45" LSID="urn:lsid:plazi:treatment:91754C375F36FFCBFF17DBDFFCA5FF69:1C03ABFA5F36FFC3FF47D8B3FEFCFD45" box="[199,271,706,731]" country="India" latitude="7.75" longLatPrecision="1303" longitude="76.96667" municipality="Off" name="Other" pageId="1" pageNumber="220" stateProvince="TAMIL NADU">Other</location>
material examined
</emphasis>
. 1 
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, 59.2 [51.5] × 33.1 MM (
<specimenCode id="497A555A5F36FFC3FCF1D8B2FBDEFD45" box="[881,1069,706,731]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="220">ZRC 2017.0335</specimenCode>
), 1 
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, 58.7 [48.7] × 29.7 MM (ZSI/ 
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/IR-INV.8236), 
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MUTTAM cOAST, 
<collectingRegion id="DB1833C35F36FFC3FD2CD896FCC6FD61" box="[684,821,743,767]" country="India" name="Tamil Nadu" pageId="1" pageNumber="220">TAMIL NADU</collectingRegion>
, 
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<collectingCountry id="61CBBDB15F36FFC3FC39D896FC00FD61" box="[953,1011,743,767]" name="India" pageId="1" pageNumber="220">INDIA</collectingCountry>
(
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, 
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), 
<collectionCode id="7FCD65E45F36FFC3FB63D896FAD2FD61" box="[1251,1313,743,767]" pageId="1" pageNumber="220">TRAWL</collectionCode>
, cA. 100 M 
<collectionCode id="7FCD65E45F36FFC3FF17D97AFF2AFCBC" box="[151,217,779,802]" pageId="1" pageNumber="220">DEPTH</collectionCode>
, cOLL. A.B. 
<collectionCode id="7FCD65E45F36FFC3FED7D97AFE56FCBD" box="[343,421,779,803]" pageId="1" pageNumber="220">KUMAR</collectionCode>
&amp; 
<collectorName id="B42998F75F36FFC3FE48D97AFE06FCBD" box="[456,501,779,803]" name="STEPHENSON" pageId="1" pageNumber="220">S.S.</collectorName>
<collectionCode id="7FCD65E45F36FFC3FE7DD97AFDC4FCBC" box="[509,567,779,802]" pageId="1" pageNumber="220">DEVI</collectionCode>
, 
<collectingDate id="7D2622095F36FFC3FDC1D97BFD1CFCBD" box="[577,751,778,803]" pageId="1" pageNumber="220" value="2016-01-06">
<date id="6D62DBE15F36FFC3FDC1D97BFD1CFCBD" box="[577,751,778,803]" pageId="1" pageNumber="220" value="2016-01-06">6 JANUARY 2016</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="1963FD215F36FFC3FF47D95FFE7CFBDD" blockId="1.[151,1438,634,2016]" pageId="1" pageNumber="220">
<materialsCitation id="A9B4F77C5F36FFC3FF47D95FFA6AFCF4" ID-GBIF-Occurrence="1653530323" collectionCode="SAKAI, MALE" country="Japan" location="JAPAN" pageId="1" pageNumber="220" specimenCount="1">
<emphasis id="2BA821335F36FFC3FF47D95FFE2BFCD9" bold="true" box="[199,472,814,839]" pageId="1" pageNumber="220">Comparative material.</emphasis>
<taxonomicName id="DEDC86A25F36FFC3FE61D95FFC6AFCD9" authority="(SAKAI, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[481,921,814,839]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="224" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F36FFC3FE61D95FFD07FCD8" box="[481,756,814,838]" italics="true" pageId="1" pageNumber="220">Laleonectes nipponensis</emphasis>
(
<bibRefCitation id="7D4D80D05F36FFC3FC83D95EFC62FCD9" author="SAKAI" box="[771,913,815,839]" pageId="1" pageNumber="220" refString="Sakai, T. (1938) On three systematically interesting crabs from Japan, one of Which is neW to science. Annotationes Zoologicae Japonenses, 17 (3), 301 - 309." type="journal article" year="1938">
<collectionCode id="7FCD65E45F36FFC3FC83D95EFCBBFCD9" box="[771,840,815,839]" pageId="1" pageNumber="220">SAKAI</collectionCode>
, 1938
</bibRefCitation>
)
</taxonomicName>
: 1 
<collectionCode id="7FCD65E45F36FFC3FC3CD95EFC04FCD9" box="[956,1015,815,839]" pageId="1" pageNumber="220">MALE</collectionCode>
, 65.8 [54.3] × 34.9 MM (RUMF-ZC- 2462), SUbMARINE cAVE (AKUMANO-YAKATA), SHIMOJI I., MIYAKO ISLAND-GROUP, RYUKYU IS., 
<collectingCountry id="61CBBDB15F36FFC3FB1FD922FB10FCF5" box="[1183,1251,851,875]" name="Japan" pageId="1" pageNumber="220">JAPAN</collectingCountry>
, ~30 M, cOLL. Y
</materialsCitation>
. FUJITA, 
<date id="6D62DBE15F36FFC3FF67D906FE88FC11" box="[231,379,887,911]" pageId="1" pageNumber="220" value="2013-01-27">27 JAN. 2013</date>
; 
<materialsCitation id="A9B4F77C5F36FFC3FE07D906FBDDFC2D" ID-GBIF-Occurrence="1653530325" collectingDate="2012-03-01" collectionCode="NEAR, DREAM, HALL, ONNA, MAR" country="Japan" location="Off" pageId="1" pageNumber="220" specimenCount="1" stateProvince="OKINAWA">
1 fEMALE, 42.2 [32.2] × 20.2 MM (RUMF-ZC-4703), SUbMARINE cAVE 
<collectionCode id="7FCD65E45F36FFC3FB18D906FB3BFC11" box="[1176,1224,887,911]" pageId="1" pageNumber="220">NEAR</collectionCode>
“ 
<collectionCode id="7FCD65E45F36FFC3FB58D906FA99FC11" box="[1240,1386,887,911]" pageId="1" pageNumber="220">DREAM HALL</collectionCode>
”, 
<location id="1C03ABFA5F36FFC3FAFCD907FA6FFC11" LSID="urn:lsid:plazi:treatment:91754C375F36FFCBFF17DBDFFCA5FF69:1C03ABFA5F36FFC3FAFCD907FA6FFC11" box="[1404,1436,886,911]" country="Japan" name="Off" pageId="1" pageNumber="220" stateProvince="OKINAWA">Off</location>
<collectionCode id="7FCD65E45F36FFC3FF17D9EAFF27FC2D" box="[151,212,923,947]" pageId="1" pageNumber="220">ONNA</collectionCode>
VILLAgE, 
<collectingRegion id="DB1833C35F36FFC3FEBDD9EAFE51FC2D" box="[317,418,923,947]" country="Japan" name="Okinawa" pageId="1" pageNumber="220">OKINAWA</collectingRegion>
I., RYUKYU IS., 
<collectingCountry id="61CBBDB15F36FFC3FDD6D9EAFD69FC2D" box="[598,666,923,947]" name="Japan" pageId="1" pageNumber="220">JAPAN</collectingCountry>
, cOLL. T. NARUSE 
<emphasis id="2BA821335F36FFC3FCD5D9EAFC74FC2D" box="[853,903,923,947]" italics="true" pageId="1" pageNumber="220">et al</emphasis>
., 
<collectingDate id="7D2622095F36FFC3FC1FD9EAFBDDFC2D" box="[927,1070,923,947]" pageId="1" pageNumber="220" value="2012-03-01">
<date id="6D62DBE15F36FFC3FC1FD9EAFBDDFC2D" box="[927,1070,923,947]" pageId="1" pageNumber="220" value="2012-03-01">
1 
<collectionCode id="7FCD65E45F36FFC3FC36D9EAFC1BFC2D" box="[950,1000,923,947]" country="Zimbabwe" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="1" pageNumber="220">MAR</collectionCode>
. 2012
</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="A9B4F77C5F36FFC3FBBBD9EAFB57FC49" ID-GBIF-Occurrence="1653530326" collectionCode="MALE, LATERAL, SPINE, TOWN, BOHOL" country="Japan" location="BOHOL PROVINcE" pageId="1" pageNumber="220" specimenCode="ZRC 2008.0101" specimenCount="1" stateProvince="BOHOL PROVINcE">
1 
<collectionCode id="7FCD65E45F36FFC3FBD2D9EAFB7EFC2D" box="[1106,1165,923,947]" pageId="1" pageNumber="220">MALE</collectionCode>
, 52.4 [47.2] × 31.4 MM (LEfT 
<collectionCode id="7FCD65E45F36FFC3FF4DD9CEFEA6FC49" box="[205,341,959,983]" pageId="1" pageNumber="220">LATERAL SPINE</collectionCode>
bROKEN Off) (
<specimenCode id="497A555A5F36FFC3FE6AD9CEFD57FC49" box="[490,676,959,983]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="220">ZRC 2008.0101</specimenCode>
), BALIcASAg I., PANgLAO 
<collectionCode id="7FCD65E45F36FFC3FC2CD9CEFC1BFC49" box="[940,1000,959,983]" pageId="1" pageNumber="220">TOWN</collectionCode>
, 
<collectingRegion id="DB1833C35F36FFC3FC72D9CEFB57FC49" box="[1010,1188,959,983]" country="Philippines" name="Bohol" pageId="1" pageNumber="220">
<collectionCode id="7FCD65E45F36FFC3FC72D9CEFBC5FC49" box="[1010,1078,959,983]" pageId="1" pageNumber="220">BOHOL</collectionCode>
PROVINcE
</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="A9B4F77C5F36FFC3FB2ED9CEFCCAFC65" ID-GBIF-Occurrence="1653530327" collectingDate="2003-07-27" collectionCode="PHILIPPINES, IN, NETS" country="Philippines" location="Ng" pageId="1" pageNumber="220" specimenCount="1">
<collectionCode id="7FCD65E45F36FFC3FB2ED9CEFAC3FC49" box="[1198,1328,959,983]" pageId="1" pageNumber="220">
<collectingCountry id="61CBBDB15F36FFC3FB2ED9CEFAC3FC49" box="[1198,1328,959,983]" name="Philippines" pageId="1" pageNumber="220">PHILIPPINES</collectingCountry>
</collectionCode>
, 
<collectionCode id="7FCD65E45F36FFC3FABBD9CEFAA2FC48" box="[1339,1361,959,982]" pageId="1" pageNumber="220">IN</collectionCode>
TANgLE 
<collectionCode id="7FCD65E45F36FFC3FF17D992FF3BFC65" box="[151,200,995,1019]" pageId="1" pageNumber="220">NETS</collectionCode>
, fROM LOcAL fISHERMEN, cOLL. P.K.L. 
<location id="1C03ABFA5F36FFC3FDEED992FD60FC65" LSID="urn:lsid:plazi:treatment:91754C375F36FFCBFF17DBDFFCA5FF69:1C03ABFA5F36FFC3FDEED992FD60FC65" box="[622,659,995,1019]" country="Philippines" name="Ng" pageId="1" pageNumber="220">Ng</location>
, 
<collectingDate id="7D2622095F36FFC3FD23D992FCCAFC65" box="[675,825,995,1019]" pageId="1" pageNumber="220" value="2003-07-27">
<date id="6D62DBE15F36FFC3FD23D992FCCAFC65" box="[675,825,995,1019]" pageId="1" pageNumber="220" value="2003-07-27">27 JUL. 2003</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="A9B4F77C5F36FFC3FCCAD992FD63FB81" ID-GBIF-Occurrence="1653530330" collectionCode="MALE, TOWN, BOHOL" country="Philippines" location="BOHOL PROVINcE" pageId="1" pageNumber="220" specimenCode="ZRC 2017.0009" specimenCount="1" stateProvince="BOHOL PROVINcE">
1 
<collectionCode id="7FCD65E45F36FFC3FCE4D992FC6CFC65" box="[868,927,995,1019]" pageId="1" pageNumber="220">MALE</collectionCode>
, 51.1 [42.8] × 27.9 MM (
<specimenCode id="497A555A5F36FFC3FB51D992FA62FC65" box="[1233,1425,995,1019]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="220">ZRC 2017.0009</specimenCode>
), BALIcASAg I., PANgLAO 
<collectionCode id="7FCD65E45F36FFC3FE13DE76FE3CFB81" box="[403,463,1031,1055]" pageId="1" pageNumber="220">TOWN</collectionCode>
, 
<collectingRegion id="DB1833C35F36FFC3FE5CDE76FD63FB81" box="[476,656,1031,1055]" country="Philippines" name="Bohol" pageId="1" pageNumber="220">
<collectionCode id="7FCD65E45F36FFC3FE5CDE76FDD3FB81" box="[476,544,1031,1055]" pageId="1" pageNumber="220">BOHOL</collectionCode>
PROVINcE
</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="A9B4F77C5F36FFC3FD1DDE76FE78FBDD" ID-GBIF-Occurrence="1653530332" collectingDate="2002-06" collectionCode="PHILIPPINES, IN, NETS, JUNE" country="Philippines" location="Ng" pageId="1" pageNumber="220" specimenCount="1">
<collectionCode id="7FCD65E45F36FFC3FD1DDE76FCECFB81" box="[669,799,1031,1055]" pageId="1" pageNumber="220">
<collectingCountry id="61CBBDB15F36FFC3FD1DDE76FCECFB81" box="[669,799,1031,1055]" name="Philippines" pageId="1" pageNumber="220">PHILIPPINES</collectingCountry>
</collectionCode>
, 
<collectionCode id="7FCD65E45F36FFC3FCACDE76FCB1FB80" box="[812,834,1031,1054]" pageId="1" pageNumber="220">IN</collectionCode>
TANgLE 
<collectionCode id="7FCD65E45F36FFC3FC1ADE76FC38FB81" box="[922,971,1031,1055]" pageId="1" pageNumber="220">NETS</collectionCode>
, 200–300 M, fROM LOcAL fISHERMEN, cOLL. P.K.L. 
<location id="1C03ABFA5F36FFC3FF64DE5AFEFAFBDD" LSID="urn:lsid:plazi:treatment:91754C375F36FFCBFF17DBDFFCA5FF69:1C03ABFA5F36FFC3FF64DE5AFEFAFBDD" box="[228,265,1067,1091]" country="Philippines" name="Ng" pageId="1" pageNumber="220">Ng</location>
, 
<collectingDate id="7D2622095F36FFC3FE95DE5AFE78FBDD" box="[277,395,1067,1091]" pageId="1" pageNumber="220" value="2002-06">
<date id="6D62DBE15F36FFC3FE95DE5AFE78FBDD" box="[277,395,1067,1091]" pageId="1" pageNumber="220" value="2002-06">
<collectionCode id="7FCD65E45F36FFC3FE95DE5AFEBBFBDC" box="[277,328,1067,1090]" pageId="1" pageNumber="220">JUNE</collectionCode>
2002
</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="51C6AEAA5F36FFCAFF47DE3FFD99FF66" lastPageId="8" lastPageNumber="227" pageId="1" pageNumber="220" type="description">
<paragraph id="1963FD215F36FFC3FF47DE3FFAEBF80B" blockId="1.[151,1438,634,2016]" pageId="1" pageNumber="220">
<emphasis id="2BA821335F36FFC3FF47DE3FFEAAFBF9" bold="true" box="[199,345,1102,1127]" pageId="1" pageNumber="220">Description.</emphasis>
CARAPAcE (
<figureCitation id="81E7E1A45F36FFC3FE58DE3EFDA1FBF8" box="[472,594,1103,1127]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="1" pageNumber="220">FIg. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FDDEDE3EFD71FBF9" box="[606,642,1103,1127]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="1" pageNumber="220">2A</figureCitation>
) TRANSVERSELY SUbOVATE, DISTINcTLY bROADER THAN LONg (CW max/CL = 1.8– 1.9; CW pen/CL = 1.6); REgIONS WELL DEfINED, SEPARATED bY DEEP, SMOOTH gROOVES; DORSAL SURfAcE cONVEX, gRANULATE, WITH LARgER gRANULES ON ANTERIOR HALf. FRONT qUADRIDENTATE, MESIAL PAIR Of TEETH TRIANgULAR WITH ROUNDED TIPS, SEPARATED bY V-SHAPED cLEfT, LATERAL TEETH MORE bROADLY TRIANgULAR, MESIAL MARgINS LONgER THAN LATERAL, SEPARATED fROM ADJAcENT MESIAL TOOTH bY A RELATIVELY WIDER, V-SHAPED cLEfT. SUPRAORbITAL TOOTH [INNER SUPRAORbITAL LObE] PROMINENT, bROADLY TRIANgULAR WITH DORSAL RIDgE, SEPARATED fROM fRONT bY bROAD DEPRESSION; SUPRAORbITAL MARgIN cONcAVE, gRANULATE, WITH TWO NARROW cLEfTS, LATERAL cLEfT SEPARATINg REST Of SUPRAORbITAL MARgIN fROM EXORbITAL TOOTH; INfRAORbITAL MARgIN (
<figureCitation id="81E7E1A45F36FFC3FEFADF21FE07FAF6" box="[378,500,1360,1384]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="1" pageNumber="220">FIg. 2A, C</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FD80DF21FDD7FAF6" box="[512,548,1360,1384]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="1" pageNumber="220">3A</figureCitation>
) gRANULATE, PROJEcTINg ANTERIORLY, VISIbLE fROM DORSAL VIEW, LATERAL JUNcTION WITH EXORbITAL TOOTH MARKED bY DEEP NARROW cLEfT, bROAD V-SHAPED cLEfT SEPARATINg LARgE TRIANgULAR INfRAORbITAL TOOTH fROM SMALLER TOOTH AND REST Of MARgIN. ANTEROLATERAL MARgIN (
<figureCitation id="81E7E1A45F36FFC3FCD5DFE9FC43FA2E" box="[853,944,1432,1456]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="1" pageNumber="220">FIg. 2A</figureCitation>
) SLIgHTLY cONVEX, ARMED WITH NINE DISTINcT, TRIANgULAR TEETH WITH SHARP APIcES, WITH 1st [EXORbITAL] TOOTH SMALLEST, NARROWEST, AND 9th TOOTH LONgEST, MOST PROMINENT, LATERALLY DIREcTED TIP fORMINg MAXIMUM EXTENT Of cARAPAcE WIDTH, MARKINg JUNcTION WITH POSTEROLATERAL MARgIN. APEX Of 2nd ANTEROLATERAL TOOTH LEVEL WITH OR SLIgHTLY ADVANcED RELATIVE TO THAT Of 1st, IN DORSAL VIEW. IN 
<typeStatus id="C66743835F36FFC3FF17DC40FF0EF9D7" box="[151,253,1585,1609]" pageId="1" pageNumber="220" type="holotype">HOLOTYPE</typeStatus>
, RIgHT 3rd, 4th TEETH MOSTLY fUSED EXcEPT AT APIcES. POSTEROLATERAL MARgIN SLIgHTLY cONcAVE, cONVERgENT POSTERIORLY, LINED WITH TWO SEPARATE ROWS Of SMALL gRANULES; JUNcTION WITH POSTERIOR MARgIN MARKED bY SMALL, SHARP- TIPPED, DORSALLY DIREcTED, cURVED TOOTH (
<figureCitation id="81E7E1A45F36FFC3FDECDC08FD3FF90F" box="[620,716,1657,1681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="1" pageNumber="220">FIg. 2D</figureCitation>
). POSTERIOR MARgIN SINUOUS, LINED WITH SMALL gRANULES. LONg, TRANSVERSELY SIgMOID gRANULAR RIDgE ON EAcH EPIbRANcHIAL REgION, EXTENDINg fROM bASE Of 9th TOOTH AND TERMINATINg JUST bEfORE cERVIcAL gROOVE; PAIRED, SHORT, HORIZONTAL gRANULAR RIDgES ON METAgASTRIc REgION. GRANULAR DENTIfORM PROJEcTIONS PRESENT ON SOME REgIONS: TWO ARRANgED SIDE-bY-SIDE ON EAcH PROTOgASTRIc REgION; ONE ON MESIAL SIDE Of EAcH EPIbRANcHIAL REgION (AT MESIAL TERMINUS Of EPIbRANcHIAL RIDgE); TWO ON RAISED PORTIONS Of EAcH MESObRANcHIAL REgION; TWO ON cARDIAc REgION. VENTRALLY, SUbHEPATIc REgION AND PTERYgOSTOME (
<figureCitation id="81E7E1A45F36FFC3FB90DD41FB94F8D6" box="[1040,1127,1840,1864]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="1" pageNumber="220">FIg. 3B</figureCitation>
) SMOOTH TO fINELY gRANULAR, WITH 26 OR 27 ObLONg TUbERcLES fANNINg AcROSS fROM 2nd TO 7th ANTEROLATERAL TEETH Of THE cARAPAcE, DEcREASINg IN SIZE TOWARDS THE 9th ANTEROLATERAL TOOTH, bEcOMINg NEARLY INDISTINgUISHAbLE fROM OTHER SURROUNDINg gRANULES.
</paragraph>
<paragraph id="1963FD215F36FFC0FF47DDD0FD19FE31" blockId="1.[151,1438,634,2016]" lastBlockId="2.[151,1438,151,431]" lastPageId="2" lastPageNumber="221" pageId="1" pageNumber="220">
EYES (
<figureCitation id="81E7E1A45F36FFC3FE90DDD0FE85F827" box="[272,374,1953,1977]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="1" pageNumber="220">FIgS. 2C</figureCitation>
, 
<figureCitation id="81E7E1A45F36FFC3FE06DDD0FE5AF827" box="[390,425,1953,1977]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="1" pageNumber="220">3A</figureCitation>
) WELL DEVELOPED, cORNEAS LARgE. ANTENNULES (
<figureCitation id="81E7E1A45F36FFC3FC38DDD0FBE7F827" box="[952,1044,1953,1977]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="1" pageNumber="220">FIg. 3A</figureCitation>
) fOLDINg TRANSVERSELY AND SLIgHTLY ObLIqUELY, 2nd AND 3rd SEgMENTS LONg, SLENDER. BASAL ANTENNAL ARTIcLE (
<figureCitation id="81E7E1A45F36FFC3FC29DDB9FBF7F87E" box="[937,1028,1992,2016]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="1" pageNumber="220">FIg. 3A</figureCitation>
) LONgITUDINALLY REcTANgULAR, SIMPLE, LODgED IN ORbITAL HIATUS, IMMObILE; fLAgELLUM RELATIVELY LONg, TIP REAcHINg cARAPAcE 2nd ANTEROLATERAL TOOTH WHEN EXTENDED. BUccAL cAVITY qUADRATE, LATERAL MARgINS PARALLEL; POSTERIOR MARgIN Of EPISTOME WITH SMALL TRIANgULAR PROJEcTION, bROADLY cONcAVE ON EITHER SIDE; ENDOSTOME WITH PAIR Of STRONg RIDgES. ENDOPOD Of fIRST MAXILLIPED, bROADENINg INTO SUbTRIANgULAR LObE DISTALLY, WITH SLENDER, SUb-ELLIPTIcAL AccESSORY LObE MESIALLY. THIRD MAXILLIPEDS (
<figureCitation id="81E7E1A45F35FFC0FF1FDB5AFF05FEDD" box="[159,246,299,323]" captionStart="FIGURE 3" captionStartId="4.[151,250,1920,1942]" captionTargetBox="[165,1429,193,1907]" captionTargetId="figure@4.[165,1429,193,1907]" captionTargetPageId="4" captionText="FIGURE 3. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm." httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="2" pageNumber="221">FIg. 3C</figureCitation>
) bROAD, fORMINg NARROW HIATUS WHEN cLOSED; IScHIUM SUbREcTANgULAR, WITH RELATIVELY SHALLOW bUT DISTINcT LONgITUDINAL MEDIAN SULcUS; MERUS RHOMbOIDAL, DISTO-LATERAL ANgLE PROJEcTINg, ROUNDED; EXOPOD STOUT, SUbREcTANgULAR, DISTAL END REAcHINg DISTO-LATERAL ANgLE Of MERUS, LATERAL MARgIN STRAIgHT, MESIAL MARgIN cONVEX PROXIMALLY, WITH SUbTRIANgULAR PROJEcTION SUbDISTALLY.
</paragraph>
<caption id="4DA3ADA95F35FFC0FF17DF3BFA60FA03" httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="2" pageNumber="221" targetBox="[151,1436,519,1290]" targetPageId="2">
<paragraph id="1963FD215F35FFC0FF17DF3BFA60FA03" blockId="2.[151,1436,1354,1437]" pageId="2" pageNumber="221">
<emphasis id="2BA821335F35FFC0FF17DF3BFEE0FAC1" bold="true" box="[151,275,1354,1376]" pageId="2" pageNumber="221">FIGURE 1.</emphasis>
Dorsal habitus. 
<taxonomicName id="DEDC86A25F35FFC0FE39DF3BFD89FAFE" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[441,634,1354,1376]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="221" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F35FFC0FE39DF3BFD89FAFE" box="[441,634,1354,1376]" italics="true" pageId="2" pageNumber="221">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F35FFC0FD00DF3BFD22FAFE" box="[640,721,1354,1376]" pageId="2" pageNumber="221" rank="species">
<emphasis id="2BA821335F35FFC0FD00DF3BFD38FAC1" bold="true" box="[640,715,1354,1375]" pageId="2" pageNumber="221">sp. nov</emphasis>
.
</taxonomicNameLabel>
(A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU-2014-18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. 
<taxonomicName id="DEDC86A25F35FFC0FC27DF19FADCFAE0" authority="(Sakai, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[935,1327,1384,1406]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="221" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F35FFC0FC27DF19FB53FAE0" box="[935,1184,1384,1406]" italics="true" pageId="2" pageNumber="221">Laleonectes nipponensis</emphasis>
(Sakai, 1938)
</taxonomicName>
(C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC-2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm.
</paragraph>
</caption>
<paragraph id="1963FD215F35FFC0FF47DFB4FB75F963" blockId="2.[151,1437,1476,2005]" pageId="2" pageNumber="221">
MALE THORAcIc STERNUM (
<figureCitation id="81E7E1A45F35FFC0FE5ADFB4FDC1FA43" box="[474,562,1477,1501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="2" pageNumber="221">FIg. 2B</figureCitation>
) bROAD, ANTERIOR THIRD fINELY gRANULAR, POSTERIOR TWO-THIRDS SMOOTH. THORAcIc STERNITES 1, 2 fUSED, fORMINg bROAD TRIANgLE. THORAcIc STERNITE 3 SEPARATED fROM STERNITE 2 bY cOMPLETE SUTURE, 
<materialsCitation id="A9B4F77C5F35FFC0FF17DC7CFAAFF9D7" ID-GBIF-Occurrence="1653530334" collectionCode="ANTERIOR, POSTERIOR, WITH, LATERAL, V-SHAPED" country="American Samoa" location="AS" pageId="2" pageNumber="221" specimenCode="STERNITE 4, STERNITE 3, SUTURE 3" specimenCount="1">
<collectionCode id="7FCD65E45F35FFC0FF17DC7CFF1DF9BB" box="[151,238,1549,1573]" pageId="2" pageNumber="221">ANTERIOR</collectionCode>
MARgIN Of fORMER RAISED SLIgHTLY AbOVE 
<collectionCode id="7FCD65E45F35FFC0FD39DC7CFCEFF9BB" box="[697,796,1549,1573]" pageId="2" pageNumber="221">POSTERIOR</collectionCode>
MARgIN Of LATTER. THORAcIc 
<specimenCode id="497A555A5F35FFC0FBD2DC7CFB48F9BB" box="[1106,1211,1549,1573]" collectionCode="STERNITE" pageId="2" pageNumber="221">STERNITE 4</specimenCode>
INcOMPLETELY fUSED 
<collectionCode id="7FCD65E45F35FFC0FF17DC40FF3AF9D6" box="[151,201,1585,1608]" pageId="2" pageNumber="221">WITH</collectionCode>
<specimenCode id="497A555A5F35FFC0FF54DC40FEB1F9D7" box="[212,322,1585,1609]" collectionCode="STERNITE" pageId="2" pageNumber="221">STERNITE 3</specimenCode>
, REMNANTS Of 
<specimenCode id="497A555A5F35FFC0FE67DC40FDB0F9D7" box="[487,579,1585,1609]" collectionCode="SUTURE" pageId="2" pageNumber="221">SUTURE 3</specimenCode>
/4 LIMITED TO 
<collectionCode id="7FCD65E45F35FFC0FD63DC40FCDAF9D7" box="[739,809,1585,1609]" pageId="2" pageNumber="221">LATERAL</collectionCode>
EDgES, cONTINUINg MESIALLY 
<collectingCountry id="61CBBDB15F35FFC0FBF4DC40FB78F9D7" box="[1140,1163,1585,1609]" name="American Samoa" pageId="2" pageNumber="221">AS</collectingCountry>
<collectionCode id="7FCD65E45F35FFC0FB17DC40FAF3F9D7" box="[1175,1280,1585,1609]" pageId="2" pageNumber="221">V-SHAPED</collectionCode>
fURROW
</materialsCitation>
; 
<materialsCitation id="A9B4F77C5F35FFC0FAEADC40FABAF9F2" ID-GBIF-Occurrence="1653530336" collectionCode="WITH, MEDIAN, EXPOSED, INTO, LINE" country="American Samoa" location="AS" pageId="2" pageNumber="221" specimenCount="1">
<collectionCode id="7FCD65E45F35FFC0FAEADC40FF19F9F3" pageId="2" pageNumber="221">WITH MEDIAN</collectionCode>
LONgITUDINAL gROOVE ON 
<collectionCode id="7FCD65E45F35FFC0FD85DC24FD92F9F3" box="[517,609,1621,1645]" pageId="2" pageNumber="221">EXPOSED</collectionCode>
REgION, cONTINUINg 
<collectionCode id="7FCD65E45F35FFC0FCC7DC24FC80F9F3" box="[839,883,1621,1645]" pageId="2" pageNumber="221">INTO</collectionCode>
STERNO-AbDOMINAL cAVITY 
<collectingCountry id="61CBBDB15F35FFC0FB1BDC24FB41F9F3" box="[1179,1202,1621,1645]" name="American Samoa" pageId="2" pageNumber="221">AS</collectingCountry>
<collectionCode id="7FCD65E45F35FFC0FB3DDC24FABAF9F2" box="[1213,1353,1621,1645]" pageId="2" pageNumber="221">MEDIAN LINE</collectionCode>
</materialsCitation>
; LATERAL MARgINS cONVEX, gRANULATE. THORAcIc STERNITE 5 WITH PROMINENT TUbERcLE NEAR SUTURE 5/6, fORMINg STERNO-AbDOMINAL LOcKINg MEcHANISM (PRESS-bUTTON). PART Of THORAcIc STERNITE 8 EXPOSED EVEN WITH PLEON fOLDED AgAINST THORAcIc STERNUM. SUTURES 4/5, 5/6 INTERRUPTED MEDIALLY, SUTURES 6/7, 7/8 cOMPLETE, JOININg MEDIALLY. MEDIAN LINE PRESENT IN STERNOAbDOMINAL cAVITY AT LEVEL Of STERNITES 4, 7, AND 8, AbSENT AT LEVEL Of STERNITES 5 AND 6.
</paragraph>
<paragraph id="1963FD215F35FFC0FF47DD78FDA8F84B" blockId="2.[151,1437,1476,2005]" pageId="2" pageNumber="221">
MALE PLEON (
<figureCitation id="81E7E1A45F35FFC0FED5DD78FE5EF8BF" box="[341,429,1801,1825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="2" pageNumber="221">FIg. 2B</figureCitation>
, 
<figureCitation id="81E7E1A45F35FFC0FE39DD78FE2AF8BF" box="[441,473,1801,1825]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="2" pageNumber="221">4E</figureCitation>
) NARROWLY TRIANgULAR; PLEOMERE 1 SMALL, THIN, PARTLY bURIED bENEATH POSTERIOR cARAPAcE MARgIN; PLEOMERE 2 SHORT, bROAD, WITH TRANSVERSE RIDgE THROUgHOUT bREADTH; PLEOMERES 3–5 fUSED, WITH NOTcH ON LATERAL MARgIN DEMARcATINg JUNcTION Of PLEOMERE 3 AND 4, WITH TRANSVERSE RIDgE AT LEVEL Of PLEOMERE 3, cOMbINED LATERAL MARgINS Of PLEOMERES 4, 5 NEARLY STRAIgHT; PLEOMERE 6 TRAPEZOIDAL, bASAL WIDTH 1.7 TIMES MEDIAN HEIgHT, LATERAL MARgIN NEARLY STRAIgHT. TELSON SUbTRIANgULAR, MEDIAN HEIgHT NEARLY EqUAL TO bASAL WIDTH, TIP ROUNDED, LATERAL MARgINS NEARLY STRAIgHT, SLIgHTLY cONVEX.
</paragraph>
<caption id="4DA3ADA95F34FFC1FF17DC49FA96F915" httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="3" pageNumber="222" targetBox="[151,1436,193,1571]" targetPageId="3">
<paragraph id="1963FD215F34FFC1FF17DC49FA96F915" blockId="3.[151,1436,1592,1675]" pageId="3" pageNumber="222">
<emphasis id="2BA821335F34FFC1FF17DC49FEE7F9D3" bold="true" box="[151,276,1592,1614]" pageId="3" pageNumber="222">FIGURE 2.</emphasis>
<taxonomicName id="DEDC86A25F34FFC1FE9DDC49FE2CF9D0" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[285,479,1592,1614]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="222" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F34FFC1FE9DDC49FE2CF9D0" box="[285,479,1592,1614]" italics="true" pageId="3" pageNumber="222">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F34FFC1FE67DC48FDC9F9D0" box="[487,570,1593,1614]" pageId="3" pageNumber="222" rank="species">
<emphasis id="2BA821335F34FFC1FE67DC48FDC7F9D0" bold="true" box="[487,564,1593,1614]" pageId="3" pageNumber="222">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype, male, 72.6 × 38.4 mm (MNHN-IU-2014-18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm.
</paragraph>
</caption>
<paragraph id="1963FD215F34FFC7FF47DCC8FAFFF874" blockId="3.[151,1437,1720,2033]" lastBlockId="5.[151,1437,1677,2026]" lastPageId="5" lastPageNumber="224" pageId="3" pageNumber="222">
CHELIPEDS (
<figureCitation id="81E7E1A45F34FFC1FEC7DCC8FE3CF94E" box="[327,463,1721,1745]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="3" pageNumber="222">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F34FFC1FE5DDCC8FDD2F94F" box="[477,545,1721,1745]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="3" pageNumber="222">2E–H</figureCitation>
, 
<figureCitation id="81E7E1A45F34FFC1FDAEDCC8FDBEF94F" box="[558,589,1721,1745]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="3" pageNumber="222">4F</figureCitation>
) SUbEqUAL, ELONgATED, RELATIVELY STOUT. BASIS fUSED WITH IScHIUM, WITH LOW, TUbERcULIfORM PROJEcTION ON ANTERO-DISTAL ANgLE. MERUS RELATIVELY STOUT, ANTERIOR (fLEXOR) MARgIN WITH THREE PROMINENT, cURVED SPINES, REST Of MARgIN LINED WITH LARgE gRANULES; POSTERIOR (EXTENSOR) MARgIN LINED WITH SHARP gRANULES, DISTAL TIP WITH PROMINENT, SHARP SPINE; DORSAL SURfAcE WITH RIDgE cOMPOSED Of 10 STRIDULATORY TUbERcLES (
<figureCitation id="81E7E1A45F34FFC1FF1FDD38FF06F8FF" box="[159,245,1865,1889]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="3" pageNumber="222">FIg. 4F</figureCitation>
) NEAR PROXIMAL END, ARRANgED DIAgONALLY fROM fLEXOR MARgIN Of MERUS (AT bASE Of MOST PROXIMAL SPINE), EAcH TUbERcLE ObLONg, bISEcTED bY NARROW SLIT (NOTE: THIS RIDgE MATcHES NEATLY WITH LONgER RIDgE Of TUbERcLES ON SUbHEPATIc REgION Of cARAPAcE; RATcHETINg SOUND cAN bE PRODUcED fROM SPEcIMEN bY ScRAPINg ARTIcULATED MERUS AcROSS SUbHEPATIc REgION). CARPUS WITH DISTINcT SPINE ON INNER DISTAL ANgLE, DORSO-EXTERNAL SURfAcE WITH TWO SPINES (ONE bIg, ONE SMALL) AND TWO LONgITUDINAL ROWS Of SHARP gRANULES. PALM Of cHELA (
<figureCitation id="81E7E1A45F34FFC1FBC2DDA8FB49F86F" box="[1090,1210,2009,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="3" pageNumber="222">FIg. 2E–H</figureCitation>
) ELONgATE; EXTERNAL SURfAcE WITH THREE LONgITUDINAL cRESTS, LOWEST cREST SURMOUNTED bY ROW Of SMALL ROUND gRANULES, UPPER AND MIDDLE SURMOUNTED bY ROW Of LARgER, SHARP gRANULES, LARgE bROAD SPINE ON POINT Of ARTIcULATION WITH cARPUS, MARKINg PROXIMAL END Of MIDDLE LONgITUDINAL cREST, SMALLER bLUNT SPINE ON DISTAL END Of UPPERMOST cREST; UPPER SURfAcE Of PALM WITH SHARP, PROMINENT cREST fORMINg INNER MARgIN, TERMINATINg DISTALLY IN SHARP SPINE, WITH SUb-MARgINAL ROW Of gRANULES; INTERNAL SURfAcE Of PALM WITH TWO SMOOTH LONgITUDINAL cRESTS, ONE MEDIALLY POSITIONED AND ONE PLAcED LOWER cONTINUINg fAINTLY INTO fIXED fINgER. FINgERS LONg, SLENDER, cURVED AND POINTED AT TIPS; EXTERNAL AND INTERNAL SURfAcES cHANNELLED WITH gROOVES; cUTTINg MARgINS LINED WITH LARgE, bROAD-bASED, POINTED TEETH INTERSPERSED WITH SMALLER, SIMILARLY SHAPED TEETH, OPPOSINg TEETH REcIPROcALLY fILLINg gAPS WHEN fINgERS cLOSED. MAJOR cHELA MUcH STOUTER THAN MINOR cHELA, cUTTINg EDgE Of DAcTYLUS WITH MODIfIED ELONgATE, ObLIqUELY ORIENTED TOOTH NEAR PROXIMAL END, OPPOSITE fLAT, MOLARIfORM TOOTH ON fIXED fINgER. FIXED fINgER Of MINOR cHELA DEfLEXED DOWNWARDS.
</paragraph>
<caption id="4DA3ADA95F33FFC6FF17DDF1FE12F84A" httpUri="https://zenodo.org/record/898946/files/figure.png" pageId="4" pageNumber="223" targetBox="[165,1429,193,1907]" targetPageId="4">
<paragraph id="1963FD215F33FFC6FF17DDF1FE12F84A" blockId="4.[151,1436,1920,2004]" pageId="4" pageNumber="223">
<emphasis id="2BA821335F33FFC6FF17DDF1FEE5F80B" bold="true" box="[151,278,1920,1942]" pageId="4" pageNumber="223">FIGURE 3.</emphasis>
<taxonomicName id="DEDC86A25F33FFC6FEA0DDF1FE17F808" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[288,484,1920,1942]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="223" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F33FFC6FEA0DDF1FE17F808" box="[288,484,1920,1942]" italics="true" pageId="4" pageNumber="223">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F33FFC6FE6DDDF0FDB2F808" box="[493,577,1921,1942]" pageId="4" pageNumber="223" rank="species">
<emphasis id="2BA821335F33FFC6FE6DDDF0FDB2F808" bold="true" box="[493,577,1921,1942]" pageId="4" pageNumber="223">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male, 72.6 × 38.4 mm (MNHN-IU-2014-18254), Réunion. A, Frontoorbital and buccal region, antero-ventral vieW; B, subhepatic stridulatory granules, antero-ventral vieW; C, third maxillipeds, external vieW. Scale bars: 5 mm.
</paragraph>
</caption>
<caption id="4DA3ADA95F32FFC7FF17DF9EFF36F9FF" httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="5" pageNumber="224" targetBox="[151,1436,167,1498]" targetPageId="5">
<paragraph id="1963FD215F32FFC7FF17DF9EFF36F9FF" blockId="5.[151,1436,1519,1633]" pageId="5" pageNumber="224">
<emphasis id="2BA821335F32FFC7FF17DF9EFEFFF99A" bold="true" box="[151,268,1519,1541]" pageId="5" pageNumber="224">FIGURE 4</emphasis>
. 
<taxonomicName id="DEDC86A25F32FFC7FE99DF9EFD53F99B" authority="(Sakai, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[281,672,1519,1541]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="224" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F32FFC7FE99DF9EFDE2F99B" box="[281,529,1519,1541]" italics="true" pageId="5" pageNumber="224">Laleonectes nipponensis</emphasis>
(Sakai, 1938)
</taxonomicName>
, male, 65.8 × 34.9 mm (RUMF-ZC-2462), Ryukyu Is. (A–D); 
<taxonomicName id="DEDC86A25F32FFC7FA9CDF81FA8BF99B" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[1308,1400,1519,1541]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="224" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F32FFC7FA9CDF81FA8BF99B" box="[1308,1400,1519,1541]" italics="true" pageId="5" pageNumber="224">L. kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F32FFC7FAFDDF9EFF31F9BD" pageId="5" pageNumber="224" rank="species">
<emphasis id="2BA821335F32FFC7FAFDDF9EFF4EF9BD" bold="true" pageId="5" pageNumber="224">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype, male, 72.6 × 38.4 mm (MNHN-IU-2014-18254), Réunion (E–H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P4, dorsal vieW; D, H, right P5, dorsal vieW. Scale bars: 10 mm.
</paragraph>
</caption>
<caption id="4DA3ADA95F31FFC4FF17DF53FB6CFAEB" httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="6" pageNumber="225" targetBox="[193,1410,216,1275]" targetPageId="6">
<paragraph id="1963FD215F31FFC4FF17DF53FB6CFAEB" blockId="6.[151,1436,1314,1397]" pageId="6" pageNumber="225">
<emphasis id="2BA821335F31FFC4FF17DF53FEE5FAA9" bold="true" box="[151,278,1314,1336]" pageId="6" pageNumber="225">FIGURE 5.</emphasis>
Gonopods (G1, G2). 
<taxonomicName id="DEDC86A25F31FFC4FD81DF53FD37FAA6" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[513,708,1314,1336]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="225" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F31FFC4FD81DF53FD37FAA6" box="[513,708,1314,1336]" italics="true" pageId="6" pageNumber="225">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F31FFC4FD4EDF53FCD1FAA6" box="[718,802,1314,1336]" pageId="6" pageNumber="225" rank="species">
<emphasis id="2BA821335F31FFC4FD4EDF53FCEFFAA9" bold="true" box="[718,796,1314,1335]" pageId="6" pageNumber="225">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype, male, 72.6 × 38.4 mm (MNHN-IU-2014-18254), Réunion (A–C); 
<taxonomicName id="DEDC86A25F31FFC4FEC7DF33FD87FAC9" authority="(Sakai, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[327,628,1345,1367]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="225" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F31FFC4FEC7DF33FE2EFAC9" box="[327,477,1346,1367]" italics="true" pageId="6" pageNumber="225">L. nipponensis</emphasis>
(Sakai, 1938)
</taxonomicName>
, male, 65.8 × 34.9 mm (RUMF-ZC-2462), Ryukyu Is. (D–F). A, D, left G1, external vieW; B, E, left G2, external vieW; C, F, distal tip of left G2, external vieW. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="1963FD215F31FFC4FF47DFD3FC6FF9F0" blockId="6.[151,1438,1442,1971]" pageId="6" pageNumber="225">
FIRST TO THIRD AMbULATORY LEgS, P2–P4 (
<figureCitation id="81E7E1A45F31FFC4FDFDDFD3FCF5FA24" box="[637,774,1442,1466]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="6" pageNumber="225">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F31FFC4FC94DFD3FCCAFA24" box="[788,825,1442,1466]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="6" pageNumber="225">4G</figureCitation>
), LONg, SLENDER, UNARMED; MERI SUb-cYLINDRIcAL, LENgTH AbOUT 4.7–5.7 TIMES WIDTH; cARPI SUbcYLINDRIcAL, WIDER DISTALLY THAN PROXIMALLY, ANTERIOR MARgIN SUb-cRISTATE, WITH LOW, ROUNDED, LONgITUDINAL RIDgE ON DORSAL SURfAcE; PROPODI LONg, SLENDER, fLATTENED, LENgTH AbOUT 4.3–5.2 TIMES WIDTH, ANTERIOR MARgIN WITH LOW cREST, DORSAL AND VENTRAL SURfAcES WITH SHALLOW LONgITUDINAL gROOVES; DAcTYLI NARROW, SPATULIfORM, ENDINg DISTALLY IN SHARP cHITINOUS TIP, WITH fIVE SMOOTH cRESTS DISTRIbUTED ON ANTERIOR MARgIN AND ON DORSAL AND VENTRAL SURfAcES, LENgTH AbOUT 6.4–7.9 TIMES WIDTH.
</paragraph>
<paragraph id="1963FD215F31FFC4FF47DC0BFAD6F960" blockId="6.[151,1438,1442,1971]" pageId="6" pageNumber="225">
LAST AMbULATORY LEg, P5 (
<figureCitation id="81E7E1A45F31FFC4FE65DC0BFD9EF90C" box="[485,621,1658,1682]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="6" pageNumber="225">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F31FFC4FDFADC0BFD6CF90C" box="[634,671,1658,1682]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="6" pageNumber="225">4H</figureCitation>
), MUcH SHORTER, STOUTER, DISTAL ARTIcLES fLATTENED AND WIDENED; MERUS AND cARPUS, SHORTER, STOUTER RELATIVE TO THOSE Of P2–P4, MERUS LENgTH AbOUT 1.9–2.4 TIMES WIDTH, cARPUS LENgTH AbOUT 1.5–1.7 TIMES WIDTH; PROPODUS AND DAcTYLUS MUcH fLATTENED AND WIDENED, ANTERIOR MARgINS Of PROPODUS AND ANTERIOR AND POSTERIOR MARgINS Of DAcTYLUS LINED WITH fINE SETAE; DAcTYLUS ENDINg DISTALLY IN A SHARP, cHITINOUS TIP.
</paragraph>
<paragraph id="1963FD215F31FFC4FF47DD7BFCF3F82C" blockId="6.[151,1438,1442,1971]" pageId="6" pageNumber="225">
G1 (
<figureCitation id="81E7E1A45F31FFC4FF78DD7BFEA3F8BD" box="[248,336,1802,1827]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="6" pageNumber="225">FIg. 5A</figureCitation>
) cURVED LATERALLY AND OUTWARDLY IN SITU; PROXIMAL HALf WIDE, DISTAL HALf SLENDER, SINUOUS, ESPEcIALLY WHEN VIEWED LATERALLY; ON DISTAL HALf Of SLENDER PORTION, MESIAL MARgIN LINED WITH LONg, STIff, SPINIfORM SETAE, ON PROXIMAL HALf Of SLENDER PORTION, MESIAL MARgIN LINED WITH SUPPLE, PLUMOSE SETAE. G2 (
<figureCitation id="81E7E1A45F31FFC4FBF7DD23FB00F8F4" box="[1143,1267,1874,1899]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="6" pageNumber="225">FIg. 5B, C</figureCitation>
) LONg, SLENDER, AbOUT TWO-THIRDS LENgTH Of G1, SLIgHTLY cURVED, DISTAL TIP bILObATE, LObES SUbEqUAL. PENIS LONg, EMERgINg fROM gONOPORE ON P5 cOXA, ANTERIOR TO cOXO-STERNAL cONDYLE.
</paragraph>
<caption id="4DA3ADA95F30FFC5FF17DDC0FB17F87B" httpUri="https://zenodo.org/record/898952/files/figure.png" pageId="7" pageNumber="226" targetBox="[211,1387,193,1949]" targetPageId="7">
<paragraph id="1963FD215F30FFC5FF17DDC0FB17F87B" blockId="7.[151,1436,1969,2022]" pageId="7" pageNumber="226">
<emphasis id="2BA821335F30FFC5FF17DDC0FEE0F858" bold="true" box="[151,275,1969,1991]" pageId="7" pageNumber="226">FIGURE 6.</emphasis>
Fresh coloration. A, 
<taxonomicName id="DEDC86A25F30FFC5FE6FDDC0FD43F859" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[495,688,1969,1991]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="226" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F30FFC5FE6FDDC0FD43F859" box="[495,688,1969,1991]" italics="true" pageId="7" pageNumber="226">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F30FFC5FD38DDC0FCFAF859" box="[696,777,1969,1991]" pageId="7" pageNumber="226" rank="species">
<emphasis id="2BA821335F30FFC5FD38DDC0FCF1F858" bold="true" box="[696,770,1969,1990]" pageId="7" pageNumber="226">sp. nov</emphasis>
.
</taxonomicNameLabel>
, male (ZRC 2017.0335), Tamil Nadu; B, 
<taxonomicName id="DEDC86A25F30FFC5FB39DDC3FF26F87B" authority="(Sakai, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="226" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F30FFC5FB39DDC3FABEF859" box="[1209,1357,1970,1991]" italics="true" pageId="7" pageNumber="226">L. nipponensis</emphasis>
(Sakai, 1938)
</taxonomicName>
, male (RUMF-ZC-2462), Ryukyu Is.; C, 
<taxonomicName id="DEDC86A25F30FFC5FDF8DDA0FC6FF87B" authority="(Sakai, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[632,924,1999,2022]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="226" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F30FFC5FDF8DDA0FD7EF878" box="[632,653,2001,2022]" italics="true" pageId="7" pageNumber="226">L.</emphasis>
<emphasis id="2BA821335F30FFC5FD13DDA0FCFFF878" box="[659,780,2001,2022]" italics="true" pageId="7" pageNumber="226">nipponensis</emphasis>
(Sakai, 1938)
</taxonomicName>
, male (ZRC 2017.0009), Bohol.
</paragraph>
</caption>
<paragraph id="1963FD215F3FFFCAFF47DAE6FD99FF66" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="227">
<emphasis id="2BA821335F3FFFCAFF47DAE6FE76FF2E" bold="true" box="[199,389,151,176]" pageId="8" pageNumber="227">Live coloration.</emphasis>
DORSAL cARAPAcE AND PEREOPODS gENERALLY YELLOWISH ORANgE. DORSAL SURfAcE Of cARAPAcE WITH IRREgULAR SPLOTcHES Of RED, PARTIcULARLY ON METAgASTRIc AND cARDIAc REgIONS, LARgE gRANULES ALSO RED. PEREOPODS WITH RED AND ORANgE bANDINg PATTERN (
<figureCitation id="81E7E1A45F3FFFCAFD82DA91FDAEFF66" box="[514,605,223,248]" captionStart="FIGURE 6" captionStartId="7.[151,250,1969,1991]" captionTargetBox="[211,1387,193,1949]" captionTargetId="figure@7.[211,1387,193,1949]" captionTargetPageId="7" captionText="FIGURE 6. Fresh coloration. A, Laleonectes kuriya sp. nov., male (ZRC 2017.0335), Tamil Nadu; B, L. nipponensis (Sakai, 1938), male (RUMF-ZC- 2462), Ryukyu Is.; C, L. nipponensis (Sakai, 1938), male (ZRC 2017.0009), Bohol." httpUri="https://zenodo.org/record/898952/files/figure.png" pageId="8" pageNumber="227">FIg. 6A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="51C6AEAA5F3FFFCAFF47DB75FEC2FEFA" pageId="8" pageNumber="227" type="etymology">
<paragraph id="1963FD215F3FFFCAFF47DB75FEC2FEFA" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="227">
<emphasis id="2BA821335F3FFFCAFF47DB75FEBDFE83" bold="true" box="[199,334,260,285]" pageId="8" pageNumber="227">Etymology.</emphasis>
THE SPEcIfIc EPITHET Of THE NEW SPEcIES IS DERIVED fROM THE MALAYALAM WORD fOR SHORT, “KURIYA”, IN ALLUSION TO THIS SPEcIES’ APPARENTLY SHORTER LEgS RELATIVE TO ITS cLOSEST cONgENER, 
<taxonomicName id="DEDC86A25F3FFFCAFB91DB58FB46FEA1" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[1041,1205,296,320]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFB91DB58FB46FEA1" box="[1041,1205,296,320]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
. IT IS USED 
<collectingCountry id="61CBBDB15F3FFFCAFAB1DB59FABBFEDE" box="[1329,1352,296,320]" name="American Samoa" pageId="8" pageNumber="227">AS</collectingCountry>
A NOUN IN APPOSITION.
</paragraph>
</subSubSection>
<subSubSection id="51C6AEAA5F3FFFCAFF47DB1EFADEFC1C" pageId="8" pageNumber="227" type="discussion">
<paragraph id="1963FD215F3FFFCAFF47DB1EFAB1FDA1" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="227">
<emphasis id="2BA821335F3FFFCAFF47DB1EFEC8FE16" bold="true" box="[199,315,367,392]" pageId="8" pageNumber="227">Remarks.</emphasis>
<taxonomicName id="DEDC86A25F3FFFCAFEC5DB01FDEDFE16" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[325,542,368,392]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F3FFFCAFEC5DB01FDEDFE16" box="[325,542,368,392]" italics="true" pageId="8" pageNumber="227">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F3FFFCAFDA8DB1EFD76FE16" box="[552,645,367,392]" pageId="8" pageNumber="227" rank="species">
<emphasis id="2BA821335F3FFFCAFDA8DB1EFD76FE16" bold="true" box="[552,645,367,392]" pageId="8" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
cAN bE DISTINgUISHED fROM THE 
<typeStatus id="C66743835F3FFFCAFC75DB01FBD6FE19" box="[1013,1061,368,391]" pageId="8" pageNumber="227">TYPE</typeStatus>
SPEcIES, 
<taxonomicName id="DEDC86A25F3FFFCAFB12DB00FAF1FE16" authorityName="A. Milne-Edwards" authorityYear="1878" box="[1170,1282,369,392]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="vocans">
<emphasis id="2BA821335F3FFFCAFB12DB00FAF1FE16" box="[1170,1282,369,392]" italics="true" pageId="8" pageNumber="227">L. vocans</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="7D4D80D05F3FFFCAFABDDB1EFF20FE32" author="RATHbUN" pageId="8" pageNumber="227" refString="Rathbun, M. J. (1930) The cancroid crabs of America of the families Euryalidae, Portunidae, Atelecyclidae, Cancridae, and Xanthidae. Bulletin of the United States National Museum, 152, 1 - 609. https: // doi. org / 10.5479 / si. 03629236.152. i" type="journal article" year="1930">RATHbUN 1930</bibRefCitation>
: PL. 25 fIg. 3; 
<bibRefCitation id="7D4D80D05F3FFFCAFEF5DBE5FD79FE32" author="MANNINg" box="[373,650,404,428]" pageId="8" pageNumber="227" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, i - v, 1 - 91." type="journal article" year="1990">MANNINg &amp; CHAcE 1990</bibRefCitation>
: fIg. 33), bY THE fOLLOWINg fEATURES: THE MORE PROJEcTINg fRONTAL TEETH, PARTIcULARLY THE LATERALS (
<figureCitation id="81E7E1A45F3FFFCAFE3BDBCAFDEAFE4D" box="[443,537,443,467]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="8" pageNumber="227">FIg. 2A</figureCitation>
) [VS. LATERALS LOW IN 
<taxonomicName id="DEDC86A25F3FFFCAFC9CDBCAFC7DFE4C" authorityName="A. Milne-Edwards" authorityYear="1878" box="[796,910,443,466]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="vocans">
<emphasis id="2BA821335F3FFFCAFC9CDBCAFC7DFE4C" box="[796,910,443,466]" italics="true" pageId="8" pageNumber="227">L. vocans</emphasis>
</taxonomicName>
]; THE bROADER PENULTIMATE (8th) TOOTH ON THE ANTEROLATERAL MARgIN Of THE cARAPAcE (
<figureCitation id="81E7E1A45F3FFFCAFDBADBAEFD60FE69" box="[570,659,479,503]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="8" pageNumber="227">FIg. 2A</figureCitation>
) [VS. bASE Of PENULTIMATE TOOTH MUcH NARROWER IN 
<taxonomicName id="DEDC86A25F3FFFCAFB46DB91FAC1FE69" authorityName="A. Milne-Edwards" authorityYear="1878" box="[1222,1330,480,503]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="vocans">
<emphasis id="2BA821335F3FFFCAFB46DB91FAC1FE69" box="[1222,1330,480,503]" italics="true" pageId="8" pageNumber="227">L. vocans</emphasis>
</taxonomicName>
]; AND THE THREE WELL-DEVELOPED SPINES ON THE fLEXOR MARgIN Of THE P1 MERUS (
<figureCitation id="81E7E1A45F3FFFCAFC12D872FBE8FD84" box="[914,1051,515,539]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="8" pageNumber="227">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F3FFFCAFBA8D872FBB4FD85" box="[1064,1095,515,539]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">4F</figureCitation>
) [VS. ONE SHARP SPINE AND ONE gRANULAR TOOTH ON fLEXOR MARgIN Of P1 MERUS IN 
<taxonomicName id="DEDC86A25F3FFFCAFD29D859FCE6FDA1" authorityName="A. Milne-Edwards" authorityYear="1878" box="[681,789,552,575]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="vocans">
<emphasis id="2BA821335F3FFFCAFD29D859FCE6FDA1" box="[681,789,552,575]" italics="true" pageId="8" pageNumber="227">L. vocans</emphasis>
</taxonomicName>
]. THE G1 MORPHOLOgY Of 
<taxonomicName id="DEDC86A25F3FFFCAFBB6D859FB52FDA1" authorityName="A. Milne-Edwards" authorityYear="1878" box="[1078,1185,552,575]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="vocans">
<emphasis id="2BA821335F3FFFCAFBB6D859FB52FDA1" box="[1078,1185,552,575]" italics="true" pageId="8" pageNumber="227">L. vocans</emphasis>
</taxonomicName>
IS NOT KNOWN.
</paragraph>
<paragraph id="1963FD215F3FFFCAFF47D83AFADEFC1C" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="227">
THE NEW SPEcIES MAY bE DISTINgUISHED fROM 
<taxonomicName id="DEDC86A25F3FFFCAFD35D83AFCDFFDFD" authorityName="Crosnier &amp; Moosa" authorityYear="2002" box="[693,812,587,611]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="stridens">
<emphasis id="2BA821335F3FFFCAFD35D83AFCDFFDFD" box="[693,812,587,611]" italics="true" pageId="8" pageNumber="227">L. stridens</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="7D4D80D05F3FFFCAFCDED83AFB83FDFD" author="CROSNIER" box="[862,1136,587,611]" pageId="8" pageNumber="227" refString="Crosnier, A. &amp; Moosa, M. K. (2002) Trois Portunidae (Crustacea, Decapoda, Brachyura) noveaux de Polynesie francaise. Zoosystema, 24 (2), 385 - 399." type="journal article" year="2002">CROSNIER &amp; MOOSA 2002</bibRefCitation>
: fIgS. 3A, 5A, B) PRIMARILY bY THE fORM Of THE G1 AND G2, bOTH Of WHIcH ARE MUcH LONgER AND MORE SLENDER, WITH THE G1 HAVINg STIff SPINIfORM SETAE ON THE MESIAL MARgIN Of ITS DISTAL HALf, IN 
<taxonomicName id="DEDC86A25F3FFFCAFD4FD8E2FCC9FD35" authorityName="Devi" authorityYear="2017" box="[719,826,659,683]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="kuriya">
<emphasis id="2BA821335F3FFFCAFD4FD8E2FCC9FD35" box="[719,826,659,683]" italics="true" pageId="8" pageNumber="227">L. kuriya</emphasis>
</taxonomicName>
(
<figureCitation id="81E7E1A45F3FFFCAFCCCD8E2FC26FD35" box="[844,981,658,683]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="8" pageNumber="227">FIgS. 5A–C</figureCitation>
) [VS. SHORTER, STOUTER, AND WITHOUT STIff SPINIfORM SETAE IN 
<taxonomicName id="DEDC86A25F3FFFCAFEE8D8C9FE2CFD51" authorityName="Crosnier &amp; Moosa" authorityYear="2002" box="[360,479,695,719]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="stridens">
<emphasis id="2BA821335F3FFFCAFEE8D8C9FE2CFD51" box="[360,479,695,719]" italics="true" pageId="8" pageNumber="227">L. stridens</emphasis>
</taxonomicName>
]. FURTHERMORE, 
<taxonomicName id="DEDC86A25F3FFFCAFD12D8C9FD0BFD51" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[658,760,695,719]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F3FFFCAFD12D8C9FD0BFD51" box="[658,760,695,719]" italics="true" pageId="8" pageNumber="227">L. kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F3FFFCAFC80D8C7FCA9FD51" box="[768,858,694,719]" pageId="8" pageNumber="227" rank="species">
<emphasis id="2BA821335F3FFFCAFC80D8C7FCA9FD51" bold="true" box="[768,858,694,719]" pageId="8" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
DIffERS fROM 
<taxonomicName id="DEDC86A25F3FFFCAFC70D8C9FB94FD51" authorityName="Crosnier &amp; Moosa" authorityYear="2002" box="[1008,1127,695,719]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="stridens">
<emphasis id="2BA821335F3FFFCAFC70D8C9FB94FD51" box="[1008,1127,695,719]" italics="true" pageId="8" pageNumber="227">L. stridens</emphasis>
</taxonomicName>
IN HAVINg: 
<emphasis id="2BA821335F3FFFCAFB68D8C7FAFBFD51" bold="true" box="[1256,1288,694,719]" pageId="8" pageNumber="227">(1)</emphasis>
THE cARAPAcE ANTEROLATERAL AND POSTEROLATERAL MARgINS LESS EXTENDED (
<figureCitation id="81E7E1A45F3FFFCAFCA0D8AAFC8DFD6D" box="[800,894,731,755]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="8" pageNumber="227">FIg. 2A</figureCitation>
) [VS. cARAPAcE ANTEROLATERAL AND POSTEROLATERAL MARgINS LONgER IN 
<taxonomicName id="DEDC86A25F3FFFCAFEE7D971FE2CFC89" authorityName="Crosnier &amp; Moosa" authorityYear="2002" box="[359,479,767,791]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="stridens">
<emphasis id="2BA821335F3FFFCAFEE7D971FE2CFC89" box="[359,479,767,791]" italics="true" pageId="8" pageNumber="227">L. stridens</emphasis>
</taxonomicName>
]; 
<emphasis id="2BA821335F3FFFCAFE79D88FFDE9FC89" bold="true" box="[505,538,766,791]" pageId="8" pageNumber="227">(2)</emphasis>
THE LAST (LARgEST) TOOTH ON THE cARAPAcE ANTEROLATERAL MARgIN SHORTER AND STOUTER (
<figureCitation id="81E7E1A45F3FFFCAFF1FD952FF09FCA5" box="[159,250,803,827]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="8" pageNumber="227">FIg. 2A</figureCitation>
) [VS. LONgER AND MORE SLENDER IN 
<taxonomicName id="DEDC86A25F3FFFCAFDFED952FD04FCA5" authorityName="Crosnier &amp; Moosa" authorityYear="2002" box="[638,759,803,827]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="stridens">
<emphasis id="2BA821335F3FFFCAFDFED952FD04FCA5" box="[638,759,803,827]" italics="true" pageId="8" pageNumber="227">L. stridens</emphasis>
</taxonomicName>
]; 
<emphasis id="2BA821335F3FFFCAFC91D953FCC1FCA5" bold="true" box="[785,818,802,827]" pageId="8" pageNumber="227">(3)</emphasis>
A MUcH MORE gRANULAR cARAPAcE, WITH DEEPER fURROWS (
<figureCitation id="81E7E1A45F3FFFCAFF1FD936FF0FFCC1" box="[159,252,839,863]" captionStart="FIGURE 2" captionStartId="3.[151,250,1592,1614]" captionTargetBox="[151,1436,193,1571]" captionTargetId="figure@3.[151,1436,193,1571]" captionTargetPageId="3" captionText="FIGURE 2. Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion. A, Carapace, dorsal vieW; B, cephalothorax and pleon, ventral vieW; C, cephalothorax, anterior vieW; D, cephalothorax, posterior vieW; right (major) chela, E, external vieW, G, internal vieW; left (minor) chela, F, external vieW, H, internal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898944/files/figure.png" pageId="8" pageNumber="227">FIg. 2A</figureCitation>
) [VS. SMALLER AND LESS WIDELY DISTRIbUTED gRANULES, SHALLOW fURROWS ON THE DORSAL cARAPAcE]; AND 
<emphasis id="2BA821335F3FFFCAFAFBD937FA6FFCC1" bold="true" box="[1403,1436,838,863]" pageId="8" pageNumber="227">(4)</emphasis>
RELATIVELY SHORTER AND STOUTER PEREOPODS (
<figureCitation id="81E7E1A45F3FFFCAFDE1D91AFD15FC1C" box="[609,742,875,899]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="8" pageNumber="227">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F3FFFCAFD71D91AFCC0FC1D" box="[753,819,875,899]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">4G, H</figureCitation>
) [VS. LONgER AND MORE SLENDER IN 
<taxonomicName id="DEDC86A25F3FFFCAFB26D91AFAEEFC1D" authorityName="Crosnier &amp; Moosa" authorityYear="2002" box="[1190,1309,875,899]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="stridens">
<emphasis id="2BA821335F3FFFCAFB26D91AFAEEFC1D" box="[1190,1309,875,899]" italics="true" pageId="8" pageNumber="227">L. stridens</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="51C6AEAA5F3FFFCBFF47D9FEFCA5FF69" lastPageId="9" lastPageNumber="228" pageId="8" pageNumber="227" type="reference_group">
<paragraph id="1963FD215F3FFFCAFF47D9FEFD9DFA09" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="227">
<taxonomicName id="DEDC86A25F3FFFCAFF47D9FEFE68FC39" ID-CoL="84FXK" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[199,411,911,935]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F3FFFCAFF47D9FEFE68FC39" box="[199,411,911,935]" italics="true" pageId="8" pageNumber="227">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F3FFFCAFE22D9FFFE0FFC39" box="[418,508,910,935]" pageId="8" pageNumber="227" rank="species">
<emphasis id="2BA821335F3FFFCAFE22D9FFFE0FFC39" bold="true" box="[418,508,910,935]" pageId="8" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
IS MOST SIMILAR TO 
<taxonomicName id="DEDC86A25F3FFFCAFD4CD9E1FBFDFC39" ID-CoL="3RYC2" authority="(SAKAI, 1938)" authorityName="Sakai" authorityYear="1938" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[716,1038,911,935]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="226" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFD4CD9E1FC83FC39" box="[716,880,912,935]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
(
<bibRefCitation id="7D4D80D05F3FFFCAFCFDD9FEFBF5FC39" author="SAKAI" box="[893,1030,911,935]" pageId="8" pageNumber="227" refString="Sakai, T. (1938) On three systematically interesting crabs from Japan, one of Which is neW to science. Annotationes Zoologicae Japonenses, 17 (3), 301 - 309." type="journal article" year="1938">SAKAI, 1938</bibRefCitation>
)
</taxonomicName>
IN THE fORM Of THE cARAPAcE AND G1 AND IN gENERAL cOLOUR PATTERN (cf. 
<bibRefCitation id="7D4D80D05F3FFFCAFD8BD9C3FD78FC54" author="SAKAI" box="[523,651,946,970]" pageId="8" pageNumber="227" refString="Sakai, T. (1938) On three systematically interesting crabs from Japan, one of Which is neW to science. Annotationes Zoologicae Japonenses, 17 (3), 301 - 309." type="journal article" year="1938">SAKAI 1938</bibRefCitation>
: TEXT-fIg. 1, PL. 16 fIg. 1; 
<bibRefCitation id="7D4D80D05F3FFFCAFC21D9C3FBD1FC54" author="SAKAI" box="[929,1058,946,970]" pageId="8" pageNumber="227" refString="Sakai, T. (1939) Studies on the Crabs of Japan IV. Brachygnatha, Brachyrhyncha. Vol. 3. Yokendo Co. Ltd, Tokyo, 377 pp., 70 pls. 23, 23 pp. index. [pp. 365 - 741 (text &amp; text-figs.), pls. 42 - 111 (plates), pp. 1 - 23 (index)]" type="book chapter" year="1939">SAKAI 1939</bibRefCitation>
: TEXT-fIgS. 6, 7, PL. 82 fIg. 1; 
<bibRefCitation id="7D4D80D05F3FFFCAFADED9C3FF20FC70" author="SAKAI" pageId="8" pageNumber="227" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd, Tokyo, 773 pp., 251 pls." type="book" year="1976">SAKAI 1976</bibRefCitation>
: TEXT-fIg. 186, PL. 119 fIg. 3; 
<bibRefCitation id="7D4D80D05F3FFFCAFDADD9A7FD39FC70" author="MIYAKE" box="[557,714,982,1006]" pageId="8" pageNumber="227" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in color. Vol. II. Brachyura (Crabs). Hoikusha, Osaka, viii + 277 pp., 64 pls. [in Japanese]" type="book" year="1983">MIYAKE 1983</bibRefCitation>
: PL. 29 fIg. 2). IT, HOWEVER, DIffERS fROM 
<taxonomicName id="DEDC86A25F3FFFCAFB28D9A7FABCFC73" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[1192,1359,982,1005]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFB28D9A7FABCFC73" box="[1192,1359,982,1005]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
IN THE fOLLOWINg MORPHOLOgIcAL fEATURES: 
<emphasis id="2BA821335F3FFFCAFD9DD988FDCEFB8C" bold="true" box="[541,573,1017,1042]" pageId="8" pageNumber="227">(1)</emphasis>
THE AMbULATORY LEgS ARE RELATIVELY SHORTER AND STOUTER: fOR INSTANcE, THE DISTAL ENDS Of THE P2 AND P3 MERI DO NOT OVERREAcH THE DISTAL END Of THE P1 MERUS (
<figureCitation id="81E7E1A45F3FFFCAFC78DE6CFB81FBAB" box="[1016,1138,1053,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="8" pageNumber="227">FIg. 1A, B</figureCitation>
) [VS. DISTAL ENDS Of P2, P3 MERI cLEARLY OVERREAcHINg DISTAL END Of P1 MERUS IN 
<taxonomicName id="DEDC86A25F3FFFCAFD64DE30FC7BFBC6" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[740,904,1089,1112]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFD64DE30FC7BFBC6" box="[740,904,1089,1112]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
; cf. 
<figureCitation id="81E7E1A45F3FFFCAFC3CDE31FBB1FBC6" box="[956,1090,1088,1112]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="8" pageNumber="227">FIgS. 1C, D</figureCitation>
; 
<bibRefCitation id="7D4D80D05F3FFFCAFBCEDE31FB22FBC6" author="SAKAI" box="[1102,1233,1088,1112]" pageId="8" pageNumber="227" refString="Sakai, T. (1939) Studies on the Crabs of Japan IV. Brachygnatha, Brachyrhyncha. Vol. 3. Yokendo Co. Ltd, Tokyo, 377 pp., 70 pls. 23, 23 pp. index. [pp. 365 - 741 (text &amp; text-figs.), pls. 42 - 111 (plates), pp. 1 - 23 (index)]" type="book chapter" year="1939">SAKAI 1939</bibRefCitation>
: TEXT-fIg. 6; PL. 82 fIg. 1); AND THE LENgTH-TO-WIDTH RATIOS Of cORRESPONDINg ARTIcLES ARE LOWER, E.g. AVERAgE Of 5.0 fOR P4 MERI, AND 4.4 fOR P4 PROPODI (
<figureCitation id="81E7E1A45F3FFFCAFECDDEF6FE55FB01" box="[333,422,1159,1183]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">FIg. 4G</figureCitation>
) (VS. HIgHER LENgTH-TO-WIDTH RATIOS IN 
<taxonomicName id="DEDC86A25F3FFFCAFCD3DEF9FC0BFB01" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[851,1016,1160,1183]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFCD3DEF9FC0BFB01" box="[851,1016,1160,1183]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
, E.g. AVERAgE Of 6.3 fOR P4 MERI, AND 6.4 fOR P4 PROPODI (
<figureCitation id="81E7E1A45F3FFFCAFEF8DEDAFE21FB5D" box="[376,466,1195,1219]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">FIg. 4E</figureCitation>
)]; 
<emphasis id="2BA821335F3FFFCAFE70DEDBFDE2FB5D" bold="true" box="[496,529,1194,1219]" pageId="8" pageNumber="227">(2)</emphasis>
THE cHELIPEDS ALSO TEND TO bE STOUTER IN 
<taxonomicName id="DEDC86A25F3FFFCAFC56DEDAFBCFFB5C" ID-CoL="84FXK" authorityName="Devi" authorityYear="2017" box="[982,1084,1194,1218]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="kuriya">
<emphasis id="2BA821335F3FFFCAFC56DEDAFBCFFB5C" box="[982,1084,1194,1218]" italics="true" pageId="8" pageNumber="227">L. kuriya</emphasis>
</taxonomicName>
(
<figureCitation id="81E7E1A45F3FFFCAFBCBDEDAFB20FB5C" box="[1099,1235,1195,1219]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="8" pageNumber="227">FIgS. 1A, B</figureCitation>
, 
<figureCitation id="81E7E1A45F3FFFCAFB60DEDAFB0CFB5D" box="[1248,1279,1195,1219]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">4F</figureCitation>
) cOMPARED TO THOSE Of 
<taxonomicName id="DEDC86A25F3FFFCAFF7EDEBEFE56FB78" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[254,421,1231,1254]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFF7EDEBEFE56FB78" box="[254,421,1231,1254]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
(
<figureCitation id="81E7E1A45F3FFFCAFE36DEBFFDB1FB78" box="[438,578,1230,1254]" captionStart="FIGURE 1" captionStartId="2.[151,250,1354,1376]" captionTargetBox="[151,1436,519,1290]" captionTargetId="figure@2.[151,1436,519,1290]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus. Laleonectes kuriya sp. nov. (A, B): A, holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion; B, non-type male, 59.2 × 33.1 mm (ZRC 2017.0335), Tamil Nadu. Laleonectes nipponensis (Sakai, 1938) (C, D): C, male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is.; D, male, 51.1 × 27.9 mm (ZRC 2017.0009), Bohol. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898942/files/figure.png" pageId="8" pageNumber="227">FIgS. 1C, D</figureCitation>
, 
<figureCitation id="81E7E1A45F3FFFCAFDD1DEBFFD86FB78" box="[593,629,1230,1254]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">4B</figureCitation>
); 
<emphasis id="2BA821335F3FFFCAFD0DDEBCFD5DFB78" bold="true" box="[653,686,1229,1254]" pageId="8" pageNumber="227">(3)</emphasis>
PLEOMERE 6 IS RELATIVELY LONgER, NARROWER AND ITS LATERAL MARgINS NEARLY STRAIgHT (
<figureCitation id="81E7E1A45F3FFFCAFEC7DE83FE6FFA94" box="[327,412,1266,1290]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">FIg. 4E</figureCitation>
) [VS. PLEOMERE 6 SHORTER &amp; WIDER; LATERAL MARgINS MORE cONVEX IN 
<taxonomicName id="DEDC86A25F3FFFCAFB03DE83FAD4FA97" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[1155,1319,1266,1289]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFB03DE83FAD4FA97" box="[1155,1319,1266,1289]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
; 
<figureCitation id="81E7E1A45F3FFFCAFAB5DE83FA63FA94" box="[1333,1424,1266,1290]" captionStart="FIGURE 4" captionStartId="5.[151,250,1519,1541]" captionTargetBox="[151,1436,167,1498]" captionTargetId="figure@5.[151,1436,167,1498]" captionTargetPageId="5" captionText="FIGURE 4. Laleonectes nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (A – D); L. kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (E – H). A, E, pleon, ventral vieW; B, F, major cheliped merus and stridulatory granules, dorsal vieW; C, G, right P 4, dorsal vieW; D, H, right P 5, dorsal vieW. Scale bars: 10 mm." httpUri="https://zenodo.org/record/898948/files/figure.png" pageId="8" pageNumber="227">FIg. 4A</figureCitation>
]; 
<emphasis id="2BA821335F3FFFCAFF17DF64FF4BFAB0" bold="true" box="[151,184,1301,1326]" pageId="8" pageNumber="227">(4)</emphasis>
THE PROXIMO-LATERAL SHELf Of THE G1 IS LONgER, AND THE DISTAL HALf IS STOUTER AND LESS cURVED (
<figureCitation id="81E7E1A45F3FFFCAFA83DF64FAACFAB0" box="[1283,1375,1301,1326]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="8" pageNumber="227">FIg. 5A</figureCitation>
) [VS. PROXIMO-LATERAL SHELf SHORTER, DISTAL HALf MORE SLENDER AND cURVED IN 
<taxonomicName id="DEDC86A25F3FFFCAFC18DF48FBCFFACE" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[920,1084,1337,1360]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFC18DF48FBCFFACE" box="[920,1084,1337,1360]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
; 
<figureCitation id="81E7E1A45F3FFFCAFBCBDF48FB56FACF" box="[1099,1189,1336,1361]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="8" pageNumber="227">FIg. 5D</figureCitation>
]; AND 
<emphasis id="2BA821335F3FFFCAFB6ADF49FAF9FACF" bold="true" box="[1258,1290,1336,1361]" pageId="8" pageNumber="227">(5)</emphasis>
ON THE DISTAL TIP Of G2, THE LATERAL LObE IS SUbEqUAL IN LENgTH TO THE MESIAL LObE (
<figureCitation id="81E7E1A45F3FFFCAFCFDDF2DFC07FAEA" box="[893,1012,1372,1397]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="8" pageNumber="227">FIg. 5B, C</figureCitation>
) [VS. LATERAL LObE IS MUcH SHORTER THAN MESIAL LObE IN 
<taxonomicName id="DEDC86A25F3FFFCAFEBBDFF1FE2DFA09" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[315,478,1408,1431]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFEBBDFF1FE2DFA09" box="[315,478,1408,1431]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
; 
<figureCitation id="81E7E1A45F3FFFCAFE6DDFF1FD92FA09" box="[493,609,1407,1432]" captionStart="FIGURE 5" captionStartId="6.[151,250,1314,1336]" captionTargetBox="[193,1410,216,1275]" captionTargetId="figure@6.[151,1436,193,1293]" captionTargetPageId="6" captionText="FIGURE 5. Gonopods (G 1, G 2). Laleonectes kuriya sp. nov., holotype, male, 72.6 × 38.4 mm (MNHN-IU- 2014 - 18254), Réunion (A – C); L. nipponensis (Sakai, 1938), male, 65.8 × 34.9 mm (RUMF-ZC- 2462), Ryukyu Is. (D – F). A, D, left G 1, external vieW; B, E, left G 2, external vieW; C, F, distal tip of left G 2, external vieW. Scale bars: 1 mm." httpUri="https://zenodo.org/record/898950/files/figure.png" pageId="8" pageNumber="227">FIg. 5E, F</figureCitation>
].
</paragraph>
<paragraph id="1963FD215F3FFFCAFF47DFD2FE54F819" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="227">
<taxonomicName id="DEDC86A25F3FFFCAFF47DFD2FE6FFA25" ID-CoL="84FXK" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[199,412,1443,1467]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F3FFFCAFF47DFD2FE6FFA25" box="[199,412,1443,1467]" italics="true" pageId="8" pageNumber="227">Laleonectes kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F3FFFCAFE24DFD3FE0CFA25" box="[420,511,1442,1467]" pageId="8" pageNumber="227" rank="species">
<emphasis id="2BA821335F3FFFCAFE24DFD3FE0CFA25" bold="true" box="[420,511,1442,1467]" pageId="8" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
IS PRESENTLY KNOWN ONLY fROM RéUNION AND fROM THE SOUTHWESTERN cOAST Of INDIA, fROM DEPTHS Of AT LEAST 100 M. BY cONTRAST, 
<taxonomicName id="DEDC86A25F3FFFCAFD08DFB6FCDEFA40" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[648,813,1479,1502]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFD08DFB6FCDEFA40" box="[648,813,1479,1502]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
APPEARS TO bE WIDESPREAD, HAVINg bEEN REPORTED IN THE PAcIfIc REgION fROM JAPAN—KII PENINSULA (
<bibRefCitation id="7D4D80D05F3FFFCAFD1ADF9BFCD6F99C" author="SAKAI" box="[666,805,1514,1538]" pageId="8" pageNumber="227" refString="Sakai, T. (1938) On three systematically interesting crabs from Japan, one of Which is neW to science. Annotationes Zoologicae Japonenses, 17 (3), 301 - 309." type="journal article" year="1938">SAKAI 1938</bibRefCitation>
, 
<bibRefCitation id="7D4D80D05F3FFFCAFCB7DF9BFC81F99C" author="SAKAI" box="[823,882,1514,1538]" pageId="8" pageNumber="227" refString="Sakai, T. (1939) Studies on the Crabs of Japan IV. Brachygnatha, Brachyrhyncha. Vol. 3. Yokendo Co. Ltd, Tokyo, 377 pp., 70 pls. 23, 23 pp. index. [pp. 365 - 741 (text &amp; text-figs.), pls. 42 - 111 (plates), pp. 1 - 23 (index)]" type="book chapter" year="1939">1939</bibRefCitation>
, 
<bibRefCitation id="7D4D80D05F3FFFCAFC04DF98FC4CF99C" author="SAKAI" box="[900,959,1513,1538]" pageId="8" pageNumber="227" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd, Tokyo, 773 pp., 251 pls." type="book" year="1976">1976</bibRefCitation>
; 
<bibRefCitation id="7D4D80D05F3FFFCAFC51DF9BFB83F99C" author="MIYAKE" box="[977,1136,1514,1538]" pageId="8" pageNumber="227" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in color. Vol. II. Brachyura (Crabs). Hoikusha, Osaka, viii + 277 pp., 64 pls. [in Japanese]" type="book" year="1983">MIYAKE 1983</bibRefCitation>
), MALAYSIAN BORNEO—Off PONTIANAK (
<bibRefCitation id="7D4D80D05F3FFFCAFE9FDC7CFE4CF9B8" author="SERENE" box="[287,447,1549,1574]" pageId="8" pageNumber="227" refString="Serene, R. (1971) Rediscovery of Portunus nipponensis (Sakai, 1938). Researches on Crustacea, 4, 71 - 74." type="journal article" year="1971">SERÈNE 1971</bibRefCitation>
); INDONESIA—BANDA &amp; TAJAUDE (=TALAUD?) ISLANDS (
<bibRefCitation id="7D4D80D05F3FFFCAFBCDDC7FFAD3F9B8" author="STEPHENSON" box="[1101,1312,1550,1574]" pageId="8" pageNumber="227" refString="Stephenson, W. (1972) Portunid crabs from the Indo-West-Pacific and Western America in the Zoological Museum, Copenhagen (Decapoda, Brachyura, Portunidae). Steenstrupia, 2 (9), 127 - 156." type="journal article" year="1972">STEPHENSON 1972</bibRefCitation>
; TUAMOTU ARcHIPELAgO—MURUROA (
<bibRefCitation id="7D4D80D05F3FFFCAFE2FDC40FDBBF9D7" author="POUPIN" box="[431,584,1584,1609]" pageId="8" pageNumber="227" refString="Poupin, J. (1996) Atlas des crustaces marins profonds de Polynesie Francaise recoltes du navire Marara (1986 / 1996). Service Mixte de Surveillance Radiologique et Biologique, Montlhery, 59 pp." type="book" year="1996">POUPIN 1996</bibRefCitation>
: PL. 17 fIgS. b, c; 
<bibRefCitation id="7D4D80D05F3FFFCAFC9ADC40FBC4F9D7" author="CROSNIER" box="[794,1079,1585,1609]" pageId="8" pageNumber="227" refString="Crosnier, A. &amp; Moosa, M. K. (2002) Trois Portunidae (Crustacea, Decapoda, Brachyura) noveaux de Polynesie francaise. Zoosystema, 24 (2), 385 - 399." type="journal article" year="2002">CROSNIER &amp; MOOSA 2002</bibRefCitation>
); cENTRAL PHILIPPINES (Ng 
<emphasis id="2BA821335F3FFFCAFAE1DC40FA66F9D7" box="[1377,1429,1585,1609]" italics="true" pageId="8" pageNumber="227">et al</emphasis>
. 2008: fIg. 117); AND EASTER ISLAND (
<bibRefCitation id="7D4D80D05F3FFFCAFDA2DC25FCC2F9F2" author="BOYKO" box="[546,817,1620,1644]" pageId="8" pageNumber="227" refString="Boyko, C. B. &amp; Liguori, A. (2014) SWimming crabs (Crustacea: Brachyura: Portunoidea) of Easter Island. Pacific Science, 68 (4), 563 - 575. https: // doi. org / 10.2984 / 68.4.10" type="journal article" year="2014">BOYKO &amp; LIgUORI 2014</bibRefCitation>
). FURTHERMORE, 
<taxonomicName id="DEDC86A25F3FFFCAFC60DC24FB76F9F2" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[992,1157,1621,1644]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFC60DC24FB76F9F2" box="[992,1157,1621,1644]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
HAS bEEN cOLLEcTED fROM SHALLOWER DEPTHS AS WELL AS fROM DEEP WATER (12–250 M). FOR INSTANcE, bOTH SPEcIMENS fROM RYUKYUS EXAMINED HERE (1 ADULT MALE AND 1 JUVENILE fEMALE) WERE cOLLEcTED bY DIVERS IN SUbMARINE cAVES AT A DEPTH Of APPROXIMATELY 30 M. THE HOLOTYPE Of 
<taxonomicName id="DEDC86A25F3FFFCAFE1CDCCEFD88F948" ID-CoL="4M2P7" authorityName="Edmondson" authorityYear="1954" box="[412,635,1726,1750]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="oahuensis">
<emphasis id="2BA821335F3FFFCAFE1CDCCEFD88F948" box="[412,635,1726,1750]" italics="true" pageId="8" pageNumber="227">Portunus oahuensis</emphasis>
</taxonomicName>
<bibRefCitation id="7D4D80D05F3FFFCAFD03DCCEFCA3F949" author="EDMONDSON" box="[643,848,1726,1751]" pageId="8" pageNumber="227" refString="Edmondson, C. H. (1954) HaWaiian Portunidae. Occasional Papers of the Bernice P. Bishop Museum, 21 (12), 217 - 274." type="journal article" year="1954">EDMONDSON, 1954</bibRefCitation>
(= 
<taxonomicName id="DEDC86A25F3FFFCAFCF9DCCEFBEEF948" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[889,1053,1727,1750]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFCF9DCCEFBEEF948" box="[889,1053,1727,1750]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
), WAS REPORTEDLY cOLLEcTED fROM A SHALLOW REEf (VIZ. 
<bibRefCitation id="7D4D80D05F3FFFCAFEE5DC93FDDCF965" author="EDMONDSON" box="[357,559,1762,1787]" pageId="8" pageNumber="227" refString="Edmondson, C. H. (1954) HaWaiian Portunidae. Occasional Papers of the Bernice P. Bishop Museum, 21 (12), 217 - 274." type="journal article" year="1954">EDMONDSON 1954</bibRefCitation>
: 245). STEPHENSON’S (1972) REcORD fROM THE TALAUD (MISSPELLED AS “TAJAUDE”) AND BANDA ISLANDS, IN INDONESIA (NOT PHILIPPINES AS MISTAKENLY REPORTED), cONSISTS Of fOUR JUVENILE SPEcIMENS cOLLEcTED AT A DEPTH Of 15–25 
<bibRefCitation id="7D4D80D05F3FFFCAFE6EDD58FCDCF8DF" author="BOYKO" box="[494,815,1833,1857]" pageId="8" pageNumber="227" refString="Boyko, C. B. &amp; Liguori, A. (2014) SWimming crabs (Crustacea: Brachyura: Portunoidea) of Easter Island. Pacific Science, 68 (4), 563 - 575. https: // doi. org / 10.2984 / 68.4.10" type="journal article" year="2014">M. BOYKO &amp; LIgUORI (2014)</bibRefCitation>
REPORTED TWO SPEcIMENS fROM EASTER ISLAND; ONE ADULT MALE WAS cOLLEcTED fROM A SUbMARINE cAVE AT A DEPTH Of 12 M, WHILE A JUVENILE MALE WAS cOLLEcTED fROM bENEATH A DOcK AT A DEPTH Of 15 M.
</paragraph>
<paragraph id="1963FD215F3FFFCBFF47DDE5FCA5FF69" blockId="8.[151,1437,151,2035]" lastBlockId="9.[151,1436,151,247]" lastPageId="9" lastPageNumber="228" pageId="8" pageNumber="227">
THERE ARE ALSO REcORDS Of 
<taxonomicName id="DEDC86A25F3FFFCAFE64DDE5FD7BF835" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[484,648,1940,1963]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFE64DDE5FD7BF835" box="[484,648,1940,1963]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
fROM THE INDIAN OcEAN. THE REcORD Of 
<bibRefCitation id="7D4D80D05F3FFFCAFBB9DDE5FAACF832" author="KATHIRVEL" box="[1081,1375,1940,1964]" pageId="8" pageNumber="227" refString="Kathirvel, M. &amp; Gokul, A. (2010) Check list of brachyuran crabs from the Gulf of Mannar Marine Biosphere Reserve. Revised Edition. The Fisheries Technocrats Forum Technical Bulletin, 4, 1 - 41." type="journal article" year="2010">KATHIRVEL &amp; GOKUL (2010)</bibRefCitation>
fROM THE GULf Of MANNAR cOULD NOT bE cONfIRMED 
<collectingCountry id="61CBBDB15F3FFFCAFD16DDC6FD5EF851" box="[662,685,1975,1999]" name="American Samoa" pageId="8" pageNumber="227">AS</collectingCountry>
THERE IS NO EVIDENcE THAT VOUcHER SPEcIMENS WERE DEPOSITED IN A REfERENcE cOLLEcTION. 
<bibRefCitation id="7D4D80D05F3FFFCAFE02DDAAFDD1F86D" author="POUPIN" box="[386,546,2011,2035]" pageId="8" pageNumber="227" refString="Poupin, J. (2010) Biodiversite de l'Indo-Pacifique tropical francais: 2514 especes de crustaces decapodes et stomatopodes. Rapport scientifique de l'Institut de Recherche de l'Ecole Navale, 76 pp. [Octobre 2010]" type="book" year="2010">POUPIN (2010)</bibRefCitation>
ALSO REPORTS 
<taxonomicName id="DEDC86A25F3FFFCAFD31DDAAFCA7F86C" ID-CoL="3RYC2" baseAuthorityName="Sakai" baseAuthorityYear="1938" box="[689,852,2011,2034]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="227" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="2BA821335F3FFFCAFD31DDAAFCA7F86C" box="[689,852,2011,2034]" italics="true" pageId="8" pageNumber="227">L. nipponensis</emphasis>
</taxonomicName>
fROM 
<collectingCountry id="61CBBDB15F3FFFCAFC17DDAAFC07F86D" box="[919,1012,2011,2035]" name="Mayotte" pageId="8" pageNumber="227">MAYOTTE</collectingCountry>
(DEPTH: 15–30 M) IN THE 
<collectingCountry id="61CBBDB15F3FFFCAFA88DDABFA6FF86D" box="[1288,1436,2010,2035]" name="Mozambique" pageId="8" pageNumber="227">MOZAMbIqUE</collectingCountry>
CHANNEL. HIS SPEcIMEN, A YOUNg MALE (SEE HTTP://cRUSTAcEAMAYOTTE.fREE.fR/ILLUSTRATION.PHP?N=1&amp;SP=398), APPEARS TO bE A SPEcIES Of 
<taxonomicName id="DEDC86A25F3EFFCBFEECDACAFE01FF4D" ID-CoL="7P858" authorityName="Devi" authorityYear="2017" box="[364,498,187,211]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="2BA821335F3EFFCBFEECDACAFE01FF4D" box="[364,498,187,211]" italics="true" pageId="9" pageNumber="228">Laleonectes</emphasis>
</taxonomicName>
, bUT THE SPEcIES cANNOT bE cONfIRMED 
<collectingCountry id="61CBBDB15F3EFFCBFC2DDACDFC37FF4A" box="[941,964,188,212]" name="American Samoa" pageId="9" pageNumber="228">AS</collectingCountry>
IT HAS NOT bEEN EXAMINED bY THE PRESENT AUTHORS. THESE REcORDS MAY REfER TO 
<taxonomicName id="DEDC86A25F3EFFCBFDAADA91FD63FF69" ID-CoL="84FXK" authority="Devi, 2017" authorityName="Devi" authorityYear="2017" box="[554,656,223,247]" class="Malacostraca" family="Portunidae" genus="Laleonectes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="228" phylum="Arthropoda" rank="species" species="kuriya" status="sp. nov.">
<emphasis id="2BA821335F3EFFCBFDAADA91FD63FF69" box="[554,656,223,247]" italics="true" pageId="9" pageNumber="228">L. kuriya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="309B9C485F3EFFCBFD16DAAFFD03FF69" box="[662,752,222,247]" pageId="9" pageNumber="228" rank="species">
<emphasis id="2BA821335F3EFFCBFD16DAAFFD03FF69" bold="true" box="[662,752,222,247]" pageId="9" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
, INSTEAD.
</paragraph>
</subSubSection>
</treatment>
</document>