495 lines
68 KiB
XML
495 lines
68 KiB
XML
<document id="5E875A96A24A41179E7C58E008EC82CB" ID-DOI="10.5281/zenodo.5311784" ID-ISSN="0374-1036" ID-Zenodo-Dep="5311784" ID-ZooBank="15CF0A9E-44F3-492B-88F7-A7922EF9F73A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630118666848" checkinUser="marcus" docAuthor="Angus, Robert B., Jia, Fenglong, Chen, Zhen-ning, Zhang, Ying, Vondráček, Dominik, Fikáček, Martin & Sw, London" docDate="2016" docId="377A879F8D71285FFE865B917070FC3F" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.109-148.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Helophorus (Lihelophorus) lamicola Zaitzev 1908" docType="treatment" docVersion="3" lastPageNumber="118" masterDocId="CB43FFE78D792856FFE05B4D7368FFAC" masterDocTitle="Taxonomy, larval morphology and cytogenetics of Lihelophorus, the Tibetan endemic subgenus of Helophorus (Coleoptera: Hydrophiloidea)" masterLastPageNumber="148" masterPageNumber="109" pageNumber="117" updateTime="1699158837459" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9E7306BAEE3B58E726CF406B6AC28808">Taxonomy, larval morphology and cytogenetics of Lihelophorus, the Tibetan endemic subgenus of Helophorus (Coleoptera: Hydrophiloidea)</mods:title>
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<mods:namePart id="71D72F86E7555EEF2FC7A3CDE924F3A7">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="21BE1CB4185171A3F2A5EDCE8CFAF9B1">Division of Life Sciences (Insects), Natural History Museum, Cromwell Road, London SW 7 5 BD, UK; e-mail: r. angus @ royalholloway. ac. uk</mods:affiliation>
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<mods:namePart id="A5C275A424D4FDC90A256E283C6BF18D">Jia, Fenglong</mods:namePart>
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<mods:affiliation id="5A7783AD480FE5F900E095EEDDCB3765">Museum of Biology, Sun Yat-sen University, Guangzhou, China; e-mail: lssjϐl @ mail. sysu. edu. cn</mods:affiliation>
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<mods:namePart id="60DF8997603A17F47383C27C7029EFA9">Chen, Zhen-ning</mods:namePart>
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<mods:affiliation id="08A0E14705787541644EF5879E02B2BC">Biology and Geography School, Qinghai Normal University, Wusi West Road 38 #, 810000, Xining, Qinghai Province, China; e-mail: 149470880 @ qq. com</mods:affiliation>
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<mods:namePart id="5D0FCCFD8B7FC7E486155D272C7FD7EA">Zhang, Ying</mods:namePart>
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<mods:affiliation id="511068AEBEC9C662D4FB070B14BAF7EA">Computer Network Information Center, Chinese Academy of Sciences, Beijing, 100190, China; e-mail: 344280438 @ qq. com</mods:affiliation>
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<mods:namePart id="9DC4457A2361CD04BBADE0FA267AE9D8">Vondráček, Dominik</mods:namePart>
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<mods:affiliation id="EC62F4AFD326AE500716921AB1AE1DF4">Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Prague 2, Czech Republic; e-mail: dominik. vondracek @ gmail. com & Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic; e-mail: mfikacek @ gmail. com</mods:affiliation>
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<mods:affiliation id="94F1E3B8522481EADAE58F0D4278FBB7">Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Prague 2, Czech Republic; e-mail: dominik. vondracek @ gmail. com & Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic; e-mail: mfikacek @ gmail. com</mods:affiliation>
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<taxonomicName id="78D34D0A8D71285EFE865B9170FFFF5F" ID-CoL="3KHKF" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[358,919,220,243]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="lamicola" subGenus="Lihelophorus">
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<emphasis id="8DA7EA9B8D71285EFE185B9171E4FF5F" bold="true" box="[504,652,220,243]" italics="true" pageId="8" pageNumber="117">Lihelophorus</emphasis>
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<emphasis id="8DA7EA9B8D71285EFD7C5B917193FF5F" bold="true" box="[668,763,220,243]" italics="true" pageId="8" pageNumber="117">lamicola</emphasis>
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<bibRefCitation id="DB424B788D71285EFCE25B9170FFFF5F" author="ZAITZEV F. A." box="[770,919,220,243]" pageId="8" pageNumber="117" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
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(
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<figureCitation id="27E82A0C8D71285EFE085BB77154FEA1" box="[488,572,250,269]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). a–b – H. yangae sp. nov. (a – ventral view of the head; b – detail of mentum and labial palps); c–d – maxillary galea and lacinia (c – H. yangae sp. nov.; d – H. ser Zaitzev, 1908); e–f – mandible (e – H. yangae sp. nov.; f – H. ser); g – labial palpus of H. ser; h–l – hind legs in dorsal view (h – H. yangae sp. nov.; i – H. ser; j – H. lamicola Zaitzev, 1908; k – detail of apex of tibia and base of tarsus of H. lamicola; l – detail of tarsomere 5 and claw of H. ser). Abbreviations: sh – swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="8" pageNumber="117">Figs 1j–k</figureCitation>
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;
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<figureCitation id="27E82A0C8D71285EFDA45BB771EAFEA1" box="[580,642,250,269]" captionStart="Fig" captionStartId="7.[113,144,1463,1482]" captionTargetBox="[133,1067,220,1453]" captionTargetId="figure-13@7.[133,1068,219,1454]" captionTargetPageId="7" captionText="Fig. 2. Adult morphology of Helophorus (Lihelophorus). a–f – details of left half of pronotum (a,c,e) and base of left elytron (b, d, f): a–b – H. yangae sp. nov.; c–d – H. ser Zaitzev, 1908; e–f – H. lamicola Zaitzev, 1908; arrow in (a) shows remnants of pronotal granulations, arrows in (b, d, f) show scutellary stria. g–h – prothoracic hypomeron and ‘suprapleural area’ (g – H. yangae sp. nov.; h – H. ser). i – detail of posterior margin of abdominal ventrite 5 of H. ser, showing the irregular denticulation. j – ventral view of the thorax of H. ser. k – dorsolateral portion of left elytron of H. lamicola. Abbreviations: ep – epipleuron; li – last elytral interval. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311790" httpUri="https://zenodo.org/record/5311790/files/figure.png" pageId="8" pageNumber="117">2e–f, k</figureCitation>
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;
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<figureCitation id="27E82A0C8D71285EFD6B5BB771DEFEA1" box="[651,694,250,269]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). a–c – H. lamicola Zaitzev, 1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Quinghai: Golo).d–g – H.ser Zaitzev, 1908 (d–e – Qinghai:Golo; f – Xizang, Nyainqentanglha Feng; g – Gansu: Xiahe env.). h–k – H. yangae sp. nov. (h – Xizang: 90 km W of Amdo; i – Xizang: 25 km W of Amdo; j – Qinghai: Golo; k – India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="8" pageNumber="117">3a–c</figureCitation>
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;
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<figureCitation id="27E82A0C8D71285EFD5F5BB77181FEA1" box="[703,745,250,269]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.a–c – H. lamicola Zaitzev,1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Qinghai: Golo). d–h – H. ser Zaitzev, 1908 (d – Qinghai: valley of Alag Hu lake (lectotype); e–f – Qinghai: Golo; g – Xizang, Nyainqentanglha Feng; h – Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="8" pageNumber="117">4a–c</figureCitation>
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<figureCitation id="27E82A0C8D71285EFD125BB7707EFEA1" box="[754,790,250,269]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="8" pageNumber="117">6f–i</figureCitation>
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)
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<emphasis id="8DA7EA9B8D71285EFF775A7A7394FEE6" box="[151,252,311,330]" italics="true" pageId="8" pageNumber="117">Helophorus</emphasis>
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(
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|
||
<bibRefCitation id="DB424B788D71285EFE9B5A7A729DFEE6" author="ANGUS R. B." box="[379,501,311,330]" pageId="8" pageNumber="117" pagination="185 - 206" refId="ref19760" refString="ANGUS R. B. 1995: Helophoridae: The Helophorus species of China, with notes on the species from neighbouring areas (Coleoptera). Pp 185 - 206. In: JACH M. A. & JI L. (eds): Water Beetles of China. Volume 1. Zoologisch- - Botanische Gesellschaft & Wiener Coleopterologenverein, Wien, 410 pp." type="book chapter" year="1995">
|
||
<collectingRegion id="7D17F86B8D71285EFE9B5A7A72DBFEE4" box="[379,435,311,330]" country="United Kingdom" name="Angus" pageId="8" pageNumber="117">ANGUS</collectingRegion>
|
||
(1995)
|
||
</bibRefCitation>
|
||
: 189
|
||
</treatmentCitation>
|
||
(redescription,
|
||
<typeStatus id="6068882B8D71285EFD435A7A7199FEE6" box="[675,753,311,330]" pageId="8" pageNumber="117" type="lectotype">lectotype</typeStatus>
|
||
designation).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F7C965028D71285EFF775A2C705FFCA8" pageId="8" pageNumber="117" type="materials_examined">
|
||
<paragraph id="BF6C36898D71285EFF775A2C70A6FED8" blockId="8.[151,1127,353,772]" box="[151,974,353,372]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF775A2C7278FED8" bold="true" box="[151,272,353,372]" pageId="8" pageNumber="117">
|
||
<typeStatus id="6068882B8D71285EFF775A2C73ACFED8" box="[151,196,353,372]" pageId="8" pageNumber="117">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="C7C476198D71285EFEF55A2C7222FED8" box="[277,330,353,372]" name="China" pageId="8" pageNumber="117">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="7D17F86B8D71285EFEB35A2C7283FED8" box="[339,491,353,372]" country="China" name="Qinghai" pageId="8" pageNumber="117">Qinghai Province</collectingRegion>
|
||
, Zhaling Hu (Lake) (= Djarin Nor),
|
||
<geoCoordinate id="DAE7504E8D71285EFCC05A2C701AFED8" box="[800,882,353,372]" degrees="34.911" direction="north" orientation="latitude" pageId="8" pageNumber="117" precision="55" value="34.911">34.911°N</geoCoordinate>
|
||
<geoCoordinate id="DAE7504E8D71285EFC975A2C70A2FED8" box="[887,970,353,372]" degrees="97.273" direction="east" orientation="longitude" pageId="8" pageNumber="117" precision="55" value="97.273">97.273°E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D71285EFF775A36705FFDD2" blockId="8.[151,1127,353,772]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF775A36721FFE22" bold="true" box="[151,375,379,398]" pageId="8" pageNumber="117">Type material examined</emphasis>
|
||
(
|
||
<specimenCount id="A9D5FD008D71285EFE655A3672AEFE22" box="[389,454,379,398]" pageId="8" pageNumber="117" type="generic">3 spec.</specimenCount>
|
||
).
|
||
<materialsCitation id="0FBB3CD48D71285EFE355A3670C2FE68" collectingDate="1900-06" collectionCode="ZIN" collectorName="Ekcp. & Eksp. Kozlova." county="Dol." location="Dzharin-nor." municipality="Khuan-khe" pageId="8" pageNumber="117" specimenCount="1" stateProvince="Angus" typeStatus="lectotype">
|
||
<typeStatus id="6068882B8D71285EFE355A367146FE20" box="[469,558,379,398]" pageId="8" pageNumber="117" type="lectotype">LECTOTYPE</typeStatus>
|
||
(designated by
|
||
<bibRefCitation id="DB424B788D71285EFD595A36703BFE22" author="ANGUS R. B." box="[697,851,379,398]" pageId="8" pageNumber="117" pagination="185 - 206" refId="ref19760" refString="ANGUS R. B. 1995: Helophoridae: The Helophorus species of China, with notes on the species from neighbouring areas (Coleoptera). Pp 185 - 206. In: JACH M. A. & JI L. (eds): Water Beetles of China. Volume 1. Zoologisch- - Botanische Gesellschaft & Wiener Coleopterologenverein, Wien, 410 pp." type="book chapter" year="1995">
|
||
<collectingRegion id="7D17F86B8D71285EFD595A36719AFE20" box="[697,754,379,398]" country="United Kingdom" name="Angus" pageId="8" pageNumber="117">ANGUS</collectingRegion>
|
||
1995: 189
|
||
</bibRefCitation>
|
||
): 1 J (
|
||
<collectionCode id="D9C2AE4C8D71285EFC735A3670D3FE22" box="[915,955,379,398]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="117" type="Museum">ZIN</collectionCode>
|
||
): ‘LECTO- / TYPE [rounded label with violet margin] // Дoл. оЗер верхн. Хуан- хе: [=
|
||
<collectingCounty id="560D4E058D71285EFD035ADB7061FE05" box="[739,777,406,425]" pageId="8" pageNumber="117">Dol.</collectingCounty>
|
||
ozer verkhn.
|
||
<collectingMunicipality id="5F08ACF38D71285EFC605ADB7089FE05" box="[896,993,406,425]" pageId="8" pageNumber="117">Khuan-khe</collectingMunicipality>
|
||
] / oЗ. Джарин- -нор. [= oz.
|
||
<location id="BA0C60528D71285EFEE25AFC721AFE68" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFEE25AFC721AFE68" box="[258,370,433,452]" county="Dol." municipality="Khuan-khe" name="Dzharin-nor." pageId="8" pageNumber="117" stateProvince="Angus">Dzharin-nor.</location>
|
||
] / Кон. [=
|
||
<location id="BA0C60528D71285EFE2E5AFC7293FE68" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFE2E5AFC7293FE68" box="[462,507,433,452]" county="Dol." municipality="Khuan-khe" name="Kon." pageId="8" pageNumber="117" stateProvince="Angus">Kon.</location>
|
||
]
|
||
<date id="CB6D10498D71285EFDE45AFC713BFE68" box="[516,595,433,452]" pageId="8" pageNumber="117" value="1900-06">
|
||
<collectingDate id="DB29E9A18D71285EFDE45AFC713BFE68" box="[516,595,433,452]" pageId="8" pageNumber="117" value="1900-06">VI. 1900</collectingDate>
|
||
</date>
|
||
. /
|
||
<collectorName id="1226535F8D71285EFD875AFC71F2FE68" box="[615,666,433,452]" pageId="8" pageNumber="117">Эkcп.</collectorName>
|
||
КoЗлoвa. [=
|
||
<collectorName id="1226535F8D71285EFCEE5AFC70FDFE68" box="[782,917,433,452]" pageId="8" pageNumber="117">Eksp. Kozlova.</collectorName>
|
||
] //
|
||
</materialsCitation>
|
||
<materialsCitation id="0FBB3CD48D71285EFC4F5AFC7185FE72" collectingDate="1970-03" collectorName="Zaitz. & R. G. BEUTEL & Zaitzev'" location="Angus" pageId="8" pageNumber="117" specimenCount="1" stateProvince="Angus" typeStatus="lectotype">
|
||
<typeStatus id="6068882B8D71285EFC4F5AFC7741FE68" box="[943,1065,433,452]" pageId="8" pageNumber="117" type="lectotype">LECTOTYPE</typeStatus>
|
||
<date id="CB6D10498D71285EFBCF5AFC73AAFE72" pageId="8" pageNumber="117" value="1970-03">
|
||
<collectingDate id="DB29E9A18D71285EFBCF5AFC73AAFE72" pageId="8" pageNumber="117" value="1970-03">March 1970</collectingDate>
|
||
</date>
|
||
/
|
||
<taxonomicName id="78D34D0A8D71285EFF325A8672ECFE72" authorityName="Zaitzev" authorityYear="1908" box="[210,388,459,478]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="lamicola">Helophorus lamicola</taxonomicName>
|
||
/
|
||
<collectorName id="1226535F8D71285EFE755A8672AFFE72" box="[405,455,459,478]" pageId="8" pageNumber="117">Zaitz.</collectorName>
|
||
/
|
||
<collectorName id="1226535F8D71285EFE365A86716BFE72" box="[470,515,459,478]" name="R. G. BEUTEL" pageId="8" pageNumber="117">R. B.</collectorName>
|
||
<collectingRegion id="7D17F86B8D71285EFDE65A867156FE72" box="[518,574,459,478]" country="United Kingdom" name="Angus" pageId="8" pageNumber="117">Angus</collectingRegion>
|
||
det. // coll.
|
||
<collectorName id="1226535F8D71285EFD435A867185FE72" box="[675,749,459,478]" pageId="8" pageNumber="117">Zaitzev’</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="0FBB3CD48D71285EFD155A86704FFDB8" collectingDate="1900-06" collectingDateMax="1970-03" collectingDateMin="1900-06" collectionCode="ZIN" collectorName="Ekcp." pageId="8" pageNumber="117" specimenCount="1" specimenCount-female="1" stateProvince="Angus" typeStatus="paralectotype">
|
||
<typeStatus id="6068882B8D71285EFD155A867016FE70" box="[757,894,459,478]" pageId="8" pageNumber="117" type="paralectotype">PARALECTOTYPES</typeStatus>
|
||
: 1 damaged specimen with dissected mouthparts,
|
||
<specimenCount id="A9D5FD008D71285EFEB55AAB721DFE54" box="[341,373,479,505]" pageId="8" pageNumber="117" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="D9C2AE4C8D71285EFE605AAB72C0FE55" box="[384,424,486,505]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="117" type="Museum">ZIN</collectionCode>
|
||
): ‘PARA- / LECTO- / TYPE [rounded label with blue margin] // Дoл. оЗер верхн. Хуан- хе: / oЗ. Джарин- нор. / Кон.
|
||
<date id="CB6D10498D71285EFE23594C7179FDB8" box="[451,529,513,532]" pageId="8" pageNumber="117" value="1900-06">
|
||
<collectingDate id="DB29E9A18D71285EFE23594C7179FDB8" box="[451,529,513,532]" pageId="8" pageNumber="117" value="1900-06">VI. 1900</collectingDate>
|
||
</date>
|
||
. /
|
||
<collectorName id="1226535F8D71285EFDC3594C713EFDB8" box="[547,598,513,532]" pageId="8" pageNumber="117">Эkcп.</collectorName>
|
||
КoЗлoвa. //
|
||
<date id="CB6D10498D71285EFD5F594C704FFDB8" box="[703,807,513,532]" pageId="8" pageNumber="117" value="1970-03">
|
||
<collectingDate id="DB29E9A18D71285EFD5F594C704FFDB8" box="[703,807,513,532]" pageId="8" pageNumber="117" value="1970-03">March 1970</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
/
|
||
<materialsCitation id="0FBB3CD48D71285EFCD6594C72D2FD82" collectorName="R. G. BEUTEL & Zaitzev'" location="Angus" pageId="8" pageNumber="117" specimenCount="1" stateProvince="Angus" typeStatus="paralectotype">
|
||
<typeStatus id="6068882B8D71285EFCD6594C70C5FDB8" box="[822,941,513,532]" pageId="8" pageNumber="117" type="paralectotype">Paralectotype</typeStatus>
|
||
. /
|
||
<taxonomicName id="78D34D0A8D71285EFC5F594C770EFDB8" authority="Zaitzev" authorityName="Zaitzev" authorityYear="1908" box="[959,1126,513,532]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="lamicola">H. lamicola Zaitzev</taxonomicName>
|
||
/
|
||
<collectorName id="1226535F8D71285EFF42595673B8FD82" box="[162,208,539,558]" name="R. G. BEUTEL" pageId="8" pageNumber="117">R. B.</collectorName>
|
||
<collectingRegion id="7D17F86B8D71285EFF3359567263FD82" box="[211,267,539,558]" country="United Kingdom" name="Angus" pageId="8" pageNumber="117">Angus</collectingRegion>
|
||
det. // coll.
|
||
<collectorName id="1226535F8D71285EFE90595672D2FD82" box="[368,442,539,558]" pageId="8" pageNumber="117">Zaitzev’</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0FBB3CD48D71285EFE225956705AFDD2" collectionCode="NHRS" county="Vatten" determinerName="Zaitzev" elevation="5073" latitude="36.266666" location="Hedin" longLatPrecision="1188" longitude="88.35" municipality="Juni" pageId="8" pageNumber="117" specimenCount="1" stateProvince="Angus" typeStatus="paralectotype">
|
||
1 J (
|
||
<collectionCode id="D9C2AE4C8D71285EFE0D59567143FD82" box="[493,555,539,558]" country="Sweden" name="Swedish Museum of Natural History, Entomology Collections" pageId="8" pageNumber="117">NHRS</collectionCode>
|
||
): ‘
|
||
<collectingCounty id="560D4E058D71285EFDDC5956711FFD82" box="[572,631,539,558]" pageId="8" pageNumber="117">Vatten</collectingCounty>
|
||
djur [= water animal] / 25
|
||
<collectingMunicipality id="5F08ACF38D71285EFCBD595670E9FD82" box="[861,897,539,558]" pageId="8" pageNumber="117">Juni</collectingMunicipality>
|
||
[19]01 / L. XXIII [= camp XXIII] /
|
||
<location id="BA0C60528D71285EFF02597B727EFDE5" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFF02597B727EFDE5" box="[226,278,566,585]" county="Vatten" latitude="36.266666" longLatPrecision="1188" longitude="88.35" municipality="Juni" name="Hedin" pageId="8" pageNumber="117" stateProvince="Angus">Hedin</location>
|
||
//
|
||
<location id="BA0C60528D71285EFECC597B723AFDE5" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFECC597B723AFDE5" box="[300,338,566,585]" county="Vatten" latitude="36.266666" longLatPrecision="1188" longitude="88.35" municipality="Juni" name="Hel." pageId="8" pageNumber="117" stateProvince="Angus">Hel.</location>
|
||
(
|
||
<taxonomicName id="78D34D0A8D71285EFEBB597B72B8FDE5" authorityName="Zaitzev" authorityYear="1908" box="[347,464,566,585]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">Lihelophorus</taxonomicName>
|
||
) / lamicola m. / III. 08
|
||
<determinerName id="4AB0DA4D8D71285EFD74597B71BCFDE5" box="[660,724,566,585]" pageId="8" pageNumber="117">Zaitzev</determinerName>
|
||
det. / 8248 // NHRS-JKLB /000021180 // 3817 / E91”.
|
||
<location id="BA0C60528D71285EFF3A591C7266FDC8" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFF3A591C7266FDC8" box="[218,270,593,612]" county="Vatten" latitude="36.266666" longLatPrecision="1188" longitude="88.35" municipality="Juni" name="Based" pageId="8" pageNumber="117" stateProvince="Angus">Based</location>
|
||
on
|
||
<bibRefCitation id="DB424B788D71285EFECF591C72BAFDC8" author="ORCHYMONT A." box="[303,466,593,612]" pageId="8" pageNumber="117" pagination="1 - 16" refId="ref20836" refString="ORCHYMONT A. d' 1943: Les Palpicornia du Tibet (Coleoptera). Bulletin du Musee Royal d'Histoire Naturelle de Belgique 19 (57): 1 - 16." type="journal article" year="1943">ORCHYMONT (1943)</bibRefCitation>
|
||
who studied
|
||
<location id="BA0C60528D71285EFDA6591C71D2FDC8" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFDA6591C71D2FDC8" box="[582,698,593,612]" county="Vatten" latitude="36.266666" longLatPrecision="1188" longitude="88.35" municipality="Juni" name="Sven Hedin's" pageId="8" pageNumber="117" stateProvince="Angus">Sven Hedinʼs</location>
|
||
journal, camp 23 was situated south of
|
||
<location id="BA0C60528D71285EFBF0591C770EFDC8" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFBF0591C770EFDC8" box="[1040,1126,593,612]" county="Vatten" latitude="36.266666" longLatPrecision="1188" longitude="88.35" municipality="Juni" name="Arka-tagh" pageId="8" pageNumber="117" stateProvince="Angus">Arka-tagh</location>
|
||
at altitude of
|
||
<elevation id="34FED1BA8D71285EFEE85926721FFDD2" box="[264,375,619,638]" metricMagnitude="3" metricUnit="m" metricValue="5.073" pageId="8" pageNumber="117" unit="m" value="5073.0">
|
||
<quantity id="782B9B6C8D71285EFEE859267221FDD2" box="[264,329,619,638]" metricMagnitude="3" metricUnit="m" metricValue="5.073" pageId="8" pageNumber="117" unit="m" value="5073.0">5073 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, at approximate GPS coordinates
|
||
<geoCoordinate id="DAE7504E8D71285EFD7A5926718AFDD2" box="[666,738,619,638]" degrees="36" direction="north" minutes="16" orientation="latitude" pageId="8" pageNumber="117" precision="925" value="36.266666">36°16ʹN</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DAE7504E8D71285EFD0B5926705AFDD2" box="[747,818,619,638]" degrees="88" direction="east" minutes="21" orientation="longitude" pageId="8" pageNumber="117" precision="925" value="88.35">88°21′E</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D71285EFF7759CB705FFCA8" blockId="8.[151,1127,353,772]" pageId="8" pageNumber="117">
|
||
<materialsCitation id="0FBB3CD48D71285EFF7759CB7155FD18" collectingDate="1970-03" collectorName="Semenov" country="China" location="Angus" pageId="8" pageNumber="117" specimenCount="10" specimenCount-female="1" stateProvince="Qinghai">
|
||
<emphasis id="8DA7EA9B8D71285EFF7759CB72C1FD35" bold="true" box="[151,425,646,665]" pageId="8" pageNumber="117">Additional material examined</emphasis>
|
||
(
|
||
<specimenCount id="A9D5FD008D71285EFE5659CB729EFD35" box="[438,502,646,665]" pageId="8" pageNumber="117" type="generic">9 spec.</specimenCount>
|
||
).
|
||
<emphasis id="8DA7EA9B8D71285EFDE459CB71C1FD35" bold="true" box="[516,681,646,665]" pageId="8" pageNumber="117">
|
||
<collectingCountry id="C7C476198D71285EFDE459CB7126FD35" box="[516,590,646,665]" name="China" pageId="8" pageNumber="117">CHINA</collectingCountry>
|
||
:
|
||
<collectingRegion id="7D17F86B8D71285EFDB959CB71C9FD3B" box="[601,673,646,665]" country="China" name="Qinghai" pageId="8" pageNumber="117">QINGHAI</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="A9D5FD008D71285EFD4F59CB71A7FD34" box="[687,719,639,665]" pageId="8" pageNumber="117" type="female">1 ♀</specimenCount>
|
||
(BMNH): same label data as the lectotype, but from
|
||
<collectorName id="1226535F8D71285EFF2659EC727DFD18" box="[198,277,673,692]" pageId="8" pageNumber="117">Semenov</collectorName>
|
||
collection and without type labels
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0FBB3CD48D71285EFDA659EC7034FD62" collectingDate="2013-06-07" collectorName="R. B. Angus & F. L. Jia & Zhang" country="China" county="Golo" elevation="4141" latitude="35.331112" location="Zuimatan" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" pageId="8" pageNumber="117" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai">
|
||
3 JJ
|
||
<specimenCount id="A9D5FD008D71285EFD9A59EC71F2FD1F" box="[634,666,666,692]" pageId="8" pageNumber="117" type="female">1 ♀</specimenCount>
|
||
(BMNH):
|
||
<collectingCounty id="560D4E058D71285EFD1859EC704DFD18" box="[760,805,673,692]" pageId="8" pageNumber="117">Golo</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="5F08ACF38D71285EFCCE59EC70E1FD18" box="[814,905,673,692]" pageId="8" pageNumber="117">Huamuxia</collectingMunicipality>
|
||
,
|
||
<location id="BA0C60528D71285EFC7159EC708DFD18" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFC7159EC708DFD18" box="[913,997,673,692]" country="China" county="Golo" latitude="35.331112" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" name="Zuimatan" pageId="8" pageNumber="117" stateProvince="Qinghai">Zuimatan</location>
|
||
, roadside pool,
|
||
<geoCoordinate id="DAE7504E8D71285EFF7759F67396FD62" box="[151,254,699,718]" degrees="35" direction="north" minutes="19" orientation="latitude" pageId="8" pageNumber="117" precision="15" seconds="52" value="35.331112">35°19′52″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DAE7504E8D71285EFEE759F67204FD62" box="[263,364,699,718]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="8" pageNumber="117" precision="15" seconds="57" value="99.065834">99°03′57″E</geoCoordinate>
|
||
,
|
||
<quantity id="782B9B6C8D71285EFE9559F672D0FD62" box="[373,440,699,718]" metricMagnitude="3" metricUnit="m" metricValue="4.141" pageId="8" pageNumber="117" unit="m" value="4141.0">
|
||
<elevation id="34FED1BA8D71285EFE9559F672D0FD62" box="[373,440,699,718]" metricMagnitude="3" metricUnit="m" metricValue="4.141" pageId="8" pageNumber="117" unit="m" value="4141.0">4141 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="CB6D10498D71285EFE2059F6717BFD62" box="[448,531,699,718]" pageId="8" pageNumber="117" value="2013-06-07">
|
||
<collectingDate id="DB29E9A18D71285EFE2059F6717BFD62" box="[448,531,699,718]" pageId="8" pageNumber="117" value="2013-06-07">7.vi.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="1226535F8D71285EFDFC59F671EEFD62" box="[540,646,699,718]" pageId="8" pageNumber="117">R. B. Angus</collectorName>
|
||
,
|
||
<collectorName id="1226535F8D71285EFD6F59F671BBFD62" box="[655,723,699,718]" pageId="8" pageNumber="117">F. L. Jia</collectorName>
|
||
& Y.
|
||
<collectorName id="1226535F8D71285EFCE559F67054FD62" box="[773,828,699,718]" pageId="8" pageNumber="117">Zhang</collectorName>
|
||
lgt
|
||
</materialsCitation>
|
||
.; 1 J (
|
||
<collectionCode id="D9C2AE4C8D71285EFC7359F670A6FD62" box="[915,974,699,718]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="8" pageNumber="117" type="Herbarium">SYSU</collectionCode>
|
||
): same label data;
|
||
<materialsCitation id="0FBB3CD48D71285EFF77599B7193FD45" box="[151,763,726,745]" collectionCode="NMPC" country="China" pageId="8" pageNumber="117" specimenCount="2" stateProvince="Qinghai">
|
||
<specimenCount id="A9D5FD008D71285EFF77599B73BDFD45" box="[151,213,726,745]" pageId="8" pageNumber="117" type="generic">2 spec.</specimenCount>
|
||
(
|
||
<collectionCode id="D9C2AE4C8D71285EFF3F599B7249FD45" box="[223,289,726,745]" country="Czech Republic" name="National Museum Prague" pageId="8" pageNumber="117">NMPC</collectionCode>
|
||
): same label data [DNA vouchers MF661 and MF662]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0FBB3CD48D71285EFCE4599B705BFCA8" collectingDate="2015-07-22" collectionCode="NMPC" collectorName="J. Schmidt" country="China" elevation="4500" latitude="35.36472" location="Tuotuohe" longLatPrecision="19" longitude="93.44111" pageId="8" pageNumber="117" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai">
|
||
<specimenCount id="A9D5FD008D71285EFCE4599B704EFD44" box="[772,806,719,745]" pageId="8" pageNumber="117" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="D9C2AE4C8D71285EFCD0599B701AFD45" box="[816,882,726,745]" country="Czech Republic" name="National Museum Prague" pageId="8" pageNumber="117">NMPC</collectionCode>
|
||
): road to
|
||
<location id="BA0C60528D71285EFC23599B777EFD45" LSID="urn:lsid:plazi:treatment:377A879F8D71285FFE865B917070FC3F:BA0C60528D71285EFC23599B777EFD45" box="[963,1046,726,745]" country="China" latitude="35.36472" longLatPrecision="19" longitude="93.44111" name="Tuotuohe" pageId="8" pageNumber="117" stateProvince="Qinghai">Tuotuohe</location>
|
||
,
|
||
<quantity id="782B9B6C8D71285EFBC0599B770BFD45" box="[1056,1123,726,745]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="117" unit="m" value="4500.0">
|
||
<elevation id="34FED1BA8D71285EFBC0599B770BFD45" box="[1056,1123,726,745]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="117" unit="m" value="4500.0">4500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="DAE7504E8D71285EFF7759BC7394FCA8" box="[151,252,753,772]" degrees="35" direction="north" minutes="21" orientation="latitude" pageId="8" pageNumber="117" precision="15" seconds="53" value="35.36472">35°21′53″N</geoCoordinate>
|
||
<geoCoordinate id="DAE7504E8D71285EFEE259BC720FFCA8" box="[258,359,753,772]" degrees="93" direction="east" minutes="26" orientation="longitude" pageId="8" pageNumber="117" precision="15" seconds="28" value="93.44111">93°26′28″E</geoCoordinate>
|
||
,
|
||
<date id="CB6D10498D71285EFE9059BC72BAFCA8" box="[368,466,753,772]" pageId="8" pageNumber="117" value="2015-07-22">
|
||
<collectingDate id="DB29E9A18D71285EFE9059BC72BAFCA8" box="[368,466,753,772]" pageId="8" pageNumber="117" value="2015-07-22">22.vii.2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="1226535F8D71285EFE3C59BC715EFCA8" box="[476,566,753,772]" pageId="8" pageNumber="117">J. Schmidt</collectorName>
|
||
lgt. [DNA voucher MF1247]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F7C965028D71285EFF77585170D8FBBF" pageId="8" pageNumber="117" type="diagnosis">
|
||
<paragraph id="BF6C36898D71285EFF77585170D8FBBF" blockId="8.[151,1127,796,1619]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF77585172E1FC9F" bold="true" box="[151,393,796,819]" pageId="8" pageNumber="117">Differential diagnosis.</emphasis>
|
||
Differs from other
|
||
<taxonomicName id="78D34D0A8D71285EFDAD585171B0FC9F" authorityName="Zaitzev" authorityYear="1908" box="[589,728,796,819]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
|
||
<emphasis id="8DA7EA9B8D71285EFDAD585171B0FC9F" box="[589,728,796,819]" italics="true" pageId="8" pageNumber="117">Lihelophorus</emphasis>
|
||
</taxonomicName>
|
||
by narrow pronotum (maximum width clearly less than length of mesotarsus + claw) with rather straight sides and almost entirely effaced pronotal grooves (
|
||
<figureCitation id="27E82A0C8D71285EFE465811729DFCDF" box="[422,501,860,883]" captionStart="Fig" captionStartId="7.[113,144,1463,1482]" captionTargetBox="[133,1067,220,1453]" captionTargetId="figure-13@7.[133,1068,219,1454]" captionTargetPageId="7" captionText="Fig. 2. Adult morphology of Helophorus (Lihelophorus). a–f – details of left half of pronotum (a,c,e) and base of left elytron (b, d, f): a–b – H. yangae sp. nov.; c–d – H. ser Zaitzev, 1908; e–f – H. lamicola Zaitzev, 1908; arrow in (a) shows remnants of pronotal granulations, arrows in (b, d, f) show scutellary stria. g–h – prothoracic hypomeron and ‘suprapleural area’ (g – H. yangae sp. nov.; h – H. ser). i – detail of posterior margin of abdominal ventrite 5 of H. ser, showing the irregular denticulation. j – ventral view of the thorax of H. ser. k – dorsolateral portion of left elytron of H. lamicola. Abbreviations: ep – epipleuron; li – last elytral interval. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311790" httpUri="https://zenodo.org/record/5311790/files/figure.png" pageId="8" pageNumber="117">Figs 2e</figureCitation>
|
||
,
|
||
<figureCitation id="27E82A0C8D71285EFE1E5811715BFCDF" box="[510,563,860,883]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). a–c – H. lamicola Zaitzev, 1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Quinghai: Golo).d–g – H.ser Zaitzev, 1908 (d–e – Qinghai:Golo; f – Xizang, Nyainqentanglha Feng; g – Gansu: Xiahe env.). h–k – H. yangae sp. nov. (h – Xizang: 90 km W of Amdo; i – Xizang: 25 km W of Amdo; j – Qinghai: Golo; k – India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="8" pageNumber="117">3a–c</figureCitation>
|
||
,
|
||
<figureCitation id="27E82A0C8D71285EFDDC5811711AFCDF" box="[572,626,860,883]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.a–c – H. lamicola Zaitzev,1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Qinghai: Golo). d–h – H. ser Zaitzev, 1908 (d – Qinghai: valley of Alag Hu lake (lectotype); e–f – Qinghai: Golo; g – Xizang, Nyainqentanglha Feng; h – Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="8" pageNumber="117">4a–c</figureCitation>
|
||
). In contrast to other
|
||
<taxonomicName id="78D34D0A8D71285EFCAA581170BFFCDF" authorityName="Zaitzev" authorityYear="1908" box="[842,983,860,883]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
|
||
<emphasis id="8DA7EA9B8D71285EFCAA581170BFFCDF" box="[842,983,860,883]" italics="true" pageId="8" pageNumber="117">Lihelophorus</emphasis>
|
||
</taxonomicName>
|
||
, elytral series are often not impressed basally, punctures of elytral series are small and interval punctures very small; because of that, the scutellary stria is sometimes rather indistinct. Meso- and metatarsal claws are long (as in
|
||
<taxonomicName id="78D34D0A8D71285EFE0C58F17145FC7F" authorityName="Zaitzev" authorityYear="1908" box="[492,557,956,979]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="ser">
|
||
<emphasis id="8DA7EA9B8D71285EFE0C58F17145FC7F" box="[492,557,956,979]" italics="true" pageId="8" pageNumber="117">H. ser</emphasis>
|
||
</taxonomicName>
|
||
, but unlike
|
||
<taxonomicName id="78D34D0A8D71285EFD4C58F17071FC7F" authority="Angus & Jia & Chen & Zhang & Vondráček & Fikáček & Sw, 2016" authorityName="Angus & Jia & Chen & Zhang & Vondráček & Fikáček & Sw" authorityYear="2016" box="[684,793,956,979]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov.">
|
||
<emphasis id="8DA7EA9B8D71285EFD4C58F17071FC7F" box="[684,793,956,979]" italics="true" pageId="8" pageNumber="117">H. yangae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="969457E08D71285EFCFF58F1701EFC7F" box="[799,886,956,979]" pageId="8" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). The aedeagus is very similar to that of
|
||
<taxonomicName id="78D34D0A8D71285EFEAA589172E3FC5F" authorityName="Zaitzev" authorityYear="1908" box="[330,395,988,1011]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="ser">
|
||
<emphasis id="8DA7EA9B8D71285EFEAA589172E3FC5F" box="[330,395,988,1011]" italics="true" pageId="8" pageNumber="117">H. ser</emphasis>
|
||
</taxonomicName>
|
||
(based on pointed parameral apices) but the median lobe is wider and the apical portion shorter (
|
||
<figureCitation id="27E82A0C8D71285EFE5158B1717EFBBF" box="[433,534,1020,1043]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="8" pageNumber="117">Figs 6f–i</figureCitation>
|
||
). For detailed comparison see
|
||
<tableCitation id="F25103328D71285EFCBA58B170C4FBBF" box="[858,940,1020,1043]" captionStart="Table 2" captionStartId="16.[153,200,220,239]" captionText="Table 2. Comparison of adult morphology of Lihelophorus species." pageId="8" pageNumber="117">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F7C965028D71285FFF775F51775AFD5F" lastPageId="9" lastPageNumber="118" pageId="8" pageNumber="117" type="description">
|
||
<paragraph id="BF6C36898D71285EFF775F5170D7FB9F" blockId="8.[151,1127,796,1619]" box="[151,959,1052,1075]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF775F517255FB9F" bold="true" box="[151,317,1052,1075]" pageId="8" pageNumber="117">Redescription.</emphasis>
|
||
Body length:
|
||
<quantity id="782B9B6C8D71285EFE345F51713CFB9F" box="[468,596,1052,1075]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.5" metricValueMin="5.1" pageId="8" pageNumber="117" unit="mm" value="5.3" valueMax="5.5" valueMin="5.1">5.1–5.5 mm</quantity>
|
||
in males,
|
||
<quantity id="782B9B6C8D71285EFD225F51702AFB9F" box="[706,834,1052,1075]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.1" metricValueMin="5.2" pageId="8" pageNumber="117" unit="mm" value="5.65" valueMax="6.1" valueMin="5.2">5.2–6.1 mm</quantity>
|
||
in females.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D71285EFF555F71729BFB5F" blockId="8.[151,1127,796,1619]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF555F71739FFBFF" box="[181,247,1084,1107]" italics="true" pageId="8" pageNumber="117">Head.</emphasis>
|
||
Dull brownish bronze with green reflections, closely and irregularly punctate, the punctures not round, giving a generally rough effect. Y-groove shallow, its stem rather indistinct, wider than the lateral arms, widened anteriorly. Maxillary palpi pale yellow, elongate, apical segment asymmetrical. Antennae yellow, 9-segmented. Maxillary galea in shape of simple pubescent lobe, without strong spines (as in
|
||
<figureCitation id="27E82A0C8D71285EFD335FF1704CFB7F" box="[723,804,1212,1235]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). a–b – H. yangae sp. nov. (a – ventral view of the head; b – detail of mentum and labial palps); c–d – maxillary galea and lacinia (c – H. yangae sp. nov.; d – H. ser Zaitzev, 1908); e–f – mandible (e – H. yangae sp. nov.; f – H. ser); g – labial palpus of H. ser; h–l – hind legs in dorsal view (h – H. yangae sp. nov.; i – H. ser; j – H. lamicola Zaitzev, 1908; k – detail of apex of tibia and base of tarsus of H. lamicola; l – detail of tarsomere 5 and claw of H. ser). Abbreviations: sh – swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="8" pageNumber="117">Fig. 1d</figureCitation>
|
||
). Mandible rather short, with simple blunt apex (as in
|
||
<figureCitation id="27E82A0C8D71285EFE7D5F91728EFB5F" box="[413,486,1244,1267]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). a–b – H. yangae sp. nov. (a – ventral view of the head; b – detail of mentum and labial palps); c–d – maxillary galea and lacinia (c – H. yangae sp. nov.; d – H. ser Zaitzev, 1908); e–f – mandible (e – H. yangae sp. nov.; f – H. ser); g – labial palpus of H. ser; h–l – hind legs in dorsal view (h – H. yangae sp. nov.; i – H. ser; j – H. lamicola Zaitzev, 1908; k – detail of apex of tibia and base of tarsus of H. lamicola; l – detail of tarsomere 5 and claw of H. ser). Abbreviations: sh – swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="8" pageNumber="117">Fig. 1f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D71285EFF555FB170C7FA7F" blockId="8.[151,1127,796,1619]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF555FB17275FABF" box="[181,285,1276,1299]" italics="true" pageId="8" pageNumber="117">Pronotum</emphasis>
|
||
. Yellow, a little wider than the head, moderately and evenly arched, widest before the middle, sides weakly curved to base. Anterior angles sharper and more pronounced than is usual in
|
||
<taxonomicName id="78D34D0A8D71285EFEE85E7172EEFAFF" authorityName="Fabricius" authorityYear="1775" box="[264,390,1340,1363]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8DA7EA9B8D71285EFEE85E7172EEFAFF" box="[264,390,1340,1363]" italics="true" pageId="8" pageNumber="117">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
. Grooves scarcely traceable, mid groove straight, narrow; submedian grooves angled outwards medially, petering out anteriorly but sinuate a quarter of the way from each end; submarginal grooves with their basal pits distinct, traceable forward from these, petering out at middle of pronotum. Narrow raised lateral margin distinct. Intervals without granules but with large punctures and smaller ones between them.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D71285EFF555E917281F9FF" blockId="8.[151,1127,796,1619]" pageId="8" pageNumber="117">
|
||
<emphasis id="8DA7EA9B8D71285EFF555E917390FA5F" box="[181,248,1500,1523]" italics="true" pageId="8" pageNumber="117">Elytra</emphasis>
|
||
. Yellow, as pronotum, without apparent darker marks, slightly dull. Striae weak and shallow, scutellary stria present but often rather indistinct due to small size of serial punctures. Interval punctures only slightly smaller than the striae, with fine erect hairs. All intervals flat, including the 10
|
||
<superScript id="48A69BC18D71285EFE365D76728AF9E5" attach="none" box="[470,482,1595,1609]" fontSize="6" pageId="8" pageNumber="117">th</superScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D70285FFF705B9172A8FE9F" blockId="9.[113,1089,220,915]" pageId="9" pageNumber="118">
|
||
<emphasis id="8DA7EA9B8D70285FFF705B9173AAFF5F" box="[144,194,220,243]" italics="true" pageId="9" pageNumber="118">Legs</emphasis>
|
||
. Long, with long swimming-hairs on meso- and metatibiae and meso- and metatarsi; metatarsomere 5 ca. as long as metatarsomeres 3–4 combined; metatarsal claw ca. 0.65× as long as utimate metatarsomere.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D70285FFF705A7171F8FE1F" blockId="9.[113,1089,220,915]" pageId="9" pageNumber="118">
|
||
<emphasis id="8DA7EA9B8D70285FFF705A71739FFEFF" box="[144,247,316,339]" italics="true" pageId="9" pageNumber="118">Aedeagus</emphasis>
|
||
. Parameres pointed, very weakly to strongly sinuate on outer margin. Phallobase ca. as long as or slightly shorther than parameres. Median lobe wide, its distal portion short (in shape of more less isosceles triangle); base of median lobe (between the struts) rather straight but with median backwardly directed spur.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D70285FFF915AF1775AFD5F" blockId="9.[113,1089,220,915]" pageId="9" pageNumber="118">
|
||
<emphasis id="8DA7EA9B8D70285FFF915AF1738CFE7F" bold="true" box="[113,228,444,467]" pageId="9" pageNumber="118">Variation.</emphasis>
|
||
The species varies considerably in size and body proportions, i.e. both smaller narrower (
|
||
<figureCitation id="27E82A0C8D70285FFF3D5A917240FE5F" box="[221,296,476,499]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). a–c – H. lamicola Zaitzev, 1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Quinghai: Golo).d–g – H.ser Zaitzev, 1908 (d–e – Qinghai:Golo; f – Xizang, Nyainqentanglha Feng; g – Gansu: Xiahe env.). h–k – H. yangae sp. nov. (h – Xizang: 90 km W of Amdo; i – Xizang: 25 km W of Amdo; j – Qinghai: Golo; k – India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="9" pageNumber="118">Fig. 3b</figureCitation>
|
||
) and larger wide (
|
||
<figureCitation id="27E82A0C8D70285FFE025A917144FE5F" box="[482,556,476,499]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). a–c – H. lamicola Zaitzev, 1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Quinghai: Golo).d–g – H.ser Zaitzev, 1908 (d–e – Qinghai:Golo; f – Xizang, Nyainqentanglha Feng; g – Gansu: Xiahe env.). h–k – H. yangae sp. nov. (h – Xizang: 90 km W of Amdo; i – Xizang: 25 km W of Amdo; j – Qinghai: Golo; k – India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="9" pageNumber="118">Fig. 3c</figureCitation>
|
||
) are known. The shape of the sides of the pronotum varies slightly from weakly arcuate (
|
||
<figureCitation id="27E82A0C8D70285FFE195AB1712DFDBF" box="[505,581,508,531]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.a–c – H. lamicola Zaitzev,1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Qinghai: Golo). d–h – H. ser Zaitzev, 1908 (d – Qinghai: valley of Alag Hu lake (lectotype); e–f – Qinghai: Golo; g – Xizang, Nyainqentanglha Feng; h – Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="9" pageNumber="118">Fig. 4a</figureCitation>
|
||
) to indistinctly sinuate (
|
||
<figureCitation id="27E82A0C8D70285FFCA95AB170F4FDBF" box="[841,924,508,531]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.a–c – H. lamicola Zaitzev,1908 (a – Qinghai: Zhaling Hu (lectotype); b–c – Qinghai: Golo). d–h – H. ser Zaitzev, 1908 (d – Qinghai: valley of Alag Hu lake (lectotype); e–f – Qinghai: Golo; g – Xizang, Nyainqentanglha Feng; h – Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="9" pageNumber="118">Figs 4c</figureCitation>
|
||
). The aedeagus varies considerably in the shape of the apical portions of parameres; they may be narrow with lateral margin indistinctly sinuate (
|
||
<figureCitation id="27E82A0C8D70285FFE3C5971712BFDFF" box="[476,579,572,595]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">Figs 6f–g</figureCitation>
|
||
) or apically widely expanded with lateral margin strongly sinuate (
|
||
<figureCitation id="27E82A0C8D70285FFEC6591172E6FDDF" box="[294,398,604,627]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">Figs 6h–i</figureCitation>
|
||
); the proportion of the phallobase to parameres varies slightly, but no specimens with extremely long phallobase were found; the width of the apical portion of the median lobe varies from very wide (
|
||
<figureCitation id="27E82A0C8D70285FFDF359D17117FD1F" box="[531,639,668,691]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">
|
||
Figs
|
||
<quantity id="782B9B6C8D70285FFDA859D1710AFD1F" box="[584,610,668,691]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="9" pageNumber="118" unit="g" value="6.0">6g</quantity>
|
||
–h
|
||
</figureCitation>
|
||
) to rather narrow (
|
||
<figureCitation id="27E82A0C8D70285FFCA259D170CBFD1F" box="[834,931,668,691]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">Figs 6f, i</figureCitation>
|
||
). The apparent paler coloration of the aedeagus of the
|
||
<typeStatus id="6068882B8D70285FFDF159F1711BFD7F" box="[529,627,700,723]" pageId="9" pageNumber="118" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<figureCitation id="27E82A0C8D70285FFD6259F171A4FD7F" box="[642,716,700,723]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">Fig. 6f</figureCitation>
|
||
) in comparison to other specimens examined (
|
||
<figureCitation id="27E82A0C8D70285FFF0759917226FD5F" box="[231,334,732,755]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. a–e – H. (L.) ser Zaitzev, 1908 (a – lectotype, Qinghai: valley of Alag Hu lake; b–c – Xizang: valley SE of Dong La pass; d – Qinghai: Golo, 20 km E of Maduo; e – Qinghai: Zuimatan). f–j – H. (L.) lamicola Zaitzev, 1908 (f – lectotype, Qinghai: Djarin Nor; g–i – Qinghai: Zuimatan; j – paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). k–o – H. (L.) yangae sp. nov. (k – holotype, Qinghai: Zuimatan; l–m – paratypes, same locality; n – Qinghai: Golo, Yematan; o – Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">
|
||
Figs
|
||
<quantity id="782B9B6C8D70285FFEFD5991725FFD5F" box="[285,311,732,755]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="9" pageNumber="118" unit="g" value="6.0">6g</quantity>
|
||
–i
|
||
</figureCitation>
|
||
) is very likely caused by the fact that the
|
||
<typeStatus id="6068882B8D70285FFCEA59917004FD5F" box="[778,876,732,755]" pageId="9" pageNumber="118" type="lectotype">lectotype</typeStatus>
|
||
is slightly teneral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F7C965028D70285FFF9159B17070FC3F" pageId="9" pageNumber="118" type="distribution">
|
||
<paragraph id="BF6C36898D70285FFF9159B17753FCFF" blockId="9.[113,1089,220,915]" pageId="9" pageNumber="118">
|
||
<emphasis id="8DA7EA9B8D70285FFF9159B17394FCBF" bold="true" box="[113,252,764,787]" pageId="9" pageNumber="118">Distribution</emphasis>
|
||
(Fig. 7a). Confined to the northern part of the Tibetan Plateau, from Zuimatan in the east, via Lake Zhaling Hu (
|
||
<typeStatus id="6068882B8D70285FFE5B58517282FC9F" box="[443,490,796,819]" pageId="9" pageNumber="118">type</typeStatus>
|
||
locality) to Sven Hedin’s camp 23, situated south of the Arka-tagh mountains. The altitudes of the localities range from about
|
||
<quantity id="782B9B6C8D70285FFCBB5871776BFCFF" box="[859,1027,828,851]" metricMagnitude="3" metricUnit="m" metricValue="4.6049999999999995" metricValueMax="5.07" metricValueMin="4.14" pageId="9" pageNumber="118" unit="m" value="4605.0" valueMax="5070.0" valueMin="4140.0">4140 to 5070 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
<paragraph id="BF6C36898D70285FFF9158117070FC3F" blockId="9.[113,1089,220,915]" pageId="9" pageNumber="118">
|
||
<emphasis id="8DA7EA9B8D70285FFF91581173C6FCDF" bold="true" box="[113,174,860,883]" pageId="9" pageNumber="118">Note.</emphasis>
|
||
The record of
|
||
<taxonomicName id="78D34D0A8D70285FFEAD581172A3FCDF" authorityName="Zaitzev" authorityYear="1908" box="[333,459,860,883]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="lamicola">
|
||
<emphasis id="8DA7EA9B8D70285FFEAD581172A3FCDF" box="[333,459,860,883]" italics="true" pageId="9" pageNumber="118">H. lamicola</emphasis>
|
||
</taxonomicName>
|
||
from Apo Tso by
|
||
<bibRefCitation id="DB424B788D70285FFD6E58117033FCDF" author="ORCHYMONT A." box="[654,859,860,883]" pageId="9" pageNumber="118" pagination="1 - 16" refId="ref20836" refString="ORCHYMONT A. d' 1943: Les Palpicornia du Tibet (Coleoptera). Bulletin du Musee Royal d'Histoire Naturelle de Belgique 19 (57): 1 - 16." type="journal article" year="1943">ORCHYMONT (1943)</bibRefCitation>
|
||
was based on a misidentified specimen of
|
||
<taxonomicName id="78D34D0A8D70285FFEBE583172F7FC3F" authorityName="Zaitzev" authorityYear="1908" box="[350,415,892,915]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="ser">
|
||
<emphasis id="8DA7EA9B8D70285FFEBE583172F7FC3F" box="[350,415,892,915]" italics="true" pageId="9" pageNumber="118">H. ser</emphasis>
|
||
</taxonomicName>
|
||
(see under that species for details).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |