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51 KiB
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510 lines
51 KiB
XML
<document ID-CLB-Dataset="21607" ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="9FED202A658955E48A38CF45E18212D9" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis tolucensis Kunth, Nov. Gen. Sp. [H. B. K.] 1: 135. 1816" docType="treatment" docVersion="6" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1732688126400" updateUser="ExternalLinkService">
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<mods:mods id="81884D5341047B276E0E60CD09457C28" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8D9E377562D05F43A1E62D99DE14EDFB">
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<mods:title id="9AB8485530E0E8B5CE924B8A90ACE879">Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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</mods:titleInfo>
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<mods:name id="7616489601CE872142CD0863FFC49A9F" type="personal">
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<mods:roleTerm id="6594868864043BC0E56B82F6A7EA6A7D">Author</mods:roleTerm>
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<mods:namePart id="86E1DDD3A0E1089764463F4B965EA01A">Sylvester, Steven P.</mods:namePart>
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<mods:affiliation id="6B23DBA2076E291F6BA249214FCCC108">College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier id="3701925C8A71C0EF5A3CD1625DF7EA56" type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier id="A3B11D89D36F194CCF2042499F8848A4" type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="883D862CDFBE658B229CB54C6CD2C638" type="personal">
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<mods:roleTerm id="8ADBA3AC27440FBB2C2FC5AEEA609A06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="80A27DB5E92FFDA68EF7C1606F9D0DC1">Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation id="5F27CF6270594DA564A168646B410F6D">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name id="62B19D768FF9DEDE68CAACBA79B33D4F" type="personal">
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<mods:role id="D6993E344DF1D4516ADE22E1397CDE5F">
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<mods:roleTerm id="28B36BAF542BFD5466D3B4ED28BB9805">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="009E11A03906115A524550DC4B9C4DD5">Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation id="2E44E25E85957F192ACD671DE540A988">Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name id="AFEEC306AFC3CD45D3AC163A8FCFAB7A" type="personal">
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<mods:roleTerm id="457A26EBA6574F7E0936E53F6FF33B30">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="86D68CF1ACCD6BB5EE36271265E983EF">Soreng, Robert J.</mods:namePart>
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<mods:affiliation id="4DE711471E87B4F012053204B3935F86">Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier id="CE36DA36A99C7630F2935B146FCF697F" type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="BEA7E3D5FAE76A4C3AB62C2D24E1FC05">text</mods:typeOfResource>
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<mods:relatedItem id="3B4A81901D68E40B8EFE3F37C5E25419" type="host">
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<mods:titleInfo id="DFA08F6462D6FAEE87B997E89D9F526B">
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<mods:title id="790C30FE5CA5D325396FCBAAA8D99576">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="2E65D1304C20A566BE791F2D60F9DBC4">
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<mods:date id="DEE8306CF73E41C42D8E616BB7B6D772">2020</mods:date>
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<mods:detail id="C806B53FFF5F66A64A58B440E757DE69" type="volume">
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<mods:number id="36BADDEEAE296B790676A2D5C6692CDA">151</mods:number>
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</mods:detail>
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<mods:extent id="2678A6BDAC096CD59A7EDE7F8127C1E6" unit="page">
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<mods:start id="34D8F47F3B375E1B97D6BAE4F5F338A2">107</mods:start>
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<mods:end id="3E899765613422C5D5850A68759061BD">160</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="4F36464BE01017F883584C4A52FCCC83">
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<mods:url id="4B6B06E2000E450DFFE835EF8224E014">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
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</mods:location>
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<mods:classification id="E09BD376F161C01B476A813D7FD29FF0">journal article</mods:classification>
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<mods:identifier id="D4D9F33FFD0DB1C5F258920FE07EB941" type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<mods:identifier id="C4EC666BD349C9C1C454F5112EF0007C" type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
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<mods:identifier id="BCC5FB5704CBE3EDBCD9F2A436DB1941" type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
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</mods:mods>
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<treatment id="9FED202A658955E48A38CF45E18212D9" ID-GBIF-Taxon="164730850" LSID="urn:lsid:plazi:treatment:9FED202A658955E48A38CF45E18212D9" httpUri="http://treatment.plazi.org/id/9FED202A658955E48A38CF45E18212D9" lastPageNumber="107" pageId="0" pageNumber="107" scope_family="Poaceae">
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<subSubSection id="E64C25BBCB65DF6107168AFCA2B3920D" pageId="0" pageNumber="107" type="nomenclature">
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<paragraph id="E32CD06A1F2A940F0582C98B85F34412" pageId="0" pageNumber="107">
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<taxonomicName id="138BE343740ADC0C59E8D5A8316EA2B3" ID-CoL="5TTHY" LSID="9FED202A-6589-55E4-8A38-CF45E18212D9" authority="Kunth, Nov. Gen. Sp. [H. B. K.] 1: 135. 1816" authorityName="Kunth, Nov. Gen. Sp. [H. B. K.] 1: 135." authorityYear="1816" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">Agrostis tolucensis Kunth, Nov. Gen. Sp. [H.B.K.] 1: 135. 1816</taxonomicName>
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<figureCitation id="EC44C5076BCE9CA36FEB7459F12F245C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agrostis tolucensis A spikelet, glumes in lateral view, floret in dorsal view, detached and raised above the glumes B whole plant. Images of Cuta-Alarcon 376 (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure10" httpUri="https://binary.pensoft.net/fig/424848" pageId="0" pageNumber="107">Fig. 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="ABC123E7F2A17CD203C346702DCFB05F" pageId="0" pageNumber="107" type="reference_group">
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<paragraph id="8627CE55C132924D71F2B2CEDCA21B33" pageId="0" pageNumber="107">
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<taxonomicName id="0337FDD2EAF6305802ECCEE570C0B9B5" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">Agrostis tolucensis</taxonomicName>
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Willd. ex Steud., Syn. Pl. Glumac. 1: 164. 1855[1854], nom. inval.
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</paragraph>
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<paragraph id="DD83A8A2CE28CC9DD67584CD255506F4" pageId="0" pageNumber="107">
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=
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<taxonomicName id="CAD6735F33B3636590B230912A1B1159" authorityName="Kunth" authorityYear="1833" baseAuthorityName="J.Presl" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis glomerata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="glomerata">Agrostis glomerata</taxonomicName>
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(J. Presl) Kunth, Enum. Pl. [Kunth] 1: 219. 1833.
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<taxonomicName id="5B4008959B71428FCBB37A64B837DD69" authorityName="J.Presl" authorityYear="1830" class="Liliopsida" family="Poaceae" genus="Vilfa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vilfa glomerata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="glomerata">Vilfa glomerata</taxonomicName>
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J. Presl, Reliq. Haenk. 1 (4-5): 239. 1830. Type: Peru. [
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<normalizedToken id="30B8120CD2FBEA82DB3784D508E1A3E0" originalValue="Huánuco">Huanuco</normalizedToken>
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] [Hab. in montanis Peruviae huanoccensibus], [1791; 1891 written on W isotype], T.P.X. Haenke s.n. [#196 written on W isotype] (holotype: [not found]; isotypes: HAL (HAL0106916 [image!]), PRC (PRC450953 [image!]), US (US00589472 fragm.), W (W0025326 [image!])).
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</paragraph>
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<paragraph id="BA783A3D3B017ADD188A0C6049072A9D" pageId="0" pageNumber="107">
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=
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<taxonomicName id="6C5DCCFCA9DA676C8C21EEFAD3E822C2" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis hoffmannii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="hoffmannii">Agrostis hoffmannii</taxonomicName>
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Mez, Repert. Spec. Nov. Regni Veg. 18: 3. 1922. Type: Costa Rica. Irazu (holotype: [not found]; isotypes: [not found]).
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</paragraph>
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<paragraph id="3E3A0CDBEABF0A48F1AE846D8007BFDD" pageId="0" pageNumber="107">
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=
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<taxonomicName id="498D9597B8D1C2ACD2ECAFCF0BA006B2" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis nana subsp. var. var. andicola" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="nana" subSpecies="var." variety="andicola">Agrostis nana var. andicola</taxonomicName>
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Pilg., Bot. Jahrb. Syst. 37 (5): 505. 1906. Type: Ecuador. Chimborazo: Chimborazo, N-Seite,
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<normalizedToken id="953AD89766F7846CB78C1315565103AF" originalValue="Páramo-Region">Paramo-Region</normalizedToken>
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, 4500 m alt., July 1903, H. Meyer 146 (syntypes: JE (JE00020226 [image!]; JE00020228 [image!])); Napo: Antisana, in frigidis alpinis, 4600 m alt., July 1903, H. Meyer 145 (syntypes: JE (JE00020225 [image!]; JE00020227 [image!])).
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</paragraph>
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<paragraph id="EA050CD8DFCF7E8FAAEB060414B1BDDE" pageId="0" pageNumber="107">
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=
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<taxonomicName id="479029C495B0366EE0AB0900529FDB44" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis nana subsp. var. var. aristata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="nana" subSpecies="var." variety="aristata">Agrostis nana var. aristata</taxonomicName>
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Griseb., Abh.
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<normalizedToken id="EB9319AF301E7B952D2DEC35F4AF1AF8" originalValue="Königl">Koenigl</normalizedToken>
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. Ges. Wiss.
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<normalizedToken id="E95231937EFA043D9B406F4597F8F1EC" originalValue="Göttingen">Goettingen</normalizedToken>
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24: 294. 1879. Type: Argentina. Salta: Umgebungen des Nevado del Castillo [around the Nevado del Castillo], 10-15000 ft. [3048-4572 m alt.], 19-23 Mar. 1873, P.G. Lorentz & G.H.E.W. Hieronymus 82 (holotype: GOET (GOET006541 [image!]); isotypes: B, BAA (BAA00000724 [image!] fragm. ex B; BAA00000723 [image!] fragm. ex B), CORD (CORD00004693 [image!]), W (W19160036588 [image!]; W19160036664 [image!])).
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</paragraph>
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<paragraph id="010F97ABF6EE207956EC0C6920CD2730" pageId="0" pageNumber="107">
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=
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<taxonomicName id="FE2728661427504FB718D152DFF3166A" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis virescens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="virescens">Agrostis virescens</taxonomicName>
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Kunth, Nov. Gen. Sp. [H.B.K.] 1: 135. 1816. Type: Mexico. Toluca, Sep., F.W.H.A. Humboldt & A.J.A. Bonpland s.n (holotype: P (P00669395 [image!]); isotypes: LE-TRIN (LE-TRIN1668.01; LE-TRIN1668.02), P (P00136912 [image!])).
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</paragraph>
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</subSubSection>
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<subSubSection id="F548F371F32E7C88B430E1B3E9514963" pageId="0" pageNumber="107" type="materials_examined">
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<paragraph id="6D4E2D1FF6732132C2B57976925C6B16" pageId="0" pageNumber="107">Type.</paragraph>
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<paragraph id="C2831DDE4642D723DB98E96C920B074E" pageId="0" pageNumber="107">Mexico. [prope Toluca et Islahuaca], F.W.H.A. Humboldt & A.J.A. Bonpland s.n. (holotype: P (P00669394 [image!]); isotypes: B (B-W1704), BAA (BAA00000235 [image!] fragm. ex P), BM, K (microfiche), LE-TRIN (LE-TRIN1660.01), P (P00136914 [image!]; P00136915 [image!]; P00740426 [image!]), US (US00156505! fragm. ex P)).</paragraph>
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</subSubSection>
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<subSubSection id="4959972467EAFF5392D79965DEAFAE48" pageId="0" pageNumber="107" type="description">
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<paragraph id="DAA8923CED4E3607F6B91ED404B178B1" pageId="0" pageNumber="107">Description.</paragraph>
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<paragraph id="E210D3EC52170073A2AE9EAD6251B2A3" pageId="0" pageNumber="107">
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<emphasis id="43539C08EF856AA2D857BFD86595A3D8" bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
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, tussock-forming or laxly to densely tufted, usually with short ascending rhizomes.
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<emphasis id="881AE8154FABA9BF5765BDFD818052D5" bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
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extravaginal and intravaginal, with cataphylls usually present.
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<emphasis id="A809CB67A346645051459BBD6DC64239" bold="true" pageId="0" pageNumber="107">Culms</emphasis>
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(3-)5.5-51(-80) cm tall, erect or arching, fairly firm, with 0(-2) nodes exerted at flowering, smooth or rarely scaberulous.
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<emphasis id="652C82689F95370AC1C38AA1AA843080" bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
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mostly basal or more-or-less evenly spread along the culm, glabrous, smooth or scabrous;
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<emphasis id="A71131E993BC6A4EAA1E5E3E2FB8D3D8" bold="true" pageId="0" pageNumber="107">ligules</emphasis>
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2-4(-6.2) mm long, of upper culm usually longer than those of lower culm and tillers, truncate to triangular, moderately to strongly decurrent with the sheath, abaxial surface usually scabrous, rarely scaberulous or smooth;
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<emphasis id="7722B6B117B8916C283C4764A68764E2" bold="true" pageId="0" pageNumber="107">blades</emphasis>
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2.5-19 cm long, 1-3(-5) mm wide, at least in the upper culm filiform, folded or flat, usually lax to sometimes firm, sometimes basal and tiller blades involute or convolute and firm to rigid, smooth throughout or scabrous only on the margin and sometimes veins, apices blunt or slightly naviculate-acute.
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<emphasis id="410E84B350E2FA4627F0236188858C79" bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
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(1-)2-15 cm long, 0.1-1.5 cm wide, moderately to densely congested, sub-spikelike or spikelike, sometimes interrupted towards the base, subincluded in the basal foliage or slightly to greatly exerted, lateral branches with spikelets almost to the base, upper lateral branches short and held close to the central inflorescence axis, central axis and panicle branches scabrous;
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<emphasis id="C25858866A7E68F178856298078BF1D5" bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
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0.7-3(-4.5) mm long, usually shorter than their spikelets, not or slightly dilated at their apex, scabrous.
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<emphasis id="A45B69248AEC986564FBE900AEDA37AB" bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
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(not including awn, if present) 2-3(-3.6) mm long;
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<emphasis id="144FDB0140585E6189B052574488B01F" bold="true" pageId="0" pageNumber="107">glumes</emphasis>
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equal or subequal, 1-veined, lower glume keel and surface usually scabrous at least in the distal half, infrequently smooth throughout, upper glume keel and surface scabrous for almost their entirety to at least in the distal half or surface sometimes smooth throughout, apices acute or acuminate;
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<emphasis id="CD3270EF1A191E118666B304908370A3" bold="true" pageId="0" pageNumber="107">floret</emphasis>
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<normalizedToken id="08A7CF0A65FDFEEA4EC53657678207F5" originalValue="½">1/2</normalizedToken>
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-2/3(-3/4) the length of the glumes;
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<emphasis id="EF90DA79E6186A1B57C9460154A7B81D" bold="true" pageId="0" pageNumber="107">calluses</emphasis>
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lightly pilose with 2 sparse tufts of short hairs on the lateral sides;
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<emphasis id="B3918AA1D3DE542798047754EFA0CAEF" bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
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1.4-2.8 mm long, glabrous, smooth, 5-veined, apex truncate, denticulate, usually awned, awn 2-3.5 mm long, inserted in the lower, middle, or upper 1/3 of the dorsal keel, exerted from the glumes, geniculate, twisted, very rarely muticous or with a short straight awn, <1 mm long, inserted in the upper half of the lemma (see notes below regarding
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<taxonomicName id="FFE1EBA54E7CE5D79714267F7077C8F8" lsidName="A. glomerata" pageId="0" pageNumber="107" rank="species" species="glomerata">
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<emphasis id="79D2B5219136ECFFA1F661DDDAAD81BA" italics="true" pageId="0" pageNumber="107">A. glomerata</emphasis>
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</taxonomicName>
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);
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<emphasis id="0EFDA2AE537F9A75650E8156EAD0027F" bold="true" pageId="0" pageNumber="107">paleas</emphasis>
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absent or 0.1-0.2 mm long, <
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<normalizedToken id="3ECC0BB0386FBE503BC5AEC273E43858" originalValue="¼">1/4</normalizedToken>
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the length of the lemma;
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<emphasis id="D187F1C2058BF9789B5DD19182BBF425" bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
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absent;
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<emphasis id="C86D0E78CDAEBACC123221F290EB31B6" bold="true" pageId="0" pageNumber="107">anthers</emphasis>
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0.5-1 mm long.
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</paragraph>
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<caption id="8A19713245085640A1BB3383BCB1A5ED" doi="10.3897/phytokeys.151.50538.figure10" httpUri="https://binary.pensoft.net/fig/424848" pageId="0" pageNumber="107" start="Figure 10" startId="F10">
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<paragraph id="068E664E7039630EB94ED76600FDC0AF" pageId="0" pageNumber="107">
|
|
<emphasis id="299EB73F94209944E4C78624BACD4D99" bold="true" pageId="0" pageNumber="107">Figure 10.</emphasis>
|
|
<taxonomicName id="EE11AB1C5EEE17C3572ADFCA5B334B60" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
|
|
<emphasis id="C3570D6FBE5D64EC94590501D0ABD30D" italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="F27AD0E6E164E07488ECE0A3DA33FD98" bold="true" pageId="0" pageNumber="107">A</emphasis>
|
|
spikelet, glumes in lateral view, floret in dorsal view, detached and raised above the glumes
|
|
<emphasis id="851FF4BB92C094B8E0CBDFA0CF4F50A0" bold="true" pageId="0" pageNumber="107">B</emphasis>
|
|
whole plant. Images of Cuta-Alarcon 376 (FMB).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="57F7B04DB63A5E257D56E52934F0C5EA" pageId="0" pageNumber="107" type="distribution">
|
|
<paragraph id="49D4307D6D456438BC04EB0D4023ADFD" pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
|
|
<paragraph id="00099E10F0FD9AE5F9C4C82ACC68D67C" pageId="0" pageNumber="107">Mexico, Guatemala, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia, Chile, Argentina. High-elevation open grasslands and forests, 2700-4900 m alt.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F97D94DDDE804299B3E48A16A957ED20" pageId="0" pageNumber="107" type="materials_examined">
|
|
<paragraph id="2073C745A2FFC7FCC211DC6923C7ACEB" pageId="0" pageNumber="107">Other specimens examined.</paragraph>
|
|
<paragraph id="B4E5386D92AECCC80E64E4AFA5B27CC8" pageId="0" pageNumber="107">
|
|
<emphasis id="116C9ADEB7E018951CAA114FAE632FD1" bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
|
|
. Sin loc., 1760-1808, J.C. Mutis 5521 (US1561906).
|
|
<emphasis id="3D22AC76F20B8D71E65818E8529D88F1" bold="true" pageId="0" pageNumber="107">
|
|
<normalizedToken id="9737EBEFE4243272C10F92400DC07986" originalValue="Boyacá">Boyaca</normalizedToken>
|
|
</emphasis>
|
|
: Munic. Arcabuco,
|
|
<normalizedToken id="FA2A7ED3CBBF263B543691ADDF7C7DFB" originalValue="Páramo">Paramo</normalizedToken>
|
|
El Valle,
|
|
<geoCoordinate id="7AD0D818882DABA2F79B6F2845889968" degrees="5.7445833" direction="north" orientation="latitude" precision="1" value="5.744583">5.7445833N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="B1C8B52972001080DD201C9C87923892" degrees="73.37118" direction="west" orientation="longitude" precision="1" value="-73.37118">73.37118W</geoCoordinate>
|
|
, 3744 m alt., 16 Nov. 2017, S.P. Sylvester 3081 (FMB, K, US); Munic. Arcabuco,
|
|
<normalizedToken id="E57EF120E1C5EA0737BC11812A157EBF" originalValue="Páramo">Paramo</normalizedToken>
|
|
El Valle, just below high point of ridge to the W of site 14,
|
|
<geoCoordinate id="0F9A5F8962444D405D4AD041A9F9738C" degrees="5.75436" direction="north" orientation="latitude" precision="1" value="5.75436">5.75436N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="423CFA2E333AEA02F8B21569018F5D8A" degrees="73.38722" direction="west" orientation="longitude" precision="1" value="-73.38722">73.38722W</geoCoordinate>
|
|
, 3747 m alt.,
|
|
<normalizedToken id="26A12A55E6C1DAD5E9AA0F3D8BADA0EB" originalValue="páramo">paramo</normalizedToken>
|
|
seco dominado por
|
|
<taxonomicName id="1CC9F3E05030A1E0F3F42D2B83C0EE5B" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia barclayana" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="barclayana">
|
|
<emphasis id="2590A2E9F4E46CAD04CA4DB7AB304635" italics="true" pageId="0" pageNumber="107">Espeletia barclayana</emphasis>
|
|
</taxonomicName>
|
|
[Cuatrec.], 16 Nov. 2017, S.P. Sylvester 3083 (K, US); Munic.
|
|
<normalizedToken id="B537F58ED73EB321CCF4265D447978C1" originalValue="Belén">Belen</normalizedToken>
|
|
,
|
|
<normalizedToken id="A5DA4B5CC8557799AC6C32A10525263E" originalValue="Páramo">Paramo</normalizedToken>
|
|
de La Rusia, border with Santander, Boqueron El Consuelo, unprotected private land,
|
|
<geoCoordinate id="3F0FA3B35F1FFBA96C910A8EA4B7B266" degrees="6.03598" direction="north" orientation="latitude" precision="1" value="6.03598">6.03598N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="2E99C84ED5676350CE458FAFFFDF2B76" degrees="72.57056" direction="west" orientation="longitude" precision="1" value="-72.57056">72.57056W</geoCoordinate>
|
|
, 3975 m alt.,
|
|
<normalizedToken id="CCD70D14AC0D1F2A1A303CA3EFE26835" originalValue="páramo">paramo</normalizedToken>
|
|
with limited grazing by horses and rodents with
|
|
<taxonomicName id="FE85FE22F28308D2B91509DE9D1A9F10" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia brachyaxiantha" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="brachyaxiantha">
|
|
<emphasis id="077EE00FDD914F592D1CDD199FB3D5BB" italics="true" pageId="0" pageNumber="107">Espeletia brachyaxiantha</emphasis>
|
|
</taxonomicName>
|
|
[S.
|
|
<normalizedToken id="10F4652DB8E2CB225ABA351A9BB1CC52" originalValue="Díaz">Diaz</normalizedToken>
|
|
] and
|
|
<taxonomicName id="47BB983F88A237975A1E4489708263D7" lsidName="E. annemariana" pageId="0" pageNumber="107" rank="species" species="annemariana">
|
|
<emphasis id="0738A4B088818F5A66806ED9B7B66659" italics="true" pageId="0" pageNumber="107">E. annemariana</emphasis>
|
|
</taxonomicName>
|
|
[Cuatrec.], steep slope on rock outcrop, 23 Nov. 2017, M. Vorontsova 2260 (FMB, K, SI, US); Munic.
|
|
<normalizedToken id="9E970B9F7A19815E7F1C82429794D39F" originalValue="Belén">Belen</normalizedToken>
|
|
,
|
|
<normalizedToken id="23D3E070E0BBDFB98E028A4121C57C66" originalValue="Páramo">Paramo</normalizedToken>
|
|
de La Rusia, near
|
|
<normalizedToken id="FE41A9CF076B2D461F7B348A85182383" originalValue="Páramo">Paramo</normalizedToken>
|
|
Le Consuelo, unprotected private land, somewhat disturbed
|
|
<normalizedToken id="CBA5466EBD6482F89BB0D389D522188C" originalValue="páramo">paramo</normalizedToken>
|
|
on top of the ridge,
|
|
<geoCoordinate id="5FA3F2CE77CB22D16D0591CDD8F28BEC" degrees="6.02416" direction="north" orientation="latitude" precision="1" value="6.02416">6.02416N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="4A46EEB476FC9D90395EFE6B55BF358D" degrees="72.57242" direction="west" orientation="longitude" precision="1" value="-72.57242">72.57242W</geoCoordinate>
|
|
, 3894 m alt., grazed by horses and rodents, with
|
|
<taxonomicName id="1CB58A97EB4ACA769782A4B49884E5BB" authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
|
<emphasis id="880500471607A673C972406FBA088EEC" italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
|
</taxonomicName>
|
|
, 22 Nov. 2017, M. Vorontsova 2231 (K, UPTC, US); Munic. Chiscas,
|
|
<normalizedToken id="1EBA5009574A82C5610B07288FD5915C" originalValue="Páramo">Paramo</normalizedToken>
|
|
de Chacaritas, asociado a rocas de 4 m de altura,
|
|
<geoCoordinate id="6BB468F221318A05F66EB907579997DF" degrees="6.62227" direction="north" orientation="latitude" precision="1" value="6.62227">6.62227N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="4BD8BFD516DAADC4451B63E0C4B33524" degrees="72.3904" direction="west" orientation="longitude" precision="5" value="-72.3904">72.3904W</geoCoordinate>
|
|
, 4192 m alt., 4 Mar. 2018, S.P. Sylvester 3115 (US); Munic. Chiscas,
|
|
<normalizedToken id="31B6FD38B6BD397B726EEFEE01A2E2E8" originalValue="Páramo">Paramo</normalizedToken>
|
|
de Chacaritas,
|
|
<normalizedToken id="50570D02F3B354B2F27679CF2751066A" originalValue="límites">limites</normalizedToken>
|
|
entre
|
|
<normalizedToken id="5B86A9CF9515CBAF9CFA9CDF2F287B4A" originalValue="páramo">paramo</normalizedToken>
|
|
y
|
|
<normalizedToken id="38F27CB4D03A7713A43B3199A90F8752" originalValue="superpáramo">superparamo</normalizedToken>
|
|
,
|
|
<geoCoordinate id="4E1A5E2744313AF5ADACEC9025F749C4" degrees="6.62865" direction="north" orientation="latitude" precision="1" value="6.62865">6.62865N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="06DBD09B707BCB7D2877944756575BF0" degrees="72.3944" direction="west" orientation="longitude" precision="5" value="-72.3944">72.3944W</geoCoordinate>
|
|
, 4064 m alt., 4 Mar. 2018, S.P. Sylvester 3104 (FMB, K, US); Munic. Chiscas,
|
|
<normalizedToken id="391E9D32D917D8AED3705D55EBB7B015" originalValue="Páramo">Paramo</normalizedToken>
|
|
El
|
|
<normalizedToken id="7EC8A4D5E75481DD9CA7AB1F33160994" originalValue="Peñon">Penon</normalizedToken>
|
|
, Chiscas,
|
|
<geoCoordinate id="AF78BFF2AE2083524AA39AACDEED1AB0" degrees="6.63012" direction="north" orientation="latitude" precision="1" value="6.63012">6.63012N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="FAE91D8C7902D8AA2819C9B5014BFC9B" degrees="72.40073" direction="west" orientation="longitude" precision="1" value="-72.40073">72.40073W</geoCoordinate>
|
|
, 4172 m alt., 5 Mar. 2018, S.P. Sylvester 3152 (UPTC); Munic. Chiscas,
|
|
<normalizedToken id="3AAFF16D36A51C31C4279C2182A39F84" originalValue="Páramo">Paramo</normalizedToken>
|
|
El
|
|
<normalizedToken id="1F130BB3F2FD1559AFCADA396420CFD3" originalValue="Peñon">Penon</normalizedToken>
|
|
, Chiscas,
|
|
<geoCoordinate id="FD521B47DAA8DE0A302DDFF65FDBFFEA" degrees="6.62876" direction="north" orientation="latitude" precision="1" value="6.62876">6.62876N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="1105FAD87E2DAF70C6A53E39AA47C6B1" degrees="72.40283" direction="west" orientation="longitude" precision="1" value="-72.40283">72.40283W</geoCoordinate>
|
|
, 4255 m alt.,
|
|
<normalizedToken id="C6743B96B3935494FD033E7331F5E637" originalValue="vegetación">vegetacion</normalizedToken>
|
|
de pajonal frailejonal,
|
|
<normalizedToken id="E9728E90E1C42D0CE4738C85BA5FA9F2" originalValue="páramo">paramo</normalizedToken>
|
|
<normalizedToken id="27D2F7B92B44F5F21E591B8CAEE7D81A" originalValue="húmedo">humedo</normalizedToken>
|
|
, 5 Mar. 2018, S.P. Sylvester 3153 (FMB, K, US); Munic. Mongua,
|
|
<normalizedToken id="D908BC6CE93D4E3D6F31A38F81F9B604" originalValue="Páramo">Paramo</normalizedToken>
|
|
de
|
|
<normalizedToken id="D0C513C438301753C04E0EA70A8800A6" originalValue="Ocetá">Oceta</normalizedToken>
|
|
, Valle de Laguna Negra,
|
|
<geoCoordinate id="A3EA5C04380CD1051F0C4C6DB21F0B60" degrees="5.69525" direction="north" orientation="latitude" precision="1" value="5.69525">5.69525N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="A034746A282CE320EC93C125D25712E1" degrees="72.79133" direction="west" orientation="longitude" precision="1" value="-72.79133">72.79133W</geoCoordinate>
|
|
, 3694 m alt., 29 Nov. 2017, L.E.
|
|
<normalizedToken id="4470BA4456D79F10AC527BB884157F13" originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
|
|
355 (FMB, K, UPTC, US); Munic. Mongui, Vereda Vallado, Sector La Pedrisca,
|
|
<normalizedToken id="068A3E5DC44746EBB3BB981E47856767" originalValue="Páramo">Paramo</normalizedToken>
|
|
de
|
|
<normalizedToken id="1D61F5C48014332DE1F214CEE79C96B7" originalValue="Ocetá">Oceta</normalizedToken>
|
|
,
|
|
<geoCoordinate id="B3DE86C946D20C8ACCE05644A24452EF" degrees="5.69969" direction="north" orientation="latitude" precision="1" value="5.69969">5.69969N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="126002DFDF5AE13DF0897C938333FC88" degrees="72.80891" direction="west" orientation="longitude" precision="1" value="-72.80891">72.80891W</geoCoordinate>
|
|
, 3751 m alt., 30 Nov. 2017, L.E.
|
|
<normalizedToken id="5290DEAEC1403275879942B5321455E7" originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
|
|
376 (FMB, US).
|
|
<emphasis id="32CCDD456BDB9758055E228EFA8C0B84" bold="true" pageId="0" pageNumber="107">Cauca</emphasis>
|
|
: Macizo Colombiano, Valle de las Papas, alrededores de Valencia, [
|
|
<geoCoordinate id="6C88F25835D1592768264F57B5CB7B7F" degrees="1.8831" direction="north" orientation="latitude" precision="5" value="1.8831">1.8831N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C279BA7956E19F92A935601D4AC145A1" degrees="76.6828" direction="west" orientation="longitude" precision="5" value="-76.6828">76.6828W</geoCoordinate>
|
|
], 2910 m alt., 11 Sep.-1 Oct. 1958, Idrobo, Pinto & Bischler 3682A (US2540528).
|
|
<emphasis id="9F16D4B81F821C5A142742FCCC02F013" bold="true" pageId="0" pageNumber="107">Cundinamarca</emphasis>
|
|
:
|
|
<normalizedToken id="B92B2B7273281B05D2FFD15BF4940682" originalValue="Páramo">Paramo</normalizedToken>
|
|
de Chisaca, [
|
|
<geoCoordinate id="6428C55DDBA2213D5235F26170A384F5" degrees="4.2747" direction="north" orientation="latitude" precision="5" value="4.2747">4.2747N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="ECE559083D3A4462F1B4E5C456B941FA" degrees="74.2006" direction="west" orientation="longitude" precision="5" value="-74.2006">74.2006W</geoCoordinate>
|
|
], 3750-3962 m alt., 5 Oct. 1966, T.R. Soderstrom 1312 (US3136708).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A9669196F69B2E3D2C96C7A0788CD286" pageId="0" pageNumber="107" type="notes">
|
|
<paragraph id="8264BE30FABBE92C94FE0E7D967E3CDC" pageId="0" pageNumber="107">Notes.</paragraph>
|
|
<paragraph id="747DCF38F018FF91246ECC4FAC435378" pageId="0" pageNumber="107">
|
|
Although in Bolivia (
|
|
<bibRefCitation id="C928553F1576964EA89BE47A002610E1" author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
|
|
) and austral South America (
|
|
<bibRefCitation id="D173292BF09AF9578BA615721558A936" author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
|
<normalizedToken id="50ADDA2AC9C750957BEE42254B089BDF" originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar 2012
|
|
</bibRefCitation>
|
|
)
|
|
<taxonomicName id="9AD2E96F6866B7428F0442C754B60897" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="478D292840A53CB5A82363AE786DD8E0" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
is mentioned to be notably rhizomatous, lateral tending rhizomes were not notable on specimens from the
|
|
<normalizedToken id="DAE6854748F7029F6C4EE1A2AB213643" originalValue="páramos">paramos</normalizedToken>
|
|
of
|
|
<normalizedToken id="B3978D3FAC8EF85E4B3EFFD4D6FD8EC5" originalValue="Boyacá">Boyaca</normalizedToken>
|
|
, which had short, ascending rhizomes/pseudostolons and formed small tussocks or were laxly to densely tufted. Type specimens and original material of
|
|
<taxonomicName id="08E66D3410504E17987F2B9BA6281A82" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="1A89B2F2D04E477BFCE6F7FFD2EE9482" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
and other species now considered synonyms of
|
|
<taxonomicName id="17CA6080DB5EBF2F9A659CFBF02B3984" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="C9086B183972AF80B753ED6BF21F0517" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
, i.e.
|
|
<taxonomicName id="94A9C7D8ABDF5796201FFD1844647A3D" authorityName="Kunth" authorityYear="1833" baseAuthorityName="J.Presl" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis glomerata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="glomerata">
|
|
<emphasis id="95C38C5AE17786D99A060FF42983478B" italics="true" pageId="0" pageNumber="107">Agrostis glomerata</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="9B0EA0DB2548C4ECBFD6D26CF2029338" lsidName="A. nana subsp. var. var. andicola" pageId="0" pageNumber="107" rank="variety" species="nana" subSpecies="var." variety="andicola">A. nana var. andicola</taxonomicName>
|
|
,
|
|
<taxonomicName id="734776EAB22377EBB2BAC7910EC6B6A2" lsidName="A. nana subsp. var. var. aristata" pageId="0" pageNumber="107" rank="variety" species="nana" subSpecies="var." variety="aristata">A. nana var. aristata</taxonomicName>
|
|
, and
|
|
<taxonomicName id="F6BEC16E884D6C5285F7F6083BFA510D" lsidName="A. virescens" pageId="0" pageNumber="107" rank="species" species="virescens">
|
|
<emphasis id="1B57BFC0CD7126FBF6F0F4BE42AB9ACF" italics="true" pageId="0" pageNumber="107">A. virescens</emphasis>
|
|
</taxonomicName>
|
|
, also had either short ascending rhizomes or lacked notable rhizomes, but with all having obvious extravaginal cataphyllous shoots.
|
|
</paragraph>
|
|
<paragraph id="3A030A2ED626600853898E5479E7B481" pageId="0" pageNumber="107">
|
|
Discrepancy was sometimes also found in the form of the leaf blades, with specimens encountered in
|
|
<normalizedToken id="F870D6BD397B6D0714AAAF545F2E241B" originalValue="Boyacá">Boyaca</normalizedToken>
|
|
, Colombia, and Venezuela sometimes having more rigid, convolute to folded, often recurved, basal leaf blades instead of the lax and filiform leaf blades more common to this species. These specimens did usually still have flat upper culm blades to help differentiate them from e.g.
|
|
<taxonomicName id="127785BB020D16E312C1F2FD52B49276" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="A1AE23943F72E2E5CC114E263AC3856C" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
(see notes on similar species below). While
|
|
<bibRefCitation id="7F9A9A08A52012A022852BBF349E06F2" author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize (1998)</bibRefCitation>
|
|
and
|
|
<bibRefCitation id="4987F6742C1E8B36F022A36385EEA5A9" author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
|
<normalizedToken id="989F9C01224693DF737F8AC462B941A9" originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar (2012)
|
|
</bibRefCitation>
|
|
state the blades in
|
|
<taxonomicName id="E0F8199B5DAEF83DD28251948B87BF87" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="432EF43AFBDAC9C0AA7692CC24C3AB60" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
to be flat, folded, or filiform and generally lax, the type material also has fairly firm blades which are narrow and folded to slightly convolute and sometimes cylindrical in outline.
|
|
<taxonomicName id="517F0C2A22097E7B42E65FCB821DBC15" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis nana" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="nana">
|
|
<emphasis id="156D9CBC7C363FDA34B0944A6481AEFD" italics="true" pageId="0" pageNumber="107">Agrostis nana</emphasis>
|
|
</taxonomicName>
|
|
, here considered a synonym of
|
|
<taxonomicName id="196F3D891DC29585FB2A5719B8926696" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="D64CEC989A5C6DEC3D14C504A6433B81" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
, also has more convolute, curved and rigid leaf blades.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BE9918D367921DBEAB8090FFFDBA8D1C" pageId="0" pageNumber="107" type="similar species">
|
|
<paragraph id="7ED39BCC8D9F6BFA2A0040B2FE1524D2" pageId="0" pageNumber="107">Similar species.</paragraph>
|
|
<paragraph id="A4B02A7D2CC5C81CB8C38B6C39469FB8" pageId="0" pageNumber="107">
|
|
See notes under
|
|
<taxonomicName id="146D4822C2358CC6122635A530363E59" lsidName="A. foliata" pageId="0" pageNumber="107" rank="species" species="foliata">
|
|
<emphasis id="6EC76150894F545158A0116C8E7E6667" italics="true" pageId="0" pageNumber="107">A. foliata</emphasis>
|
|
</taxonomicName>
|
|
for how to differentiate from that species.
|
|
<taxonomicName id="37FCB453CBE344A5757E4FD22AEAEE33" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis breviculmis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="breviculmis">
|
|
<emphasis id="9A41A79037163B862E598CCE11B04CB5" italics="true" pageId="0" pageNumber="107">Agrostis breviculmis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="8166C8B51896B89FF53D36D9C231FB37" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="EDD753D06B67650C4C8EF7DFD3A3FAC1" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
bear similarities in having mainly basal, narrow, usually convolute, rigid, leaf blades and densely congested spikelike panicles that are usually short (<5 cm long), and spikelets with an absent or reduced palea <
|
|
<normalizedToken id="F32A3B55D38FD94568F2DEF9C9B50B63" originalValue="¼">1/4</normalizedToken>
|
|
the length of the lemma.
|
|
<taxonomicName id="6F8ABD8516CA2815D63EA49439C8D393" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
|
|
<emphasis id="D396699B7C6A93AD714618CEFEA67F9C" italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
|
|
</taxonomicName>
|
|
principally differs from these by the leaf blades being usually laxer and folded or flat, at least in the upper culm. However, as noted above, specimens can be found with convolute basal blades that makes distinguishing these species more difficult. In these instances,
|
|
<taxonomicName id="A7D93056309B46016D7E2A6916195A3B" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="799120FCD34DBE72BBD031E1FB8E10D5" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
can be differentiated from
|
|
<taxonomicName id="4725397D25EE25706C07F355B678E0FB" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis id="6271A0B1EC1F0FE37476F4394DAD835C" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
by its larger spikelets 2-3(-3.6) mm long (vs. 1.5-2.1(-2.5 in Bolivia?;
|
|
<bibRefCitation id="A707154FEEB60A39952E9F9E97DC3954" author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
|
|
) mm long in
|
|
<taxonomicName id="5B2845BAE1CBCC3B7A81C070BF48FC20" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis id="03797E791F8BEAF80BE452FB38633796" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
), usually the presence of a well-developed geniculate and twisted awn inserted dorsally usually in the lower third of the lemma (lemmas very rarely muticous or with a short straight weak awn, <1 mm long, inserted in the upper
|
|
<normalizedToken id="83B57FC72F7ADACAE7971FB74DA2B268" originalValue="½">1/2</normalizedToken>
|
|
; see notes on
|
|
<taxonomicName id="DE3463B43E483325A6E8DCFE1088D91F" lsidName="A. glomerata" pageId="0" pageNumber="107" rank="species" species="glomerata">
|
|
<emphasis id="9EE2C15DFB44C3379E13A1A623CA4FDD" italics="true" pageId="0" pageNumber="107">A. glomerata</emphasis>
|
|
</taxonomicName>
|
|
below) (vs. lemmas muticous or with a short straight weak awn inserted above the middle in
|
|
<taxonomicName id="D6E67E4D9F076F5CA7B44657432E12DD" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis id="2B4A6D3B51169E4AF97AE558604EF6CB" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
), panicles generally wider, 1-15 mm wide (vs. 0.5-2(-6) mm wide in
|
|
<taxonomicName id="9E6E52A764F7288F1EA25A97CAC1E305" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis id="9F439F8EB14930484AD4C7D3F381229D" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
), and short prickle hairs on the glume keels (vs. coarse and shiny in
|
|
<taxonomicName id="619052DC3A285307629C6A6BB4364EA9" lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis id="3BDF2EA59B96B6BBC5A559A4082B55ED" italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
). While
|
|
<taxonomicName id="1FC06CA364284F601DED00617C7247A4" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="437D2D09AF45031DB62D222CBE8B97CE" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
has characters of often larger spikelets and dorsally inserted awn,
|
|
<taxonomicName id="9B2C6CFFEF6C6F46ED72952D4FCB82ED" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="819E69EB94524A4D7459FCA7459880E1" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
can be differentiated from
|
|
<taxonomicName id="B9D480E333AB5F2E80CA30F6328B83F9" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="23B46E8E1D8B390EB7B5423DBD44DD75" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
by its extravaginal cataphyllous shoots and distinct ascending rhizomes with rhizome internodes usually> 7 mm long (vs. purely intravaginal innovations, without cataphyllous shoots or distinct ascending rhizomes, rhizome internodes <2 mm long in
|
|
<taxonomicName id="D79952E13145F50CBB8B79A292705D0C" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="62F978AD23B7CD7ECC61C10461C0649D" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
), generally wider panicles, 1-15 mm wide (vs. 0.5-2(-6) mm wide in
|
|
<taxonomicName id="894E45D347A3E4135773AE1AC131A49C" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="7F3CF1EA99B6759028FB00127FB9044B" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
), and short prickle hairs on the glume keels (vs. coarse and shiny in
|
|
<taxonomicName id="50CE4C82C497D56D87990E9AD7FD1FA3" lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis id="8AB6E43559A1F28AE59B5080B19E4269" italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
). Specimens can sometimes have laxer inflorescences that could lead to confusion with
|
|
<taxonomicName id="C208DE54CCC365F21CA131F78400A0FE" lsidName="A. mertensii" pageId="0" pageNumber="107" rank="species" species="mertensii">
|
|
<emphasis id="86DAEB362AEE817793BE2B78E5E132F6" italics="true" pageId="0" pageNumber="107">A. mertensii</emphasis>
|
|
</taxonomicName>
|
|
, but this species does not form small tussocks.
|
|
</paragraph>
|
|
<paragraph id="522922B98192C8AAF73B5D3EAE3A831D" pageId="0" pageNumber="107">
|
|
<taxonomicName id="E4F7F7335AA19B523C092E62E3E54DD2" authorityName="Trin" authorityYear="1841" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis meyenii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="meyenii">
|
|
<emphasis id="991988F31CF688E6A9EAC7EB3BE29DC5" italics="true" pageId="0" pageNumber="107">Agrostis meyenii</emphasis>
|
|
</taxonomicName>
|
|
, a species known from drier high-elevation puna grassland and pampa of Argentina, Chile and Bolivia (
|
|
<bibRefCitation id="41579EF82492CFB02BC7485E5E4AF6A8" author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="D32CAC5C6392419D8F7B4FE036C59A2F" author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
|
<normalizedToken id="BD7D9C8DF91A5DF06D2DBC0B40ADD76A" originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar 2012
|
|
</bibRefCitation>
|
|
), is similar in its overall appearance, usually being tufted and with rhizomes, having similar ligules and filiform or flat leaf blades, and having a condensed spikelike panicle with spikelets of similar size.
|
|
<taxonomicName id="6B77949B76F81B55660832FF4CEBCA63" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
|
|
<emphasis id="1A669502782B2D53ADC796E699572216" italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
|
|
</taxonomicName>
|
|
can usually be differentiated from
|
|
<taxonomicName id="921BBEF83250A10EC346C85F3E5CA3A4" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
|
<emphasis id="ADA5ED051991B3BD61B8B3A2A1B3539B" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
|
</taxonomicName>
|
|
by the presence of an awn inserted in the lower third of the lemma, 2-3.5 mm long, twisted and bent and exerted from the glumes (vs. muticous or, if awn present, inserted in the middle or upper third of the lemma, to 1.2 mm long, straight or slightly flexuous in
|
|
<taxonomicName id="CBD6C1A6396D117F129E8867979749F6" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
|
<emphasis id="D49807F037C74BF35C4C26D148638AB9" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
|
</taxonomicName>
|
|
). However, specimens akin to
|
|
<taxonomicName id="CAB9C856DCBBFA74ACD26B74BA89EDED" lsidName="A. glomerata" pageId="0" pageNumber="107" rank="species" species="glomerata">
|
|
<emphasis id="4A04D2CB62DD7EE31F567E14F73A0E7F" italics="true" pageId="0" pageNumber="107">A. glomerata</emphasis>
|
|
</taxonomicName>
|
|
, a species described from Peru and here considered a synonym of
|
|
<taxonomicName id="D5CBD653C0F20CD8E7AB903684F154E3" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="3619006EC98E704765D720E9F4B89385" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
, can sometimes be found in Colombia, albeit not in the Cordillera Oriental, which have muticous lemmas or with a short straight awn inserted in the upper half of the lemma (e.g. Idrobo 3882a, Mutis 5521). These can be differentiated from
|
|
<taxonomicName id="6DE29FF5AD346D2EF5E801BF9E9F7718" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
|
<emphasis id="930707A42D21C1DCFFB5C7737404D8C9" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
|
</taxonomicName>
|
|
by the plants being generally taller, 20-60 cm tall, condensed panicles often> 10 cm long that are often interrupted and with the central inflorescence axis notably wider compared to the lateral branches, and the pedicels, panicle branches, and sometimes central inflorescence axis, being notably scabrous. The blades of these are variable and can be filiform to flat and to 5 mm wide. The characteristic of notably scabrous pedicels, panicle branches, and sometimes central inflorescence axis of
|
|
<taxonomicName id="C36C61041235C90EA8334A8C10B932C7" lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis id="428D7E596E4C9B9A6E691E42CDF3B987" italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
are considered to be key in differentiating these from
|
|
<taxonomicName id="7D4B3027CA55B19F8F10548AA5FE9443" lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
|
<emphasis id="548E51C4331358F3C2B5172463083414" italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
|
</taxonomicName>
|
|
, which are usually smooth or exceptionally lightly scaberulous.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |