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<document ID-CLB-Dataset="269602" ID-DOI="http://dx.doi.org/10.3897/dez.70.107425" ID-GBIF-Dataset="360868d3-9133-466c-bfae-e8de48028017" ID-Pensoft-Pub="1860-1324-2-311" ID-Pensoft-UUID="65D0B2390EBE553382C9727ECC9CBF8F" ID-ZooBank="8D3E277C424C440B8FE878085239C2A2" ModsDocID="1860-1324-70-2-311" checkinTime="1695893248727" checkinUser="pensoft" docAuthor="Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek &amp; Schmidt, Joachim" docDate="2023" docId="23F4CEDA79AC5113BA0F78816C8FFDA3" docLanguage="en" docName="DeutEntomolZeit 70(2): 311-335" docOrigin="Deutsche Entomologische Zeitschrift 70 (2)" docPubDate="2023-09-27" docSource="http://dx.doi.org/10.3897/dez.70.107425" docTitle="Trechus (Abunetrechus) bipartitus Raffray 1885" docType="treatment" docVersion="2" id="65D0B2390EBE553382C9727ECC9CBF8F" lastPageNumber="311" masterDocId="65D0B2390EBE553382C9727ECC9CBF8F" masterDocTitle="Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini)" masterLastPageNumber="335" masterPageNumber="311" pageNumber="311" updateTime="1695893870965" updateUser="ExternalLinkService">
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<mods:title id="BA85FCB98769D39FCDC6D4F440704FD3">Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini)</mods:title>
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<mods:namePart id="118442FC328F97945FA146CF44791105">Faille, Arnaud</mods:namePart>
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<mods:affiliation id="CB133226C870E13031243C6F9F2FA23B">Department of Entomology, Stuttgart State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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<mods:namePart id="8ACD59F651C908230FA0E7983BDB1E82">Hofmann, Sylvia</mods:namePart>
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<mods:namePart id="A3EF2F42440DF5398CF6690AAEF854C7">Merene, Yeshitla</mods:namePart>
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<mods:affiliation id="69E81DEDE059A2B1C98D9A11A8B8D759">Faculty of Biology, Philipps University Marburg, Marburg, Germany &amp; Department of Zoological Sciences, Addis Ababa University, Addis Ababa, Ethiopia &amp; Amhara Agricultural Research Institute, Bahir Dar, Ethiopia</mods:affiliation>
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<mods:namePart id="4922EE75627A66FBA2F9819BD22D5453">Hauth, David</mods:namePart>
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<mods:namePart id="395BAC7E00C642759004752FC4BE13BB">Opgenoorth, Lars</mods:namePart>
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<mods:namePart id="55032A0662F4EC761FB1A421975D8C93">Schmidt, Joachim</mods:namePart>
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<treatment id="23F4CEDA79AC5113BA0F78816C8FFDA3" LSID="urn:lsid:plazi:treatment:23F4CEDA79AC5113BA0F78816C8FFDA3" httpUri="http://treatment.plazi.org/id/23F4CEDA79AC5113BA0F78816C8FFDA3" lastPageNumber="311" pageId="0" pageNumber="311">
<subSubSection id="B8291843A81EFB37F6DBCDBD3FFCC3F9" pageId="0" pageNumber="311" type="nomenclature">
<paragraph id="C927C69D5CA0048A0E3B83F28B575079" pageId="0" pageNumber="311">
<taxonomicName id="AC21A1C8938CEA333DC420EAE37B9BFA" ID-CoL="7CNNM" LSID="23F4CEDA-79AC-5113-BA0F-78816C8FFDA3" authority="Raffray, 1885" authorityName="Raffray" authorityYear="1885" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus) bipartitus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="bipartitus" subGenus="Abunetrechus">Trechus (Abunetrechus) bipartitus Raffray, 1885</taxonomicName>
</paragraph>
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<paragraph id="BE8BFBAA856DF82C0403ACB7BF09602E" pageId="0" pageNumber="311">
<figureCitation id="F2C0AED19005C8AF14D8B074D15C786D" captionStart="Figures 17–20" captionStartId="F7" captionText="Figures 17 - 20. Trechus subgenus Abunetrechus nov., dorsal aspect of body (17, 19), head and pronotum (18, 20). 17, 18. T. bipartitus Raffray; 19, 20. T. lalibelae Queinnec &amp; Ollivier. The small white circles in Figs 18 and 20 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures17-20" httpUri="https://binary.pensoft.net/fig/913117" pageId="0" pageNumber="311">Figs 17</figureCitation>
<figureCitation id="77588D7898E5331508DD5C591728FAE2" captionStart="Figures 17–20" captionStartId="F7" captionText="Figures 17 - 20. Trechus subgenus Abunetrechus nov., dorsal aspect of body (17, 19), head and pronotum (18, 20). 17, 18. T. bipartitus Raffray; 19, 20. T. lalibelae Queinnec &amp; Ollivier. The small white circles in Figs 18 and 20 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures17-20" httpUri="https://binary.pensoft.net/fig/913117" pageId="0" pageNumber="311">, 18</figureCitation>
<figureCitation id="2B22B0C53A5913F8109896F41CF37A67" captionStart="Figures 21–23" captionStartId="F8" captionText="Figures 21 - 23. Trechus subgenus Abunetrechus nov., dorsal aspect of body (21) and head (22), and anterior part of left elytron (23). 21, 22. T. habeshaicus Queinnec &amp; Ollivier; 23. T. bipartitus Raffray. The small white circles in Fig. 22 mark the insertion points of the clypeal setae; the arrow in Fig. 23 points to the insertion of the anterior elytral discal seta." figureDoi="10.3897/dez.70.107425.figures21-23" httpUri="https://binary.pensoft.net/fig/913118" pageId="0" pageNumber="311">, 23</figureCitation>
<figureCitation id="153F1D1FC106F1C709D182226641B9CF" captionStart="Figures 24–29" captionStartId="F9" captionText="Figures 24 - 29. Trechus subgenus Abunetrechus nov., aedeagus in left lateral aspect (24, 26, 28) and dorsal aspect (25, 27, 29). 24, 25. T. bipartitus Raffray; 26, 27. T. lalibelae Queinnec &amp; Ollivier; 28, 29. T. habeshaicus Queinnec &amp; Ollivier." figureDoi="10.3897/dez.70.107425.figures24-29" httpUri="https://binary.pensoft.net/fig/913119" pageId="0" pageNumber="311">, 24-25</figureCitation>
</paragraph>
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<subSubSection id="5048A21B8F7F331FBABE3599B6494B90" pageId="0" pageNumber="311" type="reference_group">
<paragraph id="C0176369903561D569189AC0D94D3993" pageId="0" pageNumber="311">
<taxonomicName id="62B3A9898FFB5B81AB8C2B520884E49D" authorityName="Raffray" authorityYear="1885" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus bipartitus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="bipartitus">Trechus bipartitus</taxonomicName>
Raffray, 1885: 318; locus typicus: &quot;col du mont Abouna-Yousef (4024 m)&quot;.
</paragraph>
<paragraph id="CAED9C0957312FB1AFCAB5761BF590A1" pageId="0" pageNumber="311">
<taxonomicName id="795CCFEC22FD0B4D75FB206CF5F57AD5" authorityName="Raffray" authorityYear="1885" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus bipartitus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="bipartitus">Trechus bipartitus</taxonomicName>
:
<bibRefCitation id="B2DB358F524CD0CF614D108EC0B4C372" author="Jeannel, R" journalOrPublisher="Journal d'Entomologie" pageId="0" pageNumber="311" pagination="1 - 592" refId="B21" refString="Jeannel, R, 1927. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Deuxieme livraison). L'Abeille. Journal d'Entomologie 33: 1 - 592" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Deuxieme livraison). L'Abeille." volume="33" year="1927">Jeannel 1927</bibRefCitation>
: 194.
</paragraph>
<paragraph id="ED6285CC973E22930894304D6D8449C8" pageId="0" pageNumber="311">
<taxonomicName id="DFB89CC60A40254EC14377CB7A297093" authorityName="Raffray" authorityYear="1885" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus bipartitus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="bipartitus">Trechus bipartitus</taxonomicName>
:
<bibRefCitation id="69F3475F7BA958496C6072E60D40836C" author="Pawlowski, J" journalOrPublisher="Baltic Journal of Coleopterology" pageId="0" pageNumber="311" pagination="157 - 160" refId="B41" refString="Pawlowski, J, 2003. Preliminary report on Ethiopian species of the genus Trechus Clairville, 1806 (Coleoptera: Carabidae). Baltic Journal of Coleopterology 3 (2): 157 - 160" title="Preliminary report on Ethiopian species of the genus Trechus Clairville, 1806 (Coleoptera: Carabidae)." volume="3" year="2003">
<normalizedToken id="CEAABFD6FD59E9F04D1AAD655DFF8A9F" originalValue="Pawłowski">Pawlowski</normalizedToken>
2003
</bibRefCitation>
: 157.
</paragraph>
<paragraph id="3F5D1453BB2DD6D10E445B926AD8381C" pageId="0" pageNumber="311">
<taxonomicName id="44227EE50C13D491125C1195287B3664" authorityName="Raffray" authorityYear="1885" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus bipartitus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="bipartitus">Trechus bipartitus</taxonomicName>
:
<bibRefCitation id="F5363E5443A26D6E540D2ED1EE7B1ADF" DOI="https://doi.org/10.1603/AN10122" author="Ortuno, VM" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="311" pagination="132 - 140" refId="B39" refString="Ortuno, VM, Novoa, F, 2011. A new species of Trechus from the Ethiopian Highlands (Coleoptera: Carabidae: Trechinae) and key to the Trechus species of Ethiopia. Annals of the Entomological Society of America 104 (2): 132 - 140, DOI: https://doi.org/10.1603/AN10122" title="A new species of Trechus from the Ethiopian Highlands (Coleoptera: Carabidae: Trechinae) and key to the Trechus species of Ethiopia." url="https://doi.org/10.1603/AN10122" volume="104" year="2011">
<normalizedToken id="A1CFC763E25AF28AACCECB5DA758253E" originalValue="Ortuño">Ortuno</normalizedToken>
and Novoa 2011
</bibRefCitation>
: 137.
</paragraph>
<paragraph id="6727D53E59A916037618067585A88105" pageId="0" pageNumber="311">
<taxonomicName id="D76BE140DEA0A57901656947B02CA3A0" authorityName="Raffray" authorityYear="1885" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus bipartitus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="bipartitus">Trechus (s. str.) bipartitus</taxonomicName>
:
<bibRefCitation id="F09222C3F825AEA70596AF8FC813DC0C" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="CCA3FF49345DD01062A432AB6568F68E" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: 20.
</paragraph>
</subSubSection>
<subSubSection id="1D31301EC1473BAA66C4747D120A95FF" pageId="0" pageNumber="311" type="type material">
<paragraph id="08DDBE1FC5E5EA096132C24F496D5EDA" pageId="0" pageNumber="311">Type material.</paragraph>
<paragraph id="B03F2128E8D105E6B44615A083DE39F7" pageId="0" pageNumber="311">
Not studied. The lectotype was designated by
<bibRefCitation id="E2865F7347412B1F6AE15C638F19DB19" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="ACF96742BCD78165D4BBD53584A77273" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. (2021)
</bibRefCitation>
. Identification is based on the redescriptions of the species, including habitus and male genital figures of the type specimens, presented by
<bibRefCitation id="70C4D1264BEA17F907B685DC4DC8D050" author="Jeannel, R" journalOrPublisher="Journal d'Entomologie" pageId="0" pageNumber="311" pagination="1 - 592" refId="B21" refString="Jeannel, R, 1927. Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Deuxieme livraison). L'Abeille. Journal d'Entomologie 33: 1 - 592" title="Monographie des Trechinae. Morphologie comparee et distribution geographique d'un groupe de Coleopteres (Deuxieme livraison). L'Abeille." volume="33" year="1927">Jeannel (1927)</bibRefCitation>
and
<bibRefCitation id="A151FC83C35DDD0200F7B34E60B345E2" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="094E625724C0916E28F011C2BACF05BD" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. (2021)
</bibRefCitation>
, as well as on comprehensive material collected at the type locality (see below).
</paragraph>
</subSubSection>
<subSubSection id="36BA87E6F5454EC009997D2E7FA3BFDC" pageId="0" pageNumber="311" type="materials_examined">
<paragraph id="0107D19BDF67DE08817E9B0B5262294A" pageId="0" pageNumber="311">Additional material.</paragraph>
<paragraph id="3F5EFBCDB06C040069481E896F8EE53E" pageId="0" pageNumber="311">
<materialsCitation id="AD702ADC32A9E34F951F90838B1FE7B6" collectingDate="2019-03-04" collectorName="D. Hauth, Hauth, J, Yeshitla M., Yitbarek W." country="Ethiopia" elevation="3875" latitude="12.131111" location="Mt. Abuna Yosef" longLatPrecision="21" longitude="39.194168" specimenCount="15" specimenCount-female="9" specimenCount-male="6" stateProvince="Amhara">
<specimenCount id="279AB68051A0CE36EE48249A904F4496" type="male">6 males</specimenCount>
,
<specimenCount id="6356683BB49DF4C292BCA3344C3D8C98" type="female">9 females</specimenCount>
,
<collectingCountry id="407F8A7BCC0A6F9BF3F8578C662AEDCA" name="Ethiopia">Ethiopia</collectingCountry>
,
<collectingRegion id="4F5F6D1FBC1FB707E9A484FD41509AAF" country="Ethiopia" name="Amara">Amhara</collectingRegion>
,
<location id="D2E5C9476CB092FA64994E9085B09179" LSID="urn:lsid:plazi:treatment:23F4CEDA79AC5113BA0F78816C8FFDA3:D2E5C9476CB092FA64994E9085B09179" country="Ethiopia" latitude="12.131111" longLatPrecision="21" longitude="39.194168" name="Mt. Abuna Yosef" stateProvince="Amhara">Mt. Abuna Yosef</location>
, N-slope,
<quantity id="C0BD77CA4AE7BD5CF5CBBE899318CBD7" metricMagnitude="3" metricUnit="m" metricValue="3.875" metricValueMax="3.95" metricValueMin="3.8" unit="m" value="3875.0" valueMax="3950.0" valueMin="3800.0">
<elevation id="FAC24625F9AB7B75C7EDA985F37DDA27" metricMagnitude="3" metricUnit="m" metricValue="3.875" metricValueMax="3.95" metricValueMin="3.8" unit="m" value="3875.0" valueMax="3950.0" valueMin="3800.0">3800-3950 m</elevation>
</quantity>
,
<geoCoordinate id="FAD2AA1627B631E648367FCFB29BFAB7" degrees="12" direction="north" minutes="07" orientation="latitude" precision="15" seconds="52" value="12.131111">12°07'52&quot;N</geoCoordinate>
,
<geoCoordinate id="0F96AEDF51DACE208766C34CEEF099F0" degrees="39" direction="east" minutes="11" orientation="longitude" precision="15" seconds="39" value="39.194168">39°11'39&quot;E</geoCoordinate>
,
<collectingDate id="A16FA1E148B0EF1EC282E5A263150042" value="2019-03-04">4.III.2019</collectingDate>
, leg.
<collectorName id="002754E51019BC9DE7A480782D7929B2">
D.
<collectorName id="372667BAD5128A3239E522CD61799044">Hauth, J</collectorName>
</collectorName>
. Schmidt,
<collectorName id="CD4C94C5C77337F2BBB4F17BF425A6D4">Yeshitla M.</collectorName>
,
<collectorName id="759F6EFEF1385D856534035ECBD88E96">Yitbarek W.</collectorName>
(CAF, CSCHM)
</materialsCitation>
;
<materialsCitation id="E5576E67A32F07F113D5AC4FC77D803A" collectingDate="2019-03-05" collectorName="D. Hauth, Hauth, J, Yeshitla M., Yitbarek W." country="Ethiopia" elevation="3875" latitude="12.1247225" location="Mt. Abuna Yosef" longLatPrecision="21" longitude="39.189167" specimenCount="18" specimenCount-female="12" specimenCount-male="6" stateProvince="Amara">
<specimenCount id="CC73B9B856E059DAA31EC1C229C7A0CB" type="male">6 males</specimenCount>
,
<specimenCount id="10C05D41AEF92AECFCE53340CFB0637E" type="female">12 females</specimenCount>
, ditto, S-slope,
<quantity id="8A8158D5ABFC451C4E7EA5C4932AF40A" metricMagnitude="3" metricUnit="m" metricValue="3.875" metricValueMax="3.9" metricValueMin="3.85" unit="m" value="3875.0" valueMax="3900.0" valueMin="3850.0">
<elevation id="A927324A36FBFF50B7F9F7F3F787F498" metricMagnitude="3" metricUnit="m" metricValue="3.875" metricValueMax="3.9" metricValueMin="3.85" unit="m" value="3875.0" valueMax="3900.0" valueMin="3850.0">3850-3900 m</elevation>
</quantity>
,
<geoCoordinate id="F2ACFE40299C41C82C23B4A563DAEFAD" degrees="12" direction="north" minutes="07" orientation="latitude" precision="15" seconds="29" value="12.1247225">12°07'29&quot;N</geoCoordinate>
,
<geoCoordinate id="362B7596CDEDB8D097167C1ECD6E3645" degrees="39" direction="east" minutes="11" orientation="longitude" precision="15" seconds="21" value="39.189167">39°11'21&quot;E</geoCoordinate>
,
<collectingDate id="B0DA9E90A925BD01B37ECAA162D7189E" value="2019-03-05">5.III.2019</collectingDate>
, leg.
<collectorName id="09B9AB761C2E9F9D205CEF097CC67398">
D.
<collectorName id="D92F83E3F22697071917035D7CFACB79">Hauth, J</collectorName>
</collectorName>
. Schmidt,
<collectorName id="C1EC7395A171B144E11E9732818DCBB9">Yeshitla M.</collectorName>
,
<collectorName id="00712D658B94F6360981D74F837DC3D7">Yitbarek W.</collectorName>
(CSCHM, NHMAA)
</materialsCitation>
;
<materialsCitation id="4E1A93615F2E5EF09D3C97EF64E5C70B" collectingDate="2019-03-05" collectorName="D. Hauth, Hauth, J, Yeshitla M., Yitbarek W." country="Ethiopia" elevation="3775" latitude="12.152778" location="Mt. Abuna Yosef" longLatPrecision="21" longitude="39.159725" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Amara">
<specimenCount id="AD7510033BB3CBBEE42C64E6A1AC20D9" type="male">5 males</specimenCount>
,
<specimenCount id="8E021D0B00B0B17FAADED5C405D9D579" type="female">2 females</specimenCount>
, ditto, S-slope,
<quantity id="A2FD4D02FEE9A1F59DBFF00A14904C8A" metricMagnitude="3" metricUnit="m" metricValue="3.775" metricValueMax="3.85" metricValueMin="3.7" unit="m" value="3775.0" valueMax="3850.0" valueMin="3700.0">
<elevation id="1E6D3B87DAB8325236DD195546EAC368" metricMagnitude="3" metricUnit="m" metricValue="3.775" metricValueMax="3.85" metricValueMin="3.7" unit="m" value="3775.0" valueMax="3850.0" valueMin="3700.0">3700-3850 m</elevation>
</quantity>
,
<geoCoordinate id="741B90F614D848BA165B66351764301A" degrees="12" direction="north" minutes="09" orientation="latitude" precision="15" seconds="10" value="12.152778">12°09'10&quot;N</geoCoordinate>
,
<geoCoordinate id="FBE13E961795D507AD9B9E48935A465D" degrees="39" direction="east" minutes="09" orientation="longitude" precision="15" seconds="35" value="39.159725">39°09'35&quot;E</geoCoordinate>
,
<collectingDate id="B0EBC8216E6747B49D9967F0DD3A9595" value="2019-03-05">5.III.2019</collectingDate>
, leg.
<collectorName id="0988550FF3A3A4F552236384A397FB71">
D.
<collectorName id="E2BE57FC56398B003BFD7282118B2E66">Hauth, J</collectorName>
</collectorName>
. Schmidt,
<collectorName id="799081E3F42C2EDB6EC26B2FFD2241C9">Yeshitla M.</collectorName>
,
<collectorName id="F9FF6227F731408B21F5ED5953C838D9">Yitbarek W.</collectorName>
(CSCHM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E6A23F01686982EFDACE5F3AE4DF4584" pageId="0" pageNumber="311" type="description">
<paragraph id="05A763E00D58660AF38C894A30C12819" pageId="0" pageNumber="311">Additions to the species description.</paragraph>
<paragraph id="D49B28779FD4BA3DA134DF3103ED05B3" pageId="0" pageNumber="311">
Mature species with elytra including suture blackish brown (
<bibRefCitation id="842CD9E7587C36A164E97DDFF455BCDB" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="D8DF670A11C44C84D086D90C28795989" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: &quot;suture distinctly brown shiny&quot;; this information is probably based on confusion with
<taxonomicName id="DC7F66B4EB678A512957040540110A0F" family="Carabidae" kingdom="Animalia" lsidName="T. habeshanicus" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="habeshanicus">
<emphasis id="CC24E990EA0642F5E15F76FE23E83035" italics="true" pageId="0" pageNumber="311">T. habeshanicus</emphasis>
</taxonomicName>
). Body length 4.1-4.7 mm (
<bibRefCitation id="30AB7CB435422890B5B87BDC7477E132" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="08659619E88DD63963E0B0C89D1013F5" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: &quot;&gt; 4.5 mm&quot;). Length of aedeagus 0.98-1.05 mm. PW/PL = 1.29-1.37; n = 20 (
<bibRefCitation id="C42D75C18A492B8809791C38D7E0C2EA" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="4E84A99FB6A982E28A7950B5CFBD80F1" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: PW/PL = 1.6; this value is very probably based on a measurement error). PW/TL = 1.06-1.15; n = 20. EL/AL = 2.15-2.46; n = 10.
</paragraph>
</subSubSection>
<subSubSection id="95452BB5EEBC92C6031B17AC70DADC33" pageId="0" pageNumber="311" type="differential diagnosis">
<paragraph id="CB90641064EF00F17152584C40746167" pageId="0" pageNumber="311">Differential diagnosis.</paragraph>
<paragraph id="707E878E16556AE2FF2C3C79BDF8E099" pageId="0" pageNumber="311">
See Key to species of the subgenus
<taxonomicName id="049C316E8C46A4929C354CCDDCB6A1D0" authorityName="Schmidt &amp; Faille" authorityYear="2023" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus)" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subGenus" subGenus="Abunetrechus">Trechus Abunetrechus</taxonomicName>
, below.
</paragraph>
</subSubSection>
<subSubSection id="30B16BAB9D0840FE4B8B8AB4DA6EF338" pageId="0" pageNumber="311" type="distribution">
<paragraph id="C9DA8114185A7144B6CB787CCA3716DB" pageId="0" pageNumber="311">Distribution.</paragraph>
<paragraph id="DAD8F03461CAF158499F93A132B72DA4" pageId="0" pageNumber="311">Endemic to the Abune Yosef Mountains of the northern Ethiopian Highlands.</paragraph>
</subSubSection>
<subSubSection id="7423438DA8DD3CCDC245D0487BAECE34" pageId="0" pageNumber="311" type="habitat">
<paragraph id="F96C8A854910A040C40D7F50EE6C2C02" pageId="0" pageNumber="311">Habitat.</paragraph>
<paragraph id="FDA72EC477C7A4F6A3A83FAC5062C0CE" pageId="0" pageNumber="311">
As in
<taxonomicName id="953EEA1DD76653BF75130280BB2760CB" family="Carabidae" kingdom="Animalia" lsidName="T. (Abunetrechus) lalibelae" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="lalibelae" subGenus="Abunetrechus">T. (Abunetrechus) lalibelae</taxonomicName>
<normalizedToken id="BD03916838FED3E3C684311183D1080D" originalValue="Quéinnec">Queinnec</normalizedToken>
&amp; Ollivier (see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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